PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Imagotaria downsi

Mammalia - Carnivora - Odobenidae

Taxonomy

Imagotaria downsi was named by Mitchell (1968) [SBMNH = Santa Barbara Museum of Natural History]. Its type specimen is SBMNH 342, a partial skull, and it is a 3D body fossil. Its type locality is Great Lakes Carbon Co. quarry, which is in a Tortonian marine diatomite in the Monterey Formation of California. It is the type species of Imagotaria.

Synonymy list

Year	Name and author
1968	Imagotaria downsi Mitchell p. 1844 figs. Fig. 1-15
1994	Imagotaria downsi Deméré p. 113
2003	Imagotaria downsi Deméré et al. p. 49 figs. Fig. 3.3
2008	Imagotaria downsi Barnes p. 531
2011	Imagotaria downsi Boessenecker p. 2
2013	Imagotaria downsi Boessenecker and Churchill p. 10 fig. 8
2015	Imagotaria downsi Boessenecker and Churchill figs. Fig. 2
2018	Imagotaria downsi Berta p. 714 figs. Fig. 3
2018	Imagotaria downsi Berta et al. p. 217 figs. Fig. 6
2018	Imagotaria downsi Magallanes et al. p. 30 figs. Figure 14
2020	Imagotaria downsi Biewer et al. figs. Figs. 4, 5

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes
class	Mammalia

Rank	Name	Author
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Ferae	()
—	Carnivoramorpha	Wyss and Flynn 1993
—	Carnivoraformes	Flynn et al.
order	Carnivora
suborder	Caniformia	Kretzoi 1943
infraorder	Canoidea	(Simpson 1931)
superfamily	Arctoidea	Flower 1869
—	Panpinnipedia	Wolsan et al. 2020
—	Pinnipedimorpha
—	Pinnipedia	()
family	Odobenidae	()
genus	Imagotaria
species	downsi

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Imagotaria downsi Mitchell 1968

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Diagnosis

Reference	Diagnosis
J. N. Biewer et al. 2020	Imagotaria downsi shares with other known odobenids a large, dorsoventrally elliptical narial opening with thick margins (c. 2[1]), dorsoventrally thick and lat- erally broad pterygoid strut (c. 13[2]), prominent antorbital process formed by frontal and maxilla (c. 15[1]), large epitympa- nic process (c. 23[1]), bony tentorium that is appressed to the pet- rosal (c. 29[1]), medial lip of distal trochlea greater in diameter than distal capitulum (c. 84[1]), and distal end of radius expanded (c. 85[1]). It can be differentiated from more basal odobenids like Pron. repenningi and Protot. primigenia, by a flattened and plate- like paroccipital process (c. 34[3]). It can be differentiated from all other more basal odobenids (i.e., basal odobenids excluding Pontolis and Os. eodon) by its shallow glenoid fossa (c. 26[1]), lack of a metaconid on its lower postcanines (c. 70[2]), and lack of a ventral tuberosity of the zygomatic root (c. 7[0]). The latter character also differentiates it from all neodobenians except G. pugnax. Imagotaria downsi can be distinguished from species of Pontolis and Os. eodon by an anterodorsally oriented mandib- ular foramen (c. 48[1]). It can be further differentiated from Pontolis by well-developed tooth enamel (c. 62[0], which also differentiates it from neodobenians), lower postcanines with rough lingual cingula (c. 67[1]), and talonid basins in the postca- nines (c. 69[0]). It can be distinguished from Os. eodon and Neo- dobenia by having a p2 with double or bilobed roots (c. 72[0]) and a double-rooted m1 (c. 79[0]). In addition to previously listed characters (c. 7, 62, 72, 79), I. downsi can be distinguished from Neodobenia by having a narrow interorbital bar (c. 17[1]),

Measurements

No measurements are available

Composition:

hydroxyapatiteuc

Ontogeny:

modification of partsuc

Environment:

marinef

Locomotion:

actively mobilec

Life habit:

amphibiousf

Diet:

carnivoref

Reproduction:

viviparousf

Created:

2011-05-21 01:16:23

Modified:

2011-05-20 10:16:23

Source: f = family, c = class, uc = unranked clade

References: Kohno et al. 1995, Hendy et al. 2009, Carreño and Cronin 1993

Age range: base of the Tortonian to the top of the Clarendonian or 11.63000 to 9.40000 Ma

Collections (19 total)

Time interval	Ma	Country or state	Original ID and collection number
Clarendonian	12.5 - 9.4	USA (California)	Imagotaria downsi (84331)
Tortonian	11.63 - 7.246	Japan (Fukushima)	Pinnipedia indet. (227081)
Tortonian	11.63 - 7.246	USA (California)	Imagotaria downsi (40514 45489 52184 52213 type locality: 58988 59156 59157 59158 59159 59160 59161 152097 152101 215597 225292) Imagotaria downsi, Imagotaria downsi (13300) Imagotaria sp. (13062)