Parana, Pueblo Brugo to Diamante, Ituzaingo Fm. (Neogene of Argentina)

Also known as “Conglomerado osífero” (= “Mesopotamiense”, sensu Frenguelli, 1920)

Where: Entre Rios, Argentina (31.5° S, 60.2° W: paleocoordinates 31.6° S, 58.1° W)

• coordinate estimated from map

• local area-level geographic resolution

When: Huayquerian mammal zone, Ituzaingo Formation, Huayquerian (8.0 - 5.0 Ma)

• Schmidt 2015: fossils from the base of the Ituzaingo Fm.

Environment/lithology: marine; sandstone and mudstone

Size classes: macrofossils, mesofossils

Preservation: original phosphate

Primary reference: A. L. Cione, M. M. Azpelicueta, M. Bond, A. A. Carlini, J. R. Casciotta, M. A. Cozzuol, M. Fuente, Z. Gasparini, F. J. Goin, J. Noriega, G. J. Scillato-Yane, L. Soibelzon, E. P. Tonni, D. H. Verzi, and M. G. Vucetich. 2000. Miocene vertebrates from Entre Rios province, eastern Argentina. Serie Correlacion Geologica 14:191-237 [M. Uhen/M. Uhen/O. Alfaro]more details

Purpose of describing collection: taxonomic analysis

PaleoDB collection 55600: authorized by Mark Uhen, entered by Mark Uhen on 01.11.2005, edited by Philip Mannion, Juan Carrillo, Miranta Kouvari, Evangelos Vlachos, Grace Varnham and Kateryn Pino

Creative Commons license: CC BY (attribution)

Taxonomic list

unclassified
  -
Actinopteri
 Characiformes - Cynodontidae
 Characiformes - Characidae
Mammalia
 Didelphimorphia - Didelphidae
Chironectes sp.11 Illiger 1811 water opossum
Philander entrerianus11 Ameghino 1899 four-eyed opossum
Zygolestes paranensis11 Ameghino 1898 opossum
 Sparassodonta - Thylacosmilidae
Thylacosmilus atrox11 Riggs 1933 metatherian
 Sparassodonta -
Stylocynus paranensis Mercerat 1917 metatherian
"Notictis ortizi" = Notictis ortizii Ameghino 1889 metatherian
 Cingulata -
? Trachycalyptus cingulatus Ameghino 1889 edentate
Parahoplophorus paranensis Ameghino 1883 edentate
Protoglyptodon primiformis Ameghino 1885 edentate
Palaehoplophorus antiquus Ameghino 1883 edentate
Chlamyphractus pressulus Ameghino 1885 edentate
 Cingulata - Pampatheriidae
Kraglievichia paranense, "Scirrotherium carinatum n. sp." = Kraglievichia carinatum22
Kraglievichia paranense Ameghino 1883 edentate
"Scirrotherium carinatum n. sp." = Kraglievichia carinatum22 Gois et al. 2013 edentate
 Cingulata - Panochthidae
Hoplophorus verus22 Ameghino 1889 edentate
 Cingulata - Glyptodontidae
Paraglyptodon paranensis22 Castellanos 1953 glyptodon
Comaphorus concisus Ameghino 1886 glyptodon
Eleutherocercus sp.19 Koken 1888 glyptodon
Plohophorus paranensis Ameghino 1891 glyptodon
 Cingulata -
Pseudoeuryurus lelongianus Ameghino 1889 edentate
Urotherium interundatum Ameghino 1885 edentate
"Berthawyleria sp." = Pseudoplohophorus Castellanos 1926 edentate
 Cingulata - Dasypodidae
Chasicotatus spinozai22 Scillato-Yané et al. 2010 armadillo
Macroeuphractus retusus21 Ameghino 1887 armadillo
but also ref 15016
? Zaedyus sp. Ameghino 1889 armadillo
Proeuphractus limpidus Ameghino 1886 armadillo
Dasypus neogaeus Ameghino 1891 armadillo
 Megatherioidea - Megalonychidae
Protomegalonyx doellojuradoi, Protomegalonyx praecursor, "Ortotherium laticurvatum" = Ortotherium laticurvatum, "Amphiocnus seneus" = Ortotherium laticurvatum, "Ortotherium scrofum" = Ortotherium laticurvatum, "Ortotherium schlosseri" = Ortotherium laticurvatum, Ortotherium robustum, "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum, Torcellia paranense, Amphiocnus paranense, Megalonychops primigenius, Pliomorphus robustus, "Pliomorphus mutilatus" = Pliomorphus mutilatus, "Pliomorphus brevis" = Pliomorphus mutilatus, "Pliomorphus ameghinoi" = Pliomorphus mutilatus
Protomegalonyx doellojuradoi Kraglievich 1925 edentate
Protomegalonyx praecursor Kraglievich 1925 edentate
"Ortotherium laticurvatum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Amphiocnus seneus" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium scrofum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium schlosseri" = Ortotherium laticurvatum Ameghino 1885 edentate
Ortotherium robustum Ameghino 1881 edentate
"Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum Bordas 1941 edentate
Holotype: MACN Pv 13656, type of Ortotherium brevirostrum Bordas, 1942
Torcellia paranense Kraglievich 1923 edentate
Amphiocnus paranense Kraglievich 1922 edentate
Megalonychops primigenius Kraglievich 1926 edentate
Pliomorphus robustus Ameghino 1885 edentate
"Pliomorphus mutilatus" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus brevis" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus ameghinoi" = Pliomorphus mutilatus Ameghino 1885 edentate
 Megatherioidea - Nothrotheriidae
Pronothrotherium mirabilis Kraglievich 1925 edentate
Neohapalops rothi Kraglievich 1923 edentate
 Megatherioidea - Megatheriidae
Promegatherium smaltatum Ameghino 1883 edentate
Promegatherium parvulum Ameghino 1891 edentate
Megatherium antiquum3 Ameghino 1885 edentate
Pyramiodontherium sp.4 Rovereto 1914 edentate
"Promegatherium remulsum" = Pyramiodontherium remulsum3 Ameghino 1886 edentate
"Pliomegatherium lelongi" = Pliomegatherium lelongi Kraglievich 1930 edentate
"Pliomegatherium paranensis" = Pliomegatherium lelongi3 Kraglievich 1930 edentate
Eomegatherium nanum4 Burmeister 1891 edentate
  - Mylodontidae
Prolestodon paranensis Ameghino 1889 edentate
Prolestodon antiquus Ameghino 1885 edentate
Ranculcus scalabrinianus Ameghino 1891 edentate
Promylodon paranensis Ameghino 1883 edentate
Megabradys darwini Scillato-Yané 1981 edentate
Strabosodon obtusicavus Ameghino 1891 edentate
Strabosodon acuticavus Ameghino 1891 edentate
Scelidotheriinae indet. Ameghino 1889 edentate
Octomylodon aversus Ameghino 1904 edentate
"Sphenotherus paranensis" = Sphenotherus zavaletianus Ameghino 1891 edentate
 Carnivora - Procyonidae
Cyonasua argentina Ameghino 1885 procyonid carnivore
 Panameriungulata - Macraucheniidae
Paranauchenia denticulata Ameghino 1891 placental
Scalabrinitherium rothii Ameghino 1885 placental
Scalabrinitherium bravardi Ameghino 1883 placental
Promacrauchenia antiqua Ameghino 1887 placental
"Oxyodontherium zeballosi" = Oxyodontherium zeballozi Ameghino 1883 placental
"Mesorhinus piramydatus" = Oxyodontherium piramydatus Ameghino 1885 placental
Cullinia sp.20 Cabrera and Kraglievich 1931 placental
 Panameriungulata - Proterotheriidae
"Coelosoma eversa" = Diadiaphorus eversus, Proterotherium cervioides, "Brachytherium cuspidatum" = Brachytherium cuspidatum, "Licaphrium mesopotamiense" = Brachytherium cuspidatum, "Epitherium paranensis" = Brachytherium cuspidatum, "Proterotherium gradatum" = Brachytherium gradatum, Neobrachytherium ameghinoi20
"Coelosoma eversa" = Diadiaphorus eversus Soria 2001 placental
Proterotherium cervioides Ameghino 1883 placental
"Brachytherium cuspidatum" = Brachytherium cuspidatum Ameghino 1883 placental
"Licaphrium mesopotamiense" = Brachytherium cuspidatum Ameghino 1883 placental
"Epitherium paranensis" = Brachytherium cuspidatum Ameghino 1883 placental
"Proterotherium gradatum" = Brachytherium gradatum Ameghino 1891 placental
Neobrachytherium ameghinoi20 Soria 2001 placental
 Cetacea - Iniidae
Iniidae indet.12, "Saurodelphis gigas n. sp." = Saurocetes gigas8
Iniidae indet.12 Gray 1846 toothed whale
"Saurodelphis gigas n. sp." = Saurocetes gigas8 Cozzuol 1988 toothed whale
 Rodentia - Hydrochoeridae
Cantracavia sp.6, Contracavia sp., Anatochoerus sp., "Protohydrochoerus sp." = Phugatherium6, Phugatherium cataclisticum, "Anchimysops sp." = Phugatherium, "Cardiatherium sp." = Cardiatherium, "Anchimys sp." = Cardiatherium, "Procardiatherium sp." = Cardiatherium, "Kiyutherium sp." = Cardiatherium, "Plexochoerus sp." = Cardiatherium, Cardiatherium paranense n. sp.17
Cantracavia sp.6 Burmeister 1885 caviomorph
Contracavia sp. Burmeister 1885 caviomorph
Anatochoerus sp. Vucetich and Mones 1991 caviomorph
"Protohydrochoerus sp." = Phugatherium6 Ameghino 1887 caviomorph
Phugatherium cataclisticum Ameghino 1887 caviomorph
"Anchimysops sp." = Phugatherium Ameghino 1887 caviomorph
"Cardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Anchimys sp." = Cardiatherium Ameghino 1883 caviomorph
"Procardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Kiyutherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Plexochoerus sp." = Cardiatherium Ameghino 1883 caviomorph
Cardiatherium paranense n. sp.17 Ameghino 1883 caviomorph
 Rodentia - Caviidae
Cardiomys sp. Ameghino 1885 caviomorph
Caviodon sp. Ameghino 1885 caviomorph
Prodolichotis sp. Kraglievich 1932 caviomorph
Palaeocavia sp. Ameghino 1889 caviomorph
Parodimys sp. Kraglievich 1932 caviomorph
Pliodolichotis sp.5 Kraglievich 1927 mara
 Rodentia - Erethizontidae
Paradoxomys sp. Ameghino 1885 porcupine
Paradoxomys cancrivorus6 Ameghino 1885 porcupine
 Rodentia - Chinchillidae
"Lagostomopsis sp." = Lagostomus (Lagostomopsis), "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)6, "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)6
"Lagostomopsis sp." = Lagostomus (Lagostomopsis) Kraglievich 1926 viscacha
"Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)6 viscacha
"Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)6 viscacha
 Rodentia - Dinomyidae
Potamarchus sp. Burmeister 1885 caviomorph
Potamarchus murinus6 Burmeister 1885 caviomorph
Pentastylomys sp. Kraglievich 1926 caviomorph
Tetrastylus atavus6 caviomorph
T. (Protelicomys) atavus
Tetrastylus laevigatus6 Ameghino 1886 caviomorph
Tetrastylus laevigatus n. sp.23 Ameghino 1886 caviomorph
Isostylomys sp. Kraglievich 1926 caviomorph
Isostylomys laurillardi6 Ameghino 1883 caviomorph
Pseudosigmomys sp. Kraglievich 1931 caviomorph
Rusconia crassidens6 Kraglievich 1931 caviomorph
Protomegamys sp. Kraglievich 1932 caviomorph
Colpostemma sp. Ameghino 1891 caviomorph
Diaphoromys sp. Kraglievich 1931 caviomorph
Diaphoromys mesopotamicus6 Kraglievich 1932 caviomorph
Diaphoromys gamayensis6 Kraglievich 1931 caviomorph
Olenopsis typicus6 Ameghino 1889 caviomorph
Eumegamysops sp. Alvarez 1947 caviomorph
Eumegamysops praependens6 Ameghino 1886 caviomorph
Telodontomys compressidens6 Kraglievich 1931 caviomorph
Carlesia sp. Kraglievich 1926 caviomorph
Carlesia pendolai6 caviomorph
Eumegamys sp. Kraglievich 1926 caviomorph
Eumegamys dubius6 caviomorph
Eumegamys depressidens6 caviomorph
"Eumegamys" depressidens
Eumegamys contortus6 Ameghino 1889 caviomorph
Eumegamys scalabrinianus6 Kraglievich 1926 caviomorph
Eumegamys paranensis6 Kraglievich 1926 caviomorph
Briaromys sp. Ameghino 1889 caviomorph
Briaromys trouessartianus6 Ameghino 1889 caviomorph
Doellomys sp. Alvarez 1947 caviomorph
Doellomys parcus6 caviomorph
Gyriabrus sp. Ameghino 1891 caviomorph
Gyriabrus indivisus6 caviomorph
Gyriabrus holmbergi6 Ameghino 1885 caviomorph
Discolomys cuneus6 Ameghino 1889 caviomorph
"Discolomys"
Paranamys sp. Kraglievich 1934 caviomorph
"Drytomomys typicus" = Paranamys typicus7 Ameghino 1889 caviomorph
 Rodentia - Neoepiblemidae
Phoberomys sp., Phoberomys sp.6, "Phoberomys burmeisteri" = Phoberomys burmeisteri14, "Megamys burmeisteri" = Phoberomys burmeisteri13, "Phoberomys minima n. sp." = Phoberomys burmeisteri16, "Phoberomys minima" = Phoberomys burmeisteri6, "Phoberomys praecursor n. sp." = Phoberomys burmeisteri15, "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri16, "Phoberomys insolita n. sp." = Phoberomys burmeisteri16, Perimys scalabrinianus6, Neoepiblema sp., Epiblema horridula13, "Neoepiblema ambrosettianus" = Neoepiblema horridula18
Phoberomys sp. Kraglievich 1926 caviomorph
Phoberomys sp.6 Kraglievich 1926 caviomorph
"Phoberomys burmeisteri" = Phoberomys burmeisteri14 Kraglievich 1926 caviomorph
"Megamys burmeisteri" = Phoberomys burmeisteri13 Kraglievich 1926 caviomorph
collected by F. Ameghino 1886
"Phoberomys minima n. sp." = Phoberomys burmeisteri16 Kraglievich 1926 caviomorph
"Phoberomys minima" = Phoberomys burmeisteri6 Kraglievich 1926 caviomorph
"Phoberomys praecursor n. sp." = Phoberomys burmeisteri15 Kraglievich 1926 caviomorph
"Phoberomys lozanoi n. sp." = Phoberomys burmeisteri16 Kraglievich 1926 caviomorph
"Phoberomys insolita n. sp." = Phoberomys burmeisteri16 Kraglievich 1926 caviomorph
Perimys scalabrinianus6 Ameghino 1889 caviomorph
Neoepiblema sp. Ameghino 1889 caviomorph
Epiblema horridula13 Ameghino 1886 caviomorph
upper right molar - collected by F. Ameghino 1886
"Neoepiblema ambrosettianus" = Neoepiblema horridula18 Ameghino 1886 caviomorph
 Rodentia - Abrocomidae
Protabrocoma sp. Kraglievich 1927 chinchilla rat
Protabrocoma paranensis6 chinchilla rat
 Rodentia - Myocastoridae
Strophostephanos sp.5 Ameghino 1891 caviomorph
 Rodentia - Echimyidae
Myocastor sp. Kerr 1782 spiny rat
Myocastor obesus6 spiny rat
Myocastor sinuata6 spiny rat
Haplostropha sp. Ameghino 1891 spiny rat
Eumysops parodii Kraglievich 1945 spiny rat
Eumysops parodii6 Kraglievich 1945 spiny rat
“Eumysops” parodii
 Notoungulata - Interatheriidae
Munizia paranensis Kraglievich 1931 notoungulate
Protypotherium antiquum Ameghino 1882 notoungulate
 Notoungulata - Mesotheriidae
Eutypotherium sp.20 Roth 1902 notoungulate
 Notoungulata - Haplodontheriidae
Pachynodon modicus, "Haplodontherium wildei" = Haplodontherium wildei, "Toxodontherium reverendum" = Haplodontherium wildei, "Haplodontherium limun" = Haplodontherium wildei, "Toxodontherium compressum" = Haplodontherium wildei
Pachynodon modicus Burmeister 1891 notoungulate
"Haplodontherium wildei" = Haplodontherium wildei Ameghino 1885 notoungulate
"Toxodontherium reverendum" = Haplodontherium wildei Ameghino 1885 notoungulate
"Haplodontherium limun" = Haplodontherium wildei Ameghino 1885 notoungulate
"Toxodontherium compressum" = Haplodontherium wildei Ameghino 1885 notoungulate
 Notoungulata - Toxodontidae
"Dinotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate
"Palaeotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate
Dilobodon lutarius Ameghino 1886 notoungulate
Xotodon foricurvatus Ameghino 1885 notoungulate
Xotodon doellojuradi Frenguelli 1920 notoungulate
? Berroia sp. Kraglievich 1931 notoungulate
"Stenotephanos plicidens" = Stenotephanos plicidens Ameghino 1885 notoungulate
"? Palaeotoxodon virgatus" = Stenotephanos plicidens Ameghino 1885 notoungulate
"Eutomodus elautus" = Stenotephanos plicidens Ameghino 1885 notoungulate
? Palaeotoxodon protoburmeisteri Ameghino 1887 notoungulate
Reptilia
 Testudines - Chelidae
Phrynops cf. geoffroanus Schweigger 1812 side-necked turtle of Geoffroy
 Testudines - Cheloniidae
Cheloniidae indet.9 Bonaparte 1832 sea turtle
MACN Pv 14224, nearly complete right mandible
Aves
 Gruiformes - Phorusrhacidae
"Phororhacidae indet." = Phorusrhacidae, Andalgalornis sp., "? Onactornis sp." = Devincenzia
"Phororhacidae indet." = Phorusrhacidae Ameghino 1889 terror bird
Andalgalornis sp. Patterson and Kraglievich 1960 terror bird
"? Onactornis sp." = Devincenzia Kraglievich 1932 terror bird
 Gruiformes - Rallidae
Rallidae indet. Vigors 1825 rail
 Phoenicopteriformes - Phoenicopteridae
Phoenicopteridae indet. Bonaparte 1838 flamingo
 Phoenicopteriformes - Palaelodidae
Megapaloelodus sp. Miller 1944 swimming flamingo
 Rheiformes - Rheidae
Rheidae indet. Bonaparte 1849 rhea
Reptilia
 Loricata -
Crocodylia indet. crocodilian
 Crocodylia - Gavialidae
Gryposuchus neogaeus Rusconi 1933 crocodilian
 Crocodylia - Alligatoridae
Caiman jacare, Caiman cf. yacare1, Caiman gasparinae n. sp.2, Caiman australis, "Caiman latirostris" = Caiman latirostris, "Caiman lutescens n. sp." = Caiman latirostris1, "Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi10
Caiman cf. yacare1 Daudin 1802 caiman
MLP 73-IV-15-5, MLP 73-IV-15-6, MLP 73-IV-5-17, MACN PV 5417
Caiman gasparinae n. sp.2 Bona and Carabajal 2013 caiman
MLP-73-IV-15-1 - holotype (skull fragments represented by a rostrum with articulated fragments of premaxillae, maxillae, nasals, left lacrymal and a partial braincase lacking the basicranium); referred material - MACN PV 5555 (fragment of right premaxilla)
Caiman australis Bravard 1858 caiman
"Caiman latirostris" = Caiman latirostris Daudin 1801 broud-snouted caiman
"Caiman lutescens n. sp." = Caiman latirostris1 Daudin 1801 broud-snouted caiman
MACN-PV 13551
"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi10 Bocquentin Villanueva 1984 crocodilian
MLP 73-IV-15-8 - holotype (posterior portion of skull); hypodigm: MLP 73-IV-15-9 (posterior portion of skull)
 Squamata -