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Protomarctus optatus

Mammalia - Carnivora - Canidae

Taxonomy
Tomarctus optatus was named by Matthew (1924). Its type locality is Thomson Quarry, which is in a Hemingfordian terrestrial horizon in the Sheep Creek Formation of Nebraska.

It was recombined as Protomarctus optatus by Wang et al. (1999) and Wang et al. (2008).

Synonymy list
YearName and author
1924Tomarctus optatus Matthew
1999Protomarctus optatus Wang et al.
2008Protomarctus optatus Wang et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
RankNameAuthor
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyBorophaginae
tribeBorophaginiSimpson 1945
subtribeCynarctina
genusProtomarctus
speciesoptatus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Protomarctus optatus Matthew 1924
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Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: carnivoref
Diet 2: omnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-22 19:28:06
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Lillegraven 1979, Hendy et al. 2009, Carroll 1988, Nowak 1991, Ji et al. 2002

Age range: base of the Barstovian to the top of the Early/Lower Hemingfordian or 16.30000 to 15.97000 Ma

Collections (27 total)


Time interval Ma Country or state Original ID and collection number
Burdigalian20.44 - 15.97USA (California) Tomarctus optatus (19151)
Early/Lower Hemingfordian20.43 - 15.97USA (New Mexico) Protomarctus optatus (19311) Tomarctus sp. (19188)
Early/Lower Hemingfordian20.43 - 15.97USA (Nebraska) Tomarctus optatus (17864 17906)
Hemingfordian20.43 - 15.97USA (Colorado) Tomarctus sp. (17965)
Hemingfordian20.43 - 15.97USA (Wyoming) Tomarctus optatus (18873 19073 19075)
Hemingfordian20.43 - 15.97USA (Nebraska) Protomarctus optatus (17872 17890 17918 17920 17952 17961 17987 18000 18012 18016) Tomarctus optatus (17907 17917 type locality: 18001)
Hemingfordian20.43 - 15.97USA (Nevada) Tomarctus optatus (18962)
Hemingfordian - Barstovian20.43 - 12.5USA (California) Tomarctus optatus (19601)
Barstovian16.3 - 12.5USA (Nevada) Tomarctus optatus (19103)
Barstovian16.3 - 12.5USA (California) Protomarctus optatus (19510)
Barstovian16.3 - 12.5USA (New Mexico) Tomarctus optatus (19449)