PBDB Taxon

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Eubrontes

Reptilia - Theropoda - Grallatoridae

Taxonomy

Eubrontes was named by Hitchcock (1845). It is considered to be a form taxon.

It was reranked as Grallator (Eubrontes) by Olsen and Galton (1984), Gierlinski (1991) and Milàn and Gierlinski (2004).

It was assigned to Struthionoidae by Hitchcock (1845); to Carnosauria by Lapparent and Montenat (1967) and Vignaud et al. (1994); to Theropoda by Madsen and Miller (1979), Lockley (1992) and Romero Molina et al. (2003); to Grallator by Olsen and Galton (1984), Gierlinski (1991) and Milàn and Gierlinski (2004); to Theropoda by Lull (1917) and Weishampel (2006); to Prosauropoda by Weems (1987) and Weems (2006); to Theropoda by Lucas et al. (2006), Niedźwiedzki (2011) and Salisbury et al. (2016); to Eubrontidae by Lull (1904), Lull (1915), Hitchcock and Grier (1927), Lull (1953), Kuhn (1958), Kuhn (1963), Haubold (1969), Haubold (1971), Demathieu and Haubold (1972), Haubold (1974), Leonardi (1979), Zhen et al. (1986), Zhen et al. (1989), Leonardi and Mietto (2000), Gao (2007), Li et al. (2010), Marty et al. (2018) and Tsukiji et al. (2019); and to Grallatoridae by Lucas et al. (2006) and Klein and Lucas (2021).

Synonyms

Weiyuanpus was named by Gao (2007).

It was synonymized subjectively with Eubrontes by Lockley et al. (2013) and Xing et al. (2013).

It was assigned to Eubrontidae by Gao (2007).
Brontozoum isodactylum was named by Hitchcock (1858). It is considered to be a form taxon. It is a trace fossil.

It was synonymized subjectively with Anomoepus intermedius by Hitchcock (1862); it was synonymized subjectively with Brontozoum fulicoides by Quenstedt (1882); it was considered a nomen dubium by Hitchcock (1865).
Zizhongpus was named by Yang and Yang (1987). It is considered to be a form taxon. Its type is Zizhongpus wumanensis.

It was synonymized subjectively with Kayentapus by Lockley et al. (2013); it was synonymized subjectively with Eubrontes by Xing et al. (2021).

It was assigned to Anchisauripodidae by Yang and Yang (1987).
Zizhongpus wumaensis was named by Yang and Yang (1987). It is considered to be a form taxon. Its type specimen is CFZW2–8, 12–29, a footprint, and it is a trace fossil.

It was considered a nomen dubium by Xing et al. (2021); it was recombined as Kayentapus wumaensis by Lockley et al. (2013), Xing et al. (2014) and Li et al. (2021).
Tuojiangpus was named by Yang and Yang (1987). It is considered to be a form taxon.

It was considered a nomen dubium by Lockley et al. (2013); it was synonymized subjectively with Eubrontes by Xing et al. (2021).

It was assigned to Anchisauripodidae by Yang and Yang (1987).
Tuojiangpus shuinanensis was named by Yang and Yang (1987). It is considered to be a form taxon. Its type specimen is CFZW83, a footprint, and it is a trace fossil. Its type locality is Wu Ma Cun, site A (Rice cleaning), which is in an Aalenian/Bajocian terrestrial sandstone in the Xintiangou Formation of China.

It was considered a nomen dubium by Lockley et al. (2013) and Xing et al. (2021).
Chonglongpus was named by Yang and Yang (1987). It is considered to be a form taxon.

It was synonymized subjectively with Gigandipus by Lockley et al. (2013) and Xing et al. (2021).

It was assigned to Gigandipodidae by Yang and Yang (1987).
Megaichnites was named by Yang and Yang (1987). It is considered to be a form taxon.

It was synonymized subjectively with Kayentapus by Lockley et al. (2013); it was synonymized subjectively with Eubrontes by Xing et al. (2021).

It was assigned to Eubrontidae by Yang and Yang (1987).
Megaichnites jizhaoshiensis was named by Yang and Yang (1987). It is considered to be a form taxon. Its type specimen is CFZW161, 164, a footprint, and it is a trace fossil.

It was recombined as Kayentapus jizhaoshiensis by Lockley et al. (2013) and Li and Zhang (2017); it was misspelled as Kayentapus jizhaishiensis by Xing et al. (2014); it was considered a nomen dubium by Xing et al. (2021).
Gigadipus was named by Hitchcock (1855). It is considered to be a form taxon.

