Parent taxon: Trilobita according to J. D. Beech and J. C. Lamsdell 2021
Sister taxa: Aglagalia, Agraulaeva, Aguaraya, Aldonaiidae, Altajaharpes, Amechilus, Amgaspididae, Ampellatocephalina, Anacopodia, Antagmidae, Antatlaeva, Atopidae, Auritamidae, Avoninidae, Batchatiella, Bathynotidae, Beikedamaspis, Bergerolimbus, Burlingiidae, Carmon, Changshaniidae, Chinensis, Corynexochida, Crossoura, Curiaspis, Cuyanaspis, Cyclolenzellina, Dalmanites troosti, Deltacare, Devonocoryphe, Edganocare, Elleria, Elleria morani, Ellipsocephaloididae, Ellipsuella, Entomostracites, Erbenochile, Eulomella, Eutrilobita, Fabulaspis, Faciura, Gentilaspis, Gigmpeltis, Goycoia, Helioproetoides, Holocephalinidae, Howelluella, Idiorapha, Ignotogregatidae, Indigestus, Irvingelloides, Ishpella, Isocolidae, Issafenia, Ityophoridae, Jamrogiidae, Jonotus, Juriietella, Kamlacus, Kanfuqingia, Leptopilus, Libristoma, Lichida, Llanoaspididae, Loxopeltis, Lusampa, Mapaniidae, Megalaspidicus, Meneghinella, Metapopristis, Monkaspididae, Moolatrilo, Moretaeva, Moretarva, Namanoiidae, Neochilonorria, Neomalungia, Neotaenicephalus, Nepeidae, Odontopleurida, Ogygiocaridinae, Olenida, Onchonotopsidae, Opisthoparia, Orictocara, Otekmaspis, Otenaspella, Paichiashania, Paleoaethia, Parakingaspidoides, Pararania, Peculicephalina, Perissopyge, Phacopida, Piliolites, Pliomerellus, Poculicephalina, Polycyrtaspididae, Prodamesella, Proetida, Proparia, Pseudoacicaudus, Pseudoclelandia, Pseudocrecephalaspina, Pseudosalteria, Pseudosarkia, Pseudotermierella, Ptyttstrigis, Punctaspis, Redlichiida, Remopleuridacea, Repinaella, Resseria, Resseropsis, Rochaeva, Sarkiidae, Serksia, Serksioides, Sibiriopeltis, Sphaerocare, Spizharaspis, Stittaspis, Sukhanaspis, Sunaspididae, Taidonurus, Taplimella, Tazemmourtia, Thymurus, Toxotis, Trapezocephalina, Triarthrella, Trilobus, Tuberaspis, Tuyunaspis, Ulrickaspis, Utiidae, Westergardia, Winsteria, Wuaniidae, Zigzuella
Subtaxa: none
Ecology: fast-moving low-level epifaunal carnivore
Distribution:
• Devonian of Algeria (1 collection), Australia (7), Belgium (3), Brazil (1), Canada (7: Northwest Territories, Nunavut), the Czech Republic (39), France (1), Germany (6), Italy (3), Kazakhstan (1), Morocco (12), the Russian Federation (7), Thailand (1), Uzbekistan (2)
• Silurian of Australia (2), Canada (3: Northwest Territories, Quebec, Yukon Territory), China (2), the Czech Republic (27), Greenland (6), the United Kingdom (11), United States (4: Alaska, Wisconsin)
• Dobrotivian of the Czech Republic (1), France (1), Portugal (2)
• Whiterockian of Canada (16: Newfoundland and Labrador, Quebec), Norway (1), United States (19: Nevada, New York, Oklahoma, Vermont)
• Chazyan of Canada (17: Newfoundland and Labrador, Northwest Territories, Quebec), United States (18: New York)
• Volkhov of Lithuania (1), the Russian Federation (17), Sweden (2)
• Langevoja of Sweden (2)
• Arenig of Argentina (2), Canada (1: Newfoundland and Labrador), China (3), Sweden (2)
• Ordovician of Argentina (3), Australia (3), Canada (24: British Columbia, Newfoundland, Newfoundland and Labrador, Northwest Territories, Nunavut, Ontario, Quebec, Québec), China (11), the Czech Republic (6), Estonia (2), France (2), Iran (1), Ireland (2), Italy (1), Kazakhstan (1), Morocco (1), Norway (11), Portugal (1), the Russian Federation (1), Sweden (6), the United Kingdom (17), United States (27: Alaska, Idaho, Illinois, Iowa/Wisconsin, Kentucky, Maine, Minnesota, Missouri, Nevada, Oklahoma, Tennessee, Virginia)
• Tremadoc of Antarctica (1), Argentina (1), Canada (3: Alberta), China (2), the Czech Republic (2), Sweden (3)
• Skullrockian of Argentina (1)
• Taoyuanian of China (1)
• Cambrian of Canada (25: Alberta, Newfoundland and Labrador, Quebec, Yukon), China (20), Kazakhstan (22), the Russian Federation (5), United States (15: Missouri, Nevada, New York, Texas, Utah, Vermont, Wisconsin, Wyoming)
Total: 472 collections including 529 occurrences