Parent taxon: Gymnospermae according to R. Kräusel 1939
See also Srivastava 1991
Sister taxa: Condylites, Masculostrobus, Palaeocyparis, Pinopsida, Podocarpopsida, Satyrites incertus, Taxopsida, Voltziopsida, Uesuguipollenites, Birsinghpuria, Tanaidocarpidium, Alatocarpus, Barakaroxylon, Savitrispermum, Sertostrobus, Stephanostoma, Tanaidia, Entylissa, Polyloboxylon, Belemnopteris, Archaeosperma, Zalesskioxylon, Kryshtofovichiella, Nandorioxylon, Otofeistia, Cordaitopsida, Arberiella, Prototaxoxylon, Parapalaeoxylon, Carpodium, Taxopitys, Taxineae, Rotundaspermum, Pakuriospermum, Surangephyllum, Scirroma, Triangulospermum, Palaeocarpus, Voltziostrobus, Damudoxylon, Maheshwariella, Bulbospermum, Cupressoconus, Taxaliospermum, Bennettitales, Kamthioxylon, Prototaxopitys, Palispermum, Paradoxospermum, Coniferales, Platycardia, Cornucarpus, Cardiocarpum, Stereocarpus, Araucariales, Megaporoxylon, Niponophyllum, Collospermum, Pyriformispermum, Sujfunophyllum, Retortistoma, Sclerospiroxylon, Aethophyllum, Polysolenoxylon, Shivacarpus, Indoxylon, Nummulospermum, Sonajorispermum, Nidispermum, Trigonomyelon, Gondwanophyllites, Kaokoxylon, Pantiaspermum
Subtaxa: Coniferoovulites Protophyllocladoxylon
Distribution:
• Miocene of Switzerland (1 collection)
• Oligocene of the Russian Federation (1)
• Eocene of Antarctica (2), the United Kingdom (1)
• Paleogene of India (1)
• Cretaceous of Germany (1), Niger (1), United States (3: Wyoming)
• Jurassic of the Russian Federation (2)
• Permian of India (2)
• Carboniferous of United States (2: New Mexico)
Total: 17 collections each including a single occurrence