It was corrected as Gigandipus by Lull (1904), Lull (1915), Lull (1917), Bock (1952), Lull (1953), Kuhn (1958), Kuhn (1963), Haubold (1969), Haubold (1971), Haubold (1974), Yang and Yang (1987) and Romero Molina et al. (2003); it was synonymized subjectively with Eubrontes by Weems (2003), Hunt and Lucas (2006), Lucas et al. (2006) and Rainforth (2007).

It was assigned to Theropoda by Lull (1917); to Gigandipodidae by Lull (1904), Lull (1915), Bock (1952), Lull (1953), Kuhn (1958), Kuhn (1963), Haubold (1969), Haubold (1971), Haubold (1974) and Yang and Yang (1987); and to Theropoda by Romero Molina et al. (2003).
Otouphepus was named by Cushman (1904). It is considered to be a form taxon. It is the type genus of Otouphepodidae.

It was synonymized subjectively with Anchisauripus by Kuhn (1963), Haubold (1969) and Knoll (2005); it was reranked as Anchisauripus (Otouphepus) by Haubold (1971); it was considered a nomen dubium by Baird (1957); it was synonymized subjectively with Eubrontes by Rainforth (2007).

It was assigned to Theropoda by Lull (1917); to Anchisauripodidae by Lull (1953); to Otouphepodidae by Lull (1915), Bock (1952) and Kuhn (1958); to Anchisauripus by Haubold (1971); to Trisauropodia by Ellenberger (1972); and to Theropoda by Romero Molina et al. (2003).
Hyphepus was named by Hitchcock (1858).

It was synonymized subjectively with Eubrontes by Rainforth (2007).

It was assigned to Dinosauria by Hitchcock (1898); to Theropoda by Lull (1917); to Carnosauria by Kuhn (1958); to Gigandipodidae by Lull (1904), Lull (1915), Lull (1953), Kuhn (1963), Haubold (1969), Haubold (1971) and Haubold (1974); and to Theropoda by Romero Molina et al. (2003).
Paracoelurosaurichnus was named by Zhen et al. (1986). It is considered to be a form taxon.

It was synonymized subjectively with Eubrontes by Lockley et al. (2013).

It was assigned to Anchisauripodidae by Zhen et al. (1986) and Zhen et al. (1989); and to Theropoda by Romero Molina et al. (2003).
Youngichnus was named by Zhen et al. (1986). It is considered to be a form taxon.

It was synonymized subjectively with Eubrontes by Lockley et al. (2013).

It was assigned to Theropoda by Romero Molina et al. (2003); and to Eubrontidae by Zhen et al. (1986), Zhen et al. (1989) and Gao (2007).
Youngichnus xiyangensis was named by Zhen et al. (1986). It is considered to be a form taxon. Its type specimen is BPV-FP6, a footprint, and it is a trace fossil. Its type locality is Xiiyang, Jinning tracksites (BPV), which is in a Hettangian terrestrial horizon in the Fengjiahe Formation of China.

It was considered a nomen dubium by Xing et al. (2021); it was recombined as Eubrontes xiyangensis by Lockley et al. (2013), Xing et al. (2014), Li and Zhang (2017) and Li et al. (2021).
Ornithichnites was named by Hitchcock (1836).

It was synonymized subjectively with Eubrontes by Haubold (1971).

It was assigned to Aves by Owen (1861).
Brontozoum was named by Hitchcock (1847) [Haubold created two subgenera from Brontozoum: Anchisauripus (Brontozoum) and Eubrontes (Brontozoum). I have made this second subgenus a new taxon by necessity, but he did not specify as such.]. It is considered to be a form taxon.

It was synonymized subjectively with Eubrontes by Hay (1902), Lull (1904), Lull (1915), Hay (1930), Lapparent (1945), Lull (1953), Baird (1957), Haubold (1969), Haubold (1971), Olsen et al. (1998), Rainforth (2007) and Klein and Lucas (2021).

It was assigned to Struthionidae by Hitchcock (1848); to Pachydactyli by Quenstedt (1882); to Aves by Owen (1861) and Hitchcock (1898); to Theropoda by Gadow (1898) and Gadow (1901); and to Megalosaurinidae by Mansel-Pleydell (1902).
Gigantitherium was named by Hitchcock (1858). It is considered to be a form taxon.

It was synonymized subjectively with Gigadipus by Lull (1904), Lull (1915) and Kuhn (1963); it was synonymized subjectively with Gigandipus by Hay (1902), Hay (1930), Haubold (1969) and Haubold (1971).

It was assigned to Leptodactyli by Quenstedt (1882); and to Dinosauria by Hitchcock (1898).
Fulicopus was named by Hitchcock (1845). It is considered to be a form taxon. Its type is Ornithoidichnites lyellii.

It was synonymized subjectively with Sauropus by Lull (1915) and Lull (1953); it was synonymized subjectively with Eubrontes by Rainforth (2007).

It was assigned to Pteropodidae by Hitchcock (1845); and to Anomoepodidae by Lull (1904).
Amblonyx was named by Hitchcock (1858). It is considered to be a form taxon.

It was corrected as Amblyonyx by Quenstedt (1882); it was synonymized subjectively with Fulicopus by Hay (1902) and Hay (1930); it was synonymized subjectively with Sauropus by Lull (1915), Lull (1953), Kuhn (1958) and Haubold (1969).

It was assigned to Pachydactyli by Quenstedt (1882); and to Aves by Hitchcock (1858) and Hitchcock (1898).
Ichnites was named by Haubold (1971). It is considered to be a form taxon.

It was synonymized subjectively with Eubrontes by Haubold (1971).
Lufengopus was named by Lü et al. (2006).

It was synonymized subjectively with Eubrontes by Xing et al. (2021).

It was assigned to Eubrontidae by Lü et al. (2006).
Lufengopus dongi was named by Lü et al. (2006). Its type specimen is HYMVC-1, a footprint, and it is a trace fossil. Its type locality is Beikeshan, Yaozhan, Lufeng, which is in a Jurassic terrestrial mudstone/conglomerate in the Chuanjie Formation of China.

It was considered a nomen dubium by Lockley et al. (2013); it was considered a nomen dubium by Xing et al. (2021).

