Parent taxon: Trilobita according to J. D. Beech and J. C. Lamsdell 2021
Sister taxa: Aglagalia, Agraulaeva, Aguaraya, Aldonaiidae, Altajaharpes, Amechilus, Amgaspididae, Ampellatocephalina, Anacopodia, Antagmidae, Antatlaeva, Atopidae, Auritamidae, Avoninidae, Batchatiella, Bathynotidae, Beikedamaspis, Bergerolimbus, Burlingiidae, Changshaniidae, Chinensis, Corynexochida, Crossoura, Curiaspis, Cuyanaspis, Cyclolenzellina, Dalmanites troosti, Deltacare, Devonocoryphe, Edganocare, Elleria, Elleria morani, Ellipsocephaloididae, Ellipsuella, Entomostracites, Erbenochile, Eulomella, Eutrilobita, Fabulaspis, Faciura, Gentilaspis, Gigmpeltis, Goycoia, Helioproetoides, Holocephalinidae, Howelluella, Idiorapha, Ignotogregatidae, Indigestus, Irvingelloides, Ishpella, Isocolidae, Issafenia, Ityophoridae, Jamrogiidae, Jonotus, Juriietella, Kamlacus, Kanfuqingia, Leptopilus, Libristoma, Lichida, Llanoaspididae, Loxopeltis, Lusampa, Mapaniidae, Megalaspidicus, Meneghinella, Metapopristis, Monkaspididae, Moolatrilo, Moretaeva, Moretarva, Namanoiidae, Neochilonorria, Neomalungia, Neotaenicephalus, Nepeidae, Odontopleurida, Olenida, Onchonotopsidae, Opisthoparia, Orictocara, Otekmaspis, Otenaspella, Paichiashania, Paleoaethia, Parakingaspidoides, Pararania, Peculicephalina, Perissopyge, Phacopida, Phacops (Monorakos), Piliolites, Pliomerellus, Poculicephalina, Polycyrtaspididae, Prodamesella, Proetida, Proparia, Pseudoacicaudus, Pseudoclelandia, Pseudocrecephalaspina, Pseudosalteria, Pseudosarkia, Pseudotermierella, Ptyttstrigis, Punctaspis, Redlichiida, Remopleuridacea, Repinaella, Resseria, Resseropsis, Rochaeva, Sarkiidae, Serksia, Serksioides, Sibiriopeltis, Sphaerocare, Spizharaspis, Stittaspis, Sukhanaspis, Sunaspididae, Taidonurus, Taplimella, Tazemmourtia, Thymurus, Toxotis, Trapezocephalina, Triarthrella, Trilobus, Tuberaspis, Tuyunaspis, Ulrickaspis, Utiidae, Westergardia, Winsteria, Wuaniidae, Zigzuella
Subtaxa: none
Ecology: fast-moving low-level epifaunal carnivore
Distribution:
• Devonian of Algeria (1 collection), Australia (7), Belgium (3), Brazil (1), Canada (7: Northwest Territories, Nunavut), Czechia (39), France (3), Germany (16), Italy (3), Kazakhstan (1), Morocco (13), the Russian Federation (7), Thailand (1), Uzbekistan (2)
• Silurian of Australia (2), Canada (3: Northwest Territories, Quebec, Yukon Territory), China (2), Czechia (27), Greenland (6), the United Kingdom (11), United States (4: Alaska, Wisconsin)
• Dobrotivian of Czechia (1), France (1), Portugal (2)
• Whiterockian of Canada (16: Newfoundland and Labrador, Quebec), Norway (1), United States (19: Nevada, New York, Oklahoma, Vermont)
• Chazyan of Canada (17: Newfoundland and Labrador, Northwest Territories, Quebec), United States (18: New York)
• Langevoja of Sweden (2)
• Billingen of Sweden (1)
• Arenig of Argentina (2), Canada (1: Newfoundland and Labrador), China (3), Sweden (2)
• Ordovician of Antarctica (1), Argentina (4), Australia (3), Canada (27: Alberta, British Columbia, Newfoundland, Newfoundland and Labrador, Northwest Territories, Nunavut, Ontario, Quebec, Québec), China (14), Czechia (8), Estonia (2), France (2), Iran (1), Ireland (2), Italy (1), Kazakhstan (1), Lithuania (1), Morocco (2), Norway (11), Portugal (1), the Russian Federation (18), Sweden (10), the United Kingdom (17), United States (27: Alaska, Idaho, Illinois, Iowa/Wisconsin, Kentucky, Maine, Minnesota, Missouri, Nevada, Oklahoma, Tennessee, Virginia)
• Skullrockian of Argentina (1)
• Cambrian of Canada (25: Alberta, Newfoundland and Labrador, Quebec, Yukon), China (20), Kazakhstan (22), the Russian Federation (5), South Korea (10), United States (26: Missouri, Nevada, New York, Oklahoma, Texas, Utah, Vermont, Wisconsin, Wyoming)
Total: 507 collections including 570 occurrences