Chorlakki (Eocene of Pakistan)

Where: North-West Frontier, Pakistan (33.6° N, 71.9° E: paleocoordinates 14.4° N, 68.1° E)

• coordinate stated in text

• outcrop-level geographic resolution

When: Kuldana Formation, Lutetian (47.8 - 41.3 Ma)

• Gingerich (2003) places this collection in the earliest Lutetian

•shown by Gunnell et al. (2008) as being close to the top of the formation, spanning at three beds, and falling just over the Early - Middle Eocene boundary; early Lutetian

• group of beds-level stratigraphic resolution

Environment/lithology: fluvial; lithified, calcareous conglomerate

• hard, calcareous conglomerate

Size class: macrofossils

Preservation: original phosphate

Collection methods: bulk, quarrying, surface (in situ), chemical, mechanical, sieve

Primary reference: P. D. Gingerich, D. E. Russell, D. Sigogneau-Russell, J.-L. Hartenberger, S. M. Ibrahim Shah, M. Hassan, K. D. Rose and R. H. Ardrey. 1979. Reconnaissance survey and vertebrate paleontology of some Paleocene and Eocene Formations in Pakistan. Contributions from the Museum of Paleontology, University of Michigan 25(5):105-116 [M. Uhen/M. Uhen/M. Uhen]more details

Purpose of describing collection: general faunal/floral analysis

PaleoDB collection 28978: authorized by Mark Uhen, entered by Mark Uhen on 08.03.2003, edited by Will Clyde and John Alroy

Creative Commons license: CC BY (attribution)

Taxonomic list

 Testudines - Carettochelyidae
Chorlakkichelys shahi1 de Broin 1987 turtle
 Testudines - Trionychidae
Trionychinae indet.1 Gray 1825 softshell turtle
 Squamata - Agamidae
Tinosaurus sp.8 Marsh 1872 squamates
 Squamata - Boidae
Erycinae indet.8 Bonaparte 1825 sand boa
Boinae indet.8 Gray 1825 boa
 Rodentia - Chapattimyidae
"Birbalomys ijlsti" = Basalomys ijlsti5 Hussain et al. 1978 rodent
Chapattimys debruijni5 Hartenberger 1982 rodent
Chapattimys wilsoni5 Hussain et al. 1978 rodent
Birbalomys sondaari5 Hussain et al. 1978 rodent
Birbalomys sondaari7 Hussain et al. 1978 rodent
Birbalomys ? woodi5 Sahni and Khare 1973 rodent
Gumbatomys asifi5 Hartenberger 1982 rodent
 Rodentia -
cf. Tamquammys sp. Shevyreva 1971 rodent
 Rodentia - Yuomyidae
"cf. Petrokoslovia sp." = Petrokozlovia5, "cf. Petrokoslovia "sp. indet. 2"" = Petrokozlovia7
"cf. Petrokoslovia sp." = Petrokozlovia5 Shevyreva 1972 rodent
"cf. Petrokoslovia "sp. indet. 2"" = Petrokozlovia7 Shevyreva 1972 rodent
 Primates - Omomyidae
Kohatius coppensi n. gen. n. sp.10
Kohatius coppensi n. gen. n. sp.10 Russell and Gingerich 1980 tarsier
 Primates - Notharctidae
cf. Agerinia sp.12, Panobius afridi n. gen. n. sp.12, Panobius amplior n. sp.6
cf. Agerinia sp.12 Crusafont-PairĂ³ and Golpe-Posse 1973 primate
Panobius afridi n. gen. n. sp.12 Russell and Gingerich 1987 primate
Panobius amplior n. sp.6 Gunnell et al. 2008 primate
specimen referred to Adapidae indet. by Russell and Gingerich (1987) and to Panobius afridi by Thewissen et al. (2001)
 Theriamorpha - Plesiosoricidae
Pakilestes lathrius11 Russell and Gingerich 1981 placental
 Theriamorpha -
Seia shahi11 Russell and Gingerich 1981 placental
 Placentalia -
Chiroptera indet.5 Blumenbach 1779 bat
 Creodonta -
Paratritemnodon indicus5 Ranga Rao 1973 creodont
 Perissodactyla - Paraceratheriidae
Forstercooperia jigniensis12 Sahni and Khare 1973 odd-toed ungulate
 Perissodactyla - Isectolophidae
Isectolophidae indet.12 Peterson 1919 odd-toed ungulate
 Perissodactyla - Brontotheriidae
? Eotitanops dayi5 Dehm and Oettingen-Spielberg 1958 brontothere
 Perissodactyla - Anthracobunidae
"Anthracobune pinfoldi" = Anthracobune pinfoldi, "Pilgrimella pilgrimi" = Anthracobune pinfoldi5, "Lammidhania wardi" = Anthracobune wardi5
"Anthracobune pinfoldi" = Anthracobune pinfoldi Pilgrim 1940 odd-toed ungulate
"Pilgrimella pilgrimi" = Anthracobune pinfoldi5 Pilgrim 1940 odd-toed ungulate
"Lammidhania wardi" = Anthracobune wardi5 Pilgrim 1940 odd-toed ungulate
 Artiodactyla - Dichobunidae
Dichobunidae indet.12 Gill 1872 even-toed ungulate
Pakibune chorlakkiensis12 Thewissen et al. 1987 even-toed ungulate
Dulcidon gandaiensis5 even-toed ungulate
Chorlakkia hassani n. gen. n. sp.4 Gingerich et al. 1979 even-toed ungulate
 Artiodactyla - Raoellidae
Khirtharia dayi Pilgrim 1940 even-toed ungulate
Indohyus cf. indriae even-toed ungulate
Indohyus major n. sp.12 Thewissen et al. 1987 even-toed ungulate
 Artiodactyla - Diacodexeidae
"Diacodexis pakistanensis" = Gujaratia pakistanensis5
"Diacodexis pakistanensis" = Gujaratia pakistanensis5 Thewissen et al. 1983 even-toed ungulate
 Cetacea - Pakicetidae
Pakicetus inachus n. gen. n. sp.3 Gingerich and Russell 1981 whale
Ichthyolestes pinfoldi13 Dehm and Oettingen-Spielberg 1958 whale
 Cimolesta -
Basalina basalensis5 Dehm and Oettingen-Spielberg 1958 tillodont
 Lepisosteiformes - Lepisosteidae
Lepisosteus sp.2 Agassiz 1843 gar
 Osteoglossiformes - Osteoglossidae
Osteoglossidae indet.2 Bonaparte 1832 arowana
 Siluriformes - Clariidae
Clariidae indet.2 Bonaparte 1846 airbreathing catfish
 Siluriformes - Bagridae
Bagridae indet.2 Bleeker 1852 naked catfish
 Siluriformes - Ariidae
Ariidae indet.2 Berg 1958 Ariid catfish
 Cypriniformes - Cyprinidae
Cyprinidae indet.2 Rafinesque 1815 carp-like fish
 Cyprinodontiformes - Cyprinodontidae
 Anabantiformes -
 Acanthomorphata - Percichthyidae
Macquaria antiquus2 Hills 1934
 Heterostropha - Planorbidae
Planorbis sp. Muller 1774 snail