Synonymy list

Year	Name and author
1836	Ornithichnites Hitchcock p. 315
1845	Eubrontes Hitchcock p. 23
1845	Fulicopus Hitchcock p. 23
1847	Brontozoum Hitchcock p. 50
1848	Brontozoum Hitchcock p. 169
1858	Amblonyx Hitchcock pp. 63, 70
1858	Brontozoum isodactylum Hitchcock pp. 69-70
1858	Amblonyx Hitchcock p. 70
1858	Gigantitherium Hitchcock p. 93
1858	Hyphepus Hitchcock pp. 97, 180
1861	Ornithichnites Owen p. 325
1861	Brontozoum Owen p. 327
1882	Brontozoum Quenstedt p. 130
1882	Amblyonyx Quenstedt p. 131
1882	Gigantitherium Quenstedt p. 132
1889	Brontozoum isodactylum Eyerman p. 32
1894	Brontozoum isodactylum Lyman p. 205
1898	Brontozoum Gadow p. 22
1898	Amblonyx Hitchcock p. 401
1898	Brontozoum Hitchcock p. 401
1898	Gigantitherium Hitchcock p. 401
1898	Hyphepus Hitchcock p. 401
1901	Brontozoum Gadow p. 420
1902	Brontozoum Mansel-Pleydell p. lxxviii
1904	Otouphepus Cushman p. 155
1904	Gigandipus Lull p. 492
1904	Hyphepus Lull p. 493
1904	Fulicopus Lull p. 505
1904	Eubrontes Lull p. 510
1915	Otouphepus Lull p. 187
1915	Gigandipus Lull p. 191
1915	Hyphepus Lull p. 193
1915	Eubrontes Lull p. 195
1917	Otouphepus Lull p. 114
1917	Eubrontes Lull p. 115
1917	Gigandipus Lull p. 115
1917	Hyphepus Lull p. 115
1927	Eubrontes Hitchcock and Grier p. 187
1952	Gigandipus Bock p. 403
1952	Otouphebus Bock p. 407
1953	Otouphepus Lull p. 176
1953	Eubrontes Lull p. 178
1953	Gigandipus Lull p. 184
1953	Hyphepus Lull p. 186
1958	Eubrontes Kuhn p. 22
1958	Gigandipus Kuhn p. 23
1958	Hyphepus Kuhn p. 23
1958	Otouphepus Kuhn p. 23
1963	Eubrontes Kuhn p. 83
1963	Gigandipus Kuhn p. 84
1963	Hyphepus Kuhn p. 85
1967	Eubrontes Lapparent and Montenat p. 33
1969	Eubrontes Haubold p. 99
1969	Gigandipus Haubold p. 99
1969	Hyphepus Haubold p. 99
1971	Ichnites Haubold
1971	Anchisauripus (Otouphepus) Haubold p. 65
1971	Eubrontes Haubold p. 75
1971	Gigandipus Haubold p. 77
1971	Hyphepus Haubold p. 77
1972	Eubrontes Demathieu and Haubold p. 806
1972	Otouphepus Ellenberger p. 152
1974	Eubrontes Haubold p. 45
1974	Gigandipus Haubold p. 45
1974	Hyphepus Haubold p. 45
1979	Eubrontes Leonardi p. 244
1979	Eubrontes Madsen and Miller p. 36
1984	Grallator (Eubrontes) Olsen and Galton p. 97
1986	Paracoelurosaurichnus Zhen et al. p. 4
1986	Eubrontes Zhen et al. p. 9
1986	Youngichnus Zhen et al. p. 10 fig. 8
1986	Youngichnus xiyangensis Zhen et al. p. 10–11 fig. 8
1987	Eubrontes Weems p. 17
1987	Chonglongpus Yang and Yang
1987	Zizhongpus Yang and Yang p. 9 fig. 4
1987	Zizhongpus wumaensis Yang and Yang p. 9–10 fig. 4
1987	Tuojiangpus Yang and Yang p. 17 fig. 12
1987	Tuojiangpus shuinanensis Yang and Yang p. 17 fig. 12
1987	Chonglongpus Yang and Yang p. 18 fig. 13
1987	Gigandipus Yang and Yang p. 19
1987	Megaichnites Yang and Yang p. 19
1987	Megaichnites jizhaoshiensis Yang and Yang p. 19 fig. 14
1989	Eubrontes Zhen et al. pp. 192-193
1989	Paracoelurosaurichnus Zhen et al. p. 192
1989	Youngichnus Zhen et al. p. 193
1989	Youngichnus xiyangensis Zhen et al. p. 193
1991	Grallator (Eubrontes) Gierlinski p. 137
1992	Eubrontes Lockley p. 272
1994	Eubrontes Vignaud et al. p. 497
2000	Eubrontes Leonardi and Mietto p. 197
2003	Eubrontes Romero Molina et al. p. 24
2003	Gigandipus Romero Molina et al. p. 24
2003	Hyphepus Romero Molina et al. p. 24
2003	Otouphepus Romero Molina et al. p. 24
2003	Paracoelurosaurichnus Romero Molina et al. p. 24
2003	Youngichnus Romero Molina et al. p. 24
2004	Grallator (Eubrontes) Milàn and Gierlinski p. 71
2006	Eubrontes Lucas et al. p. 86–87
2006	Eubrontes Lucas et al. p. 114
2006	Lufengopus Lü et al. pp. 35-36
2006	Lufengopus dongi Lü et al. p. 36 figs. 3A-B
2006	Eubrontes Weems p. 372
2006	Eubrontes Weishampel p. 137
2007	Weiyuanpus Gao p. 342
2007	Eubrontes Gao p. 345
2007	Youngichnus Gao p. 345
2007	Youngichnus xiyangensis Gao p. 345
2007	Lufengopus dongi Lü et al. p. 441
2008	Lufengopus dongi Fujita et al. p. 33
2008	Youngichnus xiyangensis Fujita et al. p. 33
2009	Lufengopus dongi Xing et al. p. 17
2009	Youngichnus xiyangensis Xing et al. p. 23
2010	Eubrontes Li et al. p. 726
2011	Eubrontes Niedźwiedzki p. 296
2013	Eubrontes xiyangensis Lockley et al. p. 6
2013	Kayentapus jizhaoshiensis Lockley et al. p. 6
2013	Kayentapus wumaensis Lockley et al. p. 6
2014	Eubrontes xiyangensis Xing et al. p. 344
2014	Kayentapus jizhaishiensis Xing et al. p. 310
2014	Kayentapus wumaensis Xing et al. p. 310
2016	Eubrontes Salisbury et al. p. 36
2017	Eubrontes xiyangensis Li and Zhang p. 280
2017	Kayentapus jizhaoshiensis Li and Zhang p. 280
2017	Lufengopus dongi Li and Zhang p. 280
2017	Youngichnus xiyangensis Xing et al. p. 115
2018	Eubrontes Marty et al. p. 943
2019	Eubrontes Tsukiji et al. p. 6
2021	Eubrontes Klein and Lucas p. 10
2021	Eubrontes xiyangensis Li et al. p. 14
2021	Kayentapus wumaensis Li et al. p. 18
2022	Lufengopus dongi Li et al. p. 4235

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
order	Theropoda
family	Grallatoridae
genus	Eubrontes

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Eubrontes Hitchcock 1845

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Subg. †Eubrontes (Grallator) Olsen 1981

†Eubrontes euskelosauroides Huene 1941

†Eubrontes expansus Hitchcock 1841

†Eubrontes fieldi Hitchcock 1858

†Eubrontes giganteus Hitchcock 1836

Invalid names: Eubrontes approximatus Hitchcock 1865 [synonym], Eubrontes divaricatus Hitchcock 1865 [synonym]

†Eubrontes glenrosensis Shuler 1935

†Eubrontes lyellii Hitchcock 1843

†Eubrontes minusculus Hitchcock 1837

Invalid names: Eubrontes tuberatus Hitchcock 1858 [synonym], Plesiornis giganteus Hitchcock 1889 [synonym]

†Eubrontes monax Zhen et al. 1986

†Eubrontes nianpanshanensis Yang and Yang 1987

†Eubrontes nobitai Xing et al. 2021

†Eubrontes pareschequier Xing et al. 2009

†Eubrontes platypus Hitchcock 1858

†Eubrontes sillimani Hitchcock 1843

Invalid names: Anchisauripus dananus Hitchcock 1845 [synonym], Ornithichnites cuneatus Hitchcock 1837 [synonym], Otouphepus minor Lull 1915 [synonym]

†Eubrontes tatricus Michalík and Sykora 1976

†Eubrontes veillonensis Lapparent and Montenat 1967

†Eubrontes zigongensis Gao 2007

Invalid names: Brontozoum Hitchcock 1847 [synonym], Eubrontes xiyangensis Zhen et al. 1986 [nomen dubium], Fulicopus Hitchcock 1845 [synonym], Gigandipus Hitchcock 1855 [synonym], Hyphepus Hitchcock 1858 [synonym], Ichnites Haubold 1971 [synonym], Lufengopus Lü et al. 2006 [synonym], Lufengopus dongi Lü et al. 2006 [nomen dubium], Megaichnites Yang and Yang 1987 [synonym], Megaichnites jizhaoshiensis Yang and Yang 1987 [nomen dubium], Ornithichnites Hitchcock 1836 [synonym], Otouphepus Cushman 1904 [synonym], Paracoelurosaurichnus Zhen et al. 1986 [synonym], Tuojiangpus Yang and Yang 1987 [synonym], Tuojiangpus shuinanensis Yang and Yang 1987 [nomen dubium], Weiyuanpus Gao 2007 [synonym], Youngichnus Zhen et al. 1986 [synonym], Zizhongpus Yang and Yang 1987 [synonym], Zizhongpus wumaensis Yang and Yang 1987 [nomen dubium]

Diagnosis

Reference	Diagnosis
E. Hitchcock 1848 (Brontozoum)	Foot tridactylous, pachydactylous, tubercular-clawed; inner toe shortest; all of them directed forward. Phalangeal expansions on the inner toe, two; on the middle toe, three; on the outer toe, four; corresponding to the number of phalanges, except the distal expansion, which was probably made by the two extreme phalanges. Lower extremity of the tarso-metatarsal bone double-headed; rarely making a distinct impression through the cushion beneath. Cushion sloping upwards posteriorly. Claws on the lateral toes a little outside of their axes; on the middle toe, a little towards its inner side.
R. S. Lull 1915 (Gigandipus)	Those of the family, the most readily recognized being the small hallux which lies at right angles to digit II, instead of being rotated far to the rear as in Anchisauripus.
R. S. Lull 1915	Those of the family.
R. S. Lull 1915 (Hyphepus)	Bipedal, tetradactylous, hallux incumbent, curved sharply backward. Phalangeal pads obscurely defined; anterior digits apparently clawless.
H. Klein and S. G. Lucas 2021	Large (> 30 cm long), functionally tridactyl ichnite in which the digit III projection ratio is about 2.2, and the length-to-width ratio is about 1.4 to 1.5. Projection of digits II and IV along the axis of digit III about equal. Divarication of outer digits 30°-40°.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

Reference: Marsh 1875

Age range: base of the Smithian to the top of the Early/Lower Albian or 251.30000 to 109.00000 Ma

Collections (206 total)

Time interval	Ma	Country or state	Original ID and collection number
Smithian	251.3 - 247.2	USA (Utah)	E. sp. (105542)
Carnian	237.0 - 228.0	Morocco (Marrakech-Safi)	E. sp. (205518 205528 230183)
Late/Upper Triassic	237.0 - 201.4	USA (New Jersey)	Brontozoum giganteum, Brontozoum sillimanium, Brontozoum minusculum (91694)
Late/Upper Triassic	237.0 - 201.4	USA (Pennsylvania)	Anchisauripus sillimani (52494)
Norian	228.0 - 208.5	Brazil (Rio Grande do Sul)	E. sp. (157877)
Norian	228.0 - 208.5	Canada (Nova Scotia)	Grallator sillimani (138975)
Norian	228.0 - 208.5	United Kingdom (Wales)	Gigandipus sp. (57949)
Norian	228.0 - 208.5	Italy (Friuli-Venezia Giulia)	E. sp. (194710)
Norian	228.0 - 208.5	Switzerland (Graubunden)	Theropoda indet. (69623)
Norian	228.0 - 208.5	France (Occitanie)	E. sp. (194071)
Norian	228.0 - 208.5	France (Languedoc-Roussillon)	Dinosauria indet. (58623 58624)
Norian	228.0 - 208.5	USA (Pennsylvania)	Anchisauripus sillimani (55414) Brontozoum sillimanium (55413)
Norian	228.0 - 208.5	USA (New Mexico)	Coelurosaurichnus sp. (84021)
Norian	228.0 - 208.5	USA (New Jersey)	Anchisauripus minuscolus (187287) Anchisauripus sillimani (55418)
Norian	228.0 - 208.5	USA (Arizona)	Theropoda indet. (62286)
Norian - Rhaetian	228.0 - 201.3	USA (Utah)	Anchisauripus sp. (83555)
Norian - Rhaetian	228.0 - 201.3	Slovakia (Tatra)	E. sp., Coelurosaurichnus tatricus (139740)
Lacian - Alaunian	221.5 - 212.0	USA (Virginia)	Apatichnus minor (68890) Grallator sillimani (68889)
Alaunian - Sevatian	215.56 - 205.6	Switzerland (Graubünden)	E. sp. (220115)
Rhaetian	208.5 - 201.3	USA (Pennsylvania)	E. giganteus (62768)
Rhaetian	208.5 - 201.3	France (Pays de la Loire)	E. sp. (24798)
Rhaetian	208.5 - 201.3	Sweden (Kristianstads)	Theropoda indet. (51979)
Rhaetian	208.5 - 201.3	USA (Utah)	E. sp. (223701)
Rhaetian	208.5 - 201.3	USA (New Jersey)	E. giganteus, Anchisauripus sillimani (194093)
Rhaetian - Hettangian	208.5 - 199.3	USA (Utah)	E. sp. (157018)
Rhaetian - Hettangian	208.5 - 199.3	USA (Arizona)	E. sp. (38886)
Rhaetian - Early/Lower Hettangian	208.5 - 196.5	France (Occitanie)	E. giganteus, Grallator minusculus (206455)
Early/Lower Jurassic	201.4 - 174.7	China (Sichuan)	Weiyuanpus zigongensis (60499)
Early/Lower Jurassic	201.4 - 174.7	Namibia (Otjozondjupa)	E. giganteus (214423)
Early/Lower Jurassic	201.4 - 174.7	China (Nei Mongol)	E. glenrosensis (107973)
Early/Lower Jurassic	201.4 - 174.7	China (Shaanxi)	E. sp. (224366)
Hettangian	201.3 - 199.3	USA (Massachusetts)	Anchisauripus minusculus, Brontozoum giganteum, Brontozoum approximatum, Brontozoum sillimanium (53024) Brontozoum divaricatum (53030) Brontozoum giganteum (53029) Dinosauria indet. (187029) E. giganteus, E. platypus, E. divaricatus, E. tuberatus (69798) E. sp. (55260 55331) Gigadipus caudatus, Otouphepus minor, Brontozoum giganteum, Brontozoum approximatum, Brontozoum divaricatum, Brontozoum tuberatum, Hyphepus fieldi, Ornithoidichnites lyellii, Brontozoum sillimanium, Brontozoum minusculum (52986)
Hettangian	201.3 - 199.3	Italy (Trentino-Alto Adige)	E. sp. (88587)
Hettangian	201.3 - 199.3	France (Region Midi-Pyrenees)	Grallator minusculus (31307 31308)
Hettangian	201.3 - 199.3	USA (Colorado)	E. sp. (183639)
Hettangian	201.3 - 199.3	France (Aveyron)	E. divaricatus (55480)
Hettangian	201.3 - 199.3	China (Yunnan)	Paracoelurosaurichnus monax, E. platypus, Youngichnus xiyangensis (52848)
Hettangian	201.3 - 199.3	Canada (Nova Scotia)	Grallator minusculus, Grallator sillimani (227833)
Hettangian	201.3 - 199.3	USA (Connecticut)	Anchisauripus sillimani (46438) E. approximatus (51255 51263) E. approximatus, Gigandipus caudatus (51260) E. giganteus (51261) E. giganteus, Anchisauripus sillimani (51256) E. sp., E. giganteus, Anchisauripus sillimani (55741) Ornithoidichnites giganteus (91576)
Hettangian	201.3 - 199.3	USA (New Jersey)	Hyphepus sp. (55527)
Hettangian	201.3 - 199.3	USA (Arizona)	E. sp. (225154)
Hettangian	201.3 - 199.3	France (Occitanie)	E. sp. (219703 223710 223711)
Hettangian	201.3 - 199.3	USA (Utah)	E. sp. (56131 121311 121321 157013 194553 194862) E. sp., Theropoda indet. (121305) Theropoda indet. (121320)
Early/Lower Hettangian	201.3 - 196.5	USA (Connecticut)	E. sp. (192863)
Early/Lower Hettangian	201.3 - 196.5	USA (Utah)	Plateosauridae indet. (52153)
Early/Lower Hettangian	201.3 - 196.5	Poland (Swietokrzyskie)	E. sp. (84260)
Early/Lower Hettangian	201.3 - 196.5	France	E. sp. (223717 223718 223719)
Early/Lower Hettangian	201.3 - 196.5	Canada (Nova Scotia)	E. giganteus, Anchisauripus sillimani (138976 138977) Grallator minusculus, Anchisauripus sillimani, E. giganteus (138978) Grallator sillimani (227835)
Late/Upper Hettangian	201.3 - 196.5	France (Languedoc-Roussillon)	Theropoda indet. (140450)
Late/Upper Hettangian	201.3 - 196.5	Italy (Trentino-Alto Adige)	E. sp. (61851)
Hettangian - Sinemurian	201.3 - 190.8	Lesotho (Maseru)	E. sp. (214417 230716)
Hettangian - Sinemurian	201.3 - 190.8	USA (Connecticut)	Brontozoum giganteum, Brontozoum divaricatum, Brontozoum sillimanium, Brontozoum isodactylum (91574) Brontozoum giganteum, Brontozoum sillimanium (91573) E. giganteus, E. divaricatus, Brontozoum sillimanium (53023) Ornithoidichnites sillimani, Ornithoidichnites giganteus, E. dananus, Ornithoidichnites cuneatus (53033) Plesiornis giganteus, Ornithoidichnites giganteus, Brontozoum divaricatum, Brontozoum sillimanium, Ornithoidichnites expansus, Ornithoidichnites cuneatus (53000)
Hettangian - Sinemurian	201.3 - 190.8	Lesotho (Quthing)	E. sp. (56026 214416)
Hettangian - Sinemurian	201.3 - 190.8	USA (Massachusetts)	Anchisauripus minusculus, Brontozoum giganteum, Brontozoum sillimanium, Brontozoum isodactylum, Ornithoidichnites cuneatus (52996) Anchisauripus minusculus, Brontozoum giganteum, E. divaricatus, Brontozoum sillimanium, Brontozoum isodactylum, Ornithoidichnites cuneatus (52995) Anchisauripus minusculus, Brontozoum sillimanium, Ornithoidichnites cuneatus (52994) Brontozoum giganteum (91577 91666) Brontozoum giganteum, Brontozoum approximatum, Brontozoum divaricatum, Brontozoum sillimanium, Brontozoum minusculum (52985) Brontozoum giganteum, Brontozoum approximatum, Hyphepus fieldi, Amblonyx lyellianus, Brontozoum sillimanium, Brontozoum minusculum (52998) Brontozoum sillimaniium, Brontozoum giganteum, Brontozoum approximatum, Brontozoum divaricatum, Brontozoum minusculum (62844) Brontozoum sillimanium (52999 91662 91669) Brontozoum sp., Anchisauripus sillimani, Brontozoum giganteum (53028) Brontozoum tuberatum, Hyphepus fieldi, Amblonyx lyellianus, Amblonyx giganteus, Brontozoum sillimanium (52987) E. sp. (53027 222121) Ornithichnites elegantior (52993) Ornithichnites giganteus, Brontozoum approximatum, Brontozoum tuberatum, Brontozoum sillimanium, Brontozoum isodactylum, Ornithoidichnites expansus (53025) Ornithoidichnites giganteus (91398 91578)
Hettangian - Early/Lower Sinemurian	201.3 - 189.6	Italy (Trentino-Alto Adige)	E. sp. (68887 194080)
Hettangian - Pliensbachian	201.3 - 182.7	USA (Utah)	E. sp. (184775 222900)
Hettangian - Pliensbachian	201.3 - 182.7	USA (Arizona)	E. giganteus, Anchisauripus minusculus (225199)
Hettangian - Bajocian	201.3 - 168.3	India (Rajasthan)	E. giganteus (168396)
Sinemurian	199.3 - 190.8	China (Yunnan)	Changpeipus pareschequier (182648)
Sinemurian	199.3 - 190.8	USA (New Jersey)	Brontozoum approximatum (91695)
Sinemurian	199.3 - 190.8	Australia (Queensland)	E. sp. (60756)
Sinemurian - Pliensbachian	199.3 - 182.7	USA (Arizona)	E. sp. (36272)
Sinemurian - Pliensbachian	199.3 - 182.7	USA (Utah)	E. sp. (183521 184778 194145 194146 194147 194148 223699) E. sp., Grallator sp. (57067)
Early/Lower Sinemurian	196.5 - 189.6	France (Lot)	Theropoda indet. (25843)
Early/Lower Sinemurian	196.5 - 189.6	USA (Virginia)	E. giganteus, E. approximus (104029)
Pliensbachian	190.8 - 182.7	Lesotho (Leribe)	E. sp. (55996)
Pliensbachian	190.8 - 182.7	Lesotho	Brontozoum sp. (55998) Otouphepus minor, Otouphepus declivis (84073)
Pliensbachian	190.8 - 182.7	USA (Utah)	E. sp. (225150 225151 229818)
Pliensbachian - Toarcian	190.8 - 174.1	USA (Utah)	E. sp. (49586 49587 49588 49589 49590 49591 49592 49593 49594 184776 201625 222907 222908 222909 222913) Ornithopoda indet. (60602)
Pliensbachian - Toarcian	190.8 - 174.1	USA (Arizona)	Dinosauria indet. (58556) E. sp. (51976 86178)
Early/Lower Pliensbachian	189.6 - 183.0	Morocco (Er Rachida)	E. sp. (84210 91905)
Toarcian - Bajocian	182.7 - 168.3	China (Sichuan)	Theropoda indet. (103971)
Toarcian - Bathonian	182.7 - 166.1	Morocco (Azilal)	Saurischia indet. (37042 37043)
Middle Jurassic	174.7 - 161.5	China (Yunnan)	Lufengopus dongi (66927)
Middle Jurassic - Late/Upper Jurassic	174.7 - 145.0	Tajikistan (Kulyabskaya Oblast')	Brontozoum thianschanicum (53144)
Middle Jurassic - Late/Upper Jurassic	174.7 - 145.0	Zimbabwe (Mashonaland West)	E. sp. (45532)
Middle Jurassic - Late/Upper Jurassic	174.7 - 145.0	China (Yunnan)	E. sp. (86424)
Aalenian	174.1 - 170.3	United Kingdom (England)	E. sp. (202018)
Aalenian - Bajocian	174.1 - 168.3	China (Sichuan)	Jinlijingpus nianpanshanensis (60498) Megaichnites jizhaishiensis (60494) Tuojiangpus shuinanensis, Chonglongpus hei, Zizhongpus wumanensis (60493)
Early/Lower Bathonian	167.7 - 164.7	France (Occitanie)	E. sp. (223713) Grallator minusculus (223714)
Middle Bathonian	167.7 - 164.7	United Kingdom (Scotland)	E. sp. (32318)
Middle Bathonian - Late/Upper Bathonian	167.7 - 164.7	United Kingdom (Scotland)	Gigandipus sp. (55423)
Oxfordian - Early/Lower Kimmeridgian	163.5 - 150.8	USA (Colorado)	E. giganteus (183640)
Berriasian	145.0 - 139.8	China (Hebei)	E. sp. (52850)
Berriasian - Hauterivian	145.0 - 129.4	Brazil (Paraíba)	E. sp. (58787 58800)
Berriasian - Barremian	145.0 - 125.0	China (Sichuan)	Theropoda indet. (182655)
Early/Lower Cretaceous	145.0 - 100.5	China (Sichuan)	E. sp. (220010)
Valanginian	139.8 - 132.9	China (Beijing)	E. sp. (153397 220020)
Valanginian - Hauterivian	139.8 - 129.4	China (Sichuan)	E. sp. (220045)
Late/Upper Barremian	130.0 - 125.45	China (Nei Mongol)	E. sp. (49342 60511 66928)
Barremian	129.4 - 125.0	China (Shaanxi)	E. sp. (219887)
Barremian - Albian	129.4 - 100.5	China (Sichuan)	E. nobitai (217078) E. sp. (201877 201878 219971 219999)
Barremian - Albian	129.4 - 100.5	China (Guizhou)	Gigandipus chiappei (220000)
Aptian	125.0 - 113.0	China (Hebei)	Theropoda indet. (60460)
Late/Upper Aptian	122.46 - 113.0	USA (Maryland)	Brontozoum sp. (75480)
Albian - Early/Lower Cenomanian	113.0 - 93.5	United Kingdom (England)	Ornithoidichnites giganteus (232275)
Early/Lower Albian	112.03 - 109.0	USA (Texas)	Dinosauria indet. (52540) E. sp. (51421) Irenesauripus glenrosensis (57674) Irenesauripus sp. (52148) Theropoda indet. (52532 52535 52537 52538)
Early/Lower Albian	112.03 - 109.0	USA (Arkansas)	E. glenrosensis (223076)