PBDB Taxon

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Mosasauridae (mosasaur)

Reptilia - Mosasauridae

Taxonomy

Mosasauridae was named by Gervais (1852). