Also known as “Conglomerado osífero” (= “Mesopotamiense”, sensu Frenguelli, 1920)
Where: Entre Rios, Argentina (31.5° S, 60.2° W: paleocoordinates 31.6° S, 58.1° W)
• coordinate estimated from map
• local area-level geographic resolution
When: Huayquerian mammal zone, Ituzaingo Formation, Huayquerian (8.0 - 5.0 Ma)
• Schmidt 2015: fossils from the base of the Ituzaingo Fm.
Environment/lithology: marine; sandstone and mudstone
Size classes: macrofossils, mesofossils
Preservation: original phosphate
Primary reference: A. L. Cione, M. M. Azpelicueta, M. Bond, A. A. Carlini, J. R. Casciotta, M. A. Cozzuol, M. Fuente, Z. Gasparini, F. J. Goin, J. Noriega, G. J. Scillato-Yane, L. Soibelzon, E. P. Tonni, D. H. Verzi, and M. G. Vucetich. 2000. Miocene vertebrates from Entre Rios province, eastern Argentina. Serie Correlacion Geologica 14:191-237 [M. Uhen/M. Uhen/O. Alfaro]more details
Purpose of describing collection: taxonomic analysis
PaleoDB collection 55600: authorized by Mark Uhen, entered by Mark Uhen on 01.11.2005, edited by Philip Mannion, Juan Carrillo, Miranta Kouvari, Evangelos Vlachos, Grace Varnham and Kateryn Pino
Creative Commons license: CC BY (attribution)
Taxonomic list
unclassified | |
Actinopteri | |
Mammalia | |
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Cullinia sp.21, Paranauchenia denticulata, "Mesorhinus piramydatus" = Oxyodontherium piramydatus, "Oxyodontherium zeballosi" = Oxyodontherium zeballozi, Promacrauchenia antiqua, Scalabrinitherium rothii, Scalabrinitherium bravardi
Paranauchenia denticulata Ameghino 1891 placental
"Mesorhinus piramydatus" = Oxyodontherium piramydatus Ameghino 1885 placental
"Oxyodontherium zeballosi" = Oxyodontherium zeballozi Ameghino 1883 placental
Promacrauchenia antiqua Ameghino 1887 placental
Scalabrinitherium rothii Ameghino 1885 placental
Scalabrinitherium bravardi Ameghino 1883 placental | |
Neobrachytherium ameghinoi21, "Proterotherium gradatum" = Brachytherium gradatum, "Brachytherium cuspidatum" = Brachytherium cuspidatum, "Epitherium paranensis" = Brachytherium cuspidatum, "Licaphrium mesopotamiense" = Brachytherium cuspidatum, "Coelosoma eversa" = Diadiaphorus eversus, Proterotherium cervioides
"Proterotherium gradatum" = Brachytherium gradatum Ameghino 1891 placental
"Brachytherium cuspidatum" = Brachytherium cuspidatum Ameghino 1883 placental
"Epitherium paranensis" = Brachytherium cuspidatum Ameghino 1883 placental
"Licaphrium mesopotamiense" = Brachytherium cuspidatum Ameghino 1883 placental
"Coelosoma eversa" = Diadiaphorus eversus Soria 2001 placental
Proterotherium cervioides Ameghino 1883 placental | |
Cyonasua argentina Ameghino 1885 procyonid carnivore | |
Paradoxomys sp. Ameghino 1885 porcupine | |
Perimys scalabrinianus7, Neoepiblema sp., "Neoepiblema ambrosettianus" = Neoepiblema horridula19, Epiblema horridula14, Phoberomys sp., Phoberomys sp.7, "Phoberomys burmeisteri" = Phoberomys burmeisteri15, "Phoberomys minima n. sp." = Phoberomys burmeisteri17, "Phoberomys minima" = Phoberomys burmeisteri7, "Megamys burmeisteri" = Phoberomys burmeisteri14, "Phoberomys praecursor n. sp." = Phoberomys burmeisteri16, "Phoberomys insolita n. sp." = Phoberomys burmeisteri17, "Phoberomys lozanoi n. sp." = Phoberomys burmeisteri17
Neoepiblema sp. Ameghino 1889 caviomorph
Phoberomys sp. Kraglievich 1926 caviomorph
"Megamys burmeisteri" = Phoberomys burmeisteri14 Kraglievich 1926 caviomorph collected by F. Ameghino 1886
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Telodontomys compressidens7, Pentastylomys sp., Pentastylomys seriei7, Carlesia sp., Carlesia pendolai7, Olenopsis typicus7, Potamarchus sp., Potamarchus sigmodon7, Potamarchus murinus7, Pseudosigmomys sp., Pseudosigmomys paranensis7, Rusconia crassidens7, Gyriabrus sp., Gyriabrus glutinatus7, Gyriabrus indivisus7, Gyriabrus rebagliattii7, Gyriabrus holmbergi7, Diaphoromys sp., Diaphoromys compressidens7, Diaphoromys mesopotamicus7, Diaphoromys gamayensis7, Paranamys sp., "Drytomomys typicus" = Paranamys typicus8, Protomegamys sp., Protomegamys coligatus7, Briaromys sp., Briaromys trouessartianus7, Eumegamysops sp., Eumegamysops praependens7, Colpostemma sp., Discolomys cuneus7, Tetrastylomys castellanosi7, Doellomys sp., Doellomys parcus7, Eumegamys sp., Eumegamys dubius7, Eumegamys depressidens7, Eumegamys contortus7, Eumegamys paranensis7, Eumegamys scalabrinianus7, Pentastylodon racedi7, Isostylomys sp., Isostylomys ameghinoi7, Isostylomys laurillardi7, Tetrastylus aguilari7, Tetrastylus diffusus7, Tetrastylus robustus7, Tetrastylus atavus7, Tetrastylus laevigatus7, Tetrastylus laevigatus n. sp.24
Pentastylomys sp. Kraglievich 1926 caviomorph
Carlesia sp. Kraglievich 1926 caviomorph
Potamarchus sp. Burmeister 1885 caviomorph
Pseudosigmomys sp. Kraglievich 1931 caviomorph
Gyriabrus sp. Ameghino 1891 caviomorph
Diaphoromys sp. Kraglievich 1931 caviomorph
Paranamys sp. Kraglievich 1934 caviomorph
Protomegamys sp. Kraglievich 1932 caviomorph
Briaromys sp. Ameghino 1889 caviomorph
Eumegamysops sp. Alvarez 1947 caviomorph
Colpostemma sp. Ameghino 1891 caviomorph
Doellomys sp. Alvarez 1947 caviomorph
Eumegamys sp. Kraglievich 1926 caviomorph
Isostylomys sp. Kraglievich 1926 caviomorph | |
"Lagostomopsis sp." = Lagostomus (Lagostomopsis), "Lagostomopsis pallidens" = Lagostomus (Lagostomopsis)7, "Lagostomopsis antiquus" = Lagostomus (Lagostomopsis)7
"Lagostomopsis sp." = Lagostomus (Lagostomopsis) Kraglievich 1926 viscacha | |
Contracavia sp., "Anchimysops sp." = Phugatherium, "Protohydrochoerus sp." = Phugatherium7, Phugatherium cataclisticum, "Cardiatherium sp." = Cardiatherium, "Kiyutherium sp." = Cardiatherium, "Procardiatherium sp." = Cardiatherium, "Anchimys sp." = Cardiatherium, "Plexochoerus sp." = Cardiatherium, Cardiatherium paranense n. sp.18, Anatochoerus sp., Cantracavia sp.7
Contracavia sp. Burmeister 1885 caviomorph
"Anchimysops sp." = Phugatherium Ameghino 1887 caviomorph
Phugatherium cataclisticum Ameghino 1887 caviomorph
"Cardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Kiyutherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Procardiatherium sp." = Cardiatherium Ameghino 1883 caviomorph
"Anchimys sp." = Cardiatherium Ameghino 1883 caviomorph
"Plexochoerus sp." = Cardiatherium Ameghino 1883 caviomorph
Anatochoerus sp. Vucetich and Mones 1991 caviomorph | |
Prodolichotis sp. Kraglievich 1932 caviomorph
Cardiomys sp. Ameghino 1885 caviomorph
Caviodon sp. Ameghino 1885 caviomorph
Parodimys sp. Kraglievich 1932 caviomorph
Palaeocavia sp. Ameghino 1889 caviomorph | |
Myocastor sp., Myocastor paranensis7, Myocastor obesus7, Myocastor sinuata7, Haplostropha sp., Haplostropha sacabriniana7, Eumysops parodii, Eumysops parodii7
Myocastor sp. Kerr 1782 spiny rat
Haplostropha sp. Ameghino 1891 spiny rat
Eumysops parodii Kraglievich 1945 spiny rat | |
Protabrocoma sp. Kraglievich 1927 chinchilla rat | |
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Munizia paranensis Kraglievich 1931 notoungulate
Protypotherium antiquum Ameghino 1882 notoungulate | |
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"Haplodontherium wildei" = Haplodontherium wildei, "Haplodontherium limun" = Haplodontherium wildei, "Toxodontherium reverendum" = Haplodontherium wildei, "Toxodontherium compressum" = Haplodontherium wildei, Pachynodon modicus
"Haplodontherium wildei" = Haplodontherium wildei Ameghino 1885 notoungulate
"Haplodontherium limun" = Haplodontherium wildei Ameghino 1885 notoungulate
"Toxodontherium reverendum" = Haplodontherium wildei Ameghino 1885 notoungulate
"Toxodontherium compressum" = Haplodontherium wildei Ameghino 1885 notoungulate
Pachynodon modicus Burmeister 1891 notoungulate | |
Xotodon foricurvatus, Xotodon doellojuradi, "Stenotephanos plicidens" = Stenotephanos plicidens, "Eutomodus elautus" = Stenotephanos plicidens, "? Palaeotoxodon virgatus" = Stenotephanos plicidens, ? Palaeotoxodon protoburmeisteri, ? Adinotherium paranense, ? Berroia sp., "Dinotoxodon paranensis" = Dinotoxodon paranensis, "Palaeotoxodon paranensis" = Dinotoxodon paranensis, Dilobodon lutarius
Xotodon foricurvatus Ameghino 1885 notoungulate
Xotodon doellojuradi Frenguelli 1920 notoungulate
"Stenotephanos plicidens" = Stenotephanos plicidens Ameghino 1885 notoungulate
"Eutomodus elautus" = Stenotephanos plicidens Ameghino 1885 notoungulate
"? Palaeotoxodon virgatus" = Stenotephanos plicidens Ameghino 1885 notoungulate
? Palaeotoxodon protoburmeisteri Ameghino 1887 notoungulate
? Adinotherium paranense notoungulate
? Berroia sp. Kraglievich 1931 notoungulate
"Dinotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate
"Palaeotoxodon paranensis" = Dinotoxodon paranensis Burmeister 1885 notoungulate
Dilobodon lutarius Ameghino 1886 notoungulate | |
Pronothrotherium mirabilis Kraglievich 1925 edentate
Neohapalops rothi Kraglievich 1923 edentate | |
Promegatherium smaltatum, Promegatherium parvulum, Pyramiodontherium sp.5, "Promegatherium remulsum" = Pyramiodontherium remulsum4, "Pliomegatherium lelongi" = Pliomegatherium lelongi, "Pliomegatherium paranensis" = Pliomegatherium lelongi4, Megatherium antiquum4, Eomegatherium nanum5
Promegatherium smaltatum Ameghino 1883 edentate
Promegatherium parvulum Ameghino 1891 edentate
"Pliomegatherium lelongi" = Pliomegatherium lelongi Kraglievich 1930 edentate | |
Protomegalonyx praecursor, Protomegalonyx doellojuradoi, "Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum, Torcellia paranense, Pliomorphus robustus, "Pliomorphus mutilatus" = Pliomorphus mutilatus, "Pliomorphus ameghinoi" = Pliomorphus mutilatus, "Pliomorphus brevis" = Pliomorphus mutilatus, Megalonychops primigenius, Ortotherium robustum, "Ortotherium laticurvatum" = Ortotherium laticurvatum, "Ortotherium scrofum" = Ortotherium laticurvatum, "Ortotherium schlosseri" = Ortotherium laticurvatum, "Amphiocnus seneus" = Ortotherium laticurvatum, Amphiocnus paranense
Protomegalonyx praecursor Kraglievich 1925 edentate
Protomegalonyx doellojuradoi Kraglievich 1925 edentate
"Ortotherium brevirostrum n. sp." = Mesopotamocnus brevirostrum Bordas 1941 edentate Holotype: MACN Pv 13656, type of Ortotherium brevirostrum Bordas, 1942
Torcellia paranense Kraglievich 1923 edentate
Pliomorphus robustus Ameghino 1885 edentate
"Pliomorphus mutilatus" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus ameghinoi" = Pliomorphus mutilatus Ameghino 1885 edentate
"Pliomorphus brevis" = Pliomorphus mutilatus Ameghino 1885 edentate
Megalonychops primigenius Kraglievich 1926 edentate
Ortotherium robustum Ameghino 1881 edentate
"Ortotherium laticurvatum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium scrofum" = Ortotherium laticurvatum Ameghino 1885 edentate
"Ortotherium schlosseri" = Ortotherium laticurvatum Ameghino 1885 edentate
"Amphiocnus seneus" = Ortotherium laticurvatum Ameghino 1885 edentate
Amphiocnus paranense Kraglievich 1922 edentate | |
Megabradys darwini, Promylodon paranensis, Ranculcus scalabrinianus, Prolestodon paranensis, Prolestodon antiquus, Strabosodon acuticavus, Strabosodon obtusicavus, Scelidotheriinae indet., "Sphenotherus paranensis" = Sphenotherus zavaletianus, Octomylodon aversus
Megabradys darwini Scillato-Yané 1981 edentate
Promylodon paranensis Ameghino 1883 edentate
Ranculcus scalabrinianus Ameghino 1891 edentate
Prolestodon paranensis Ameghino 1889 edentate
Prolestodon antiquus Ameghino 1885 edentate
Strabosodon acuticavus Ameghino 1891 edentate
Strabosodon obtusicavus Ameghino 1891 edentate
Scelidotheriinae indet. Ameghino 1889 edentate
"Sphenotherus paranensis" = Sphenotherus zavaletianus Ameghino 1891 edentate
Octomylodon aversus Ameghino 1904 edentate | |
Dasypus neogaeus, Chasicotatus spinozai23, Proeuphractus limpidus, ? Zaedyus sp., Macroeuphractus retusus22
Dasypus neogaeus Ameghino 1891 armadillo
Proeuphractus limpidus Ameghino 1886 armadillo
? Zaedyus sp. Ameghino 1889 armadillo | |
? Trachycalyptus cingulatus, Urotherium interundatum, Pseudoeuryurus lelongianus, "Berthawyleria sp." = Pseudoplohophorus, Protoglyptodon primiformis, Palaehoplophorus antiquus, Chlamyphractus pressulus
? Trachycalyptus cingulatus Ameghino 1889 edentate
Urotherium interundatum Ameghino 1885 edentate
Pseudoeuryurus lelongianus Ameghino 1889 edentate
"Berthawyleria sp." = Pseudoplohophorus Castellanos 1926 edentate
Protoglyptodon primiformis Ameghino 1885 edentate
Palaehoplophorus antiquus Ameghino 1883 edentate
Chlamyphractus pressulus Ameghino 1885 edentate | |
Kraglievichia paranense Ameghino 1883 edentate | |
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Comaphorus concisus Ameghino 1886 glyptodon
Plohophorus paranensis Ameghino 1891 glyptodon | |
Parahoplophorus paranensis Ameghino 1883 edentate | |
"Notictis ortizi" = Notictis ortizii, Stylocynus paranensis
"Notictis ortizi" = Notictis ortizii Ameghino 1889 metatherian
Stylocynus paranensis Mercerat 1917 metatherian | |
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Reptilia | |
Phrynops cf. geoffroanus Schweigger 1812 side-necked turtle of Geoffroy | |
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Crocodylia indet. crocodilian | |
Gryposuchus neogaeus Rusconi 1933 crocodilian | |
"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi11, "Caiman gasparinae n. sp." = Paranasuchus gasparinae3, Caiman jacare, Caiman australis, Caiman latirostris, Caiman cf. yacare1, Paranacaiman bravardi n. gen. n. sp.2
"Carandaisuchus nativus n. gen. n. sp." = Mourasuchus arendsi11 Bocquentin Villanueva 1984 crocodilian MLP 73-IV-15-8 - holotype (posterior portion of skull); hypodigm: MLP 73-IV-15-9 (posterior portion of skull)
"Caiman gasparinae n. sp." = Paranasuchus gasparinae3 Bona and Carabajal 2013 crocodilian MLP-73-IV-15-1 - holotype (skull fragments represented by a rostrum with articulated fragments of premaxillae, maxillae, nasals, left lacrymal and a partial braincase lacking the basicranium); referred material - MACN PV 5555 (fragment of right premaxilla)
Caiman jacare caiman
Caiman australis Bravard 1858 caiman
Caiman latirostris Daudin 1801 broud-snouted caiman | |
Aves | |
Rallidae indet. Vigors 1825 rail | |
"Phororhacidae indet." = Phorusrhacidae Ameghino 1889 terror bird
"? Onactornis sp." = Devincenzia Kraglievich 1932 terror bird
Andalgalornis sp. Patterson and Kraglievich 1960 terror bird | |
Rheidae indet. Bonaparte 1849 rhea | |
Megapaloelodus sp. Miller 1944 swimming flamingo | |
Phoenicopteridae indet. Bonaparte 1838 flamingo | |
Reptilia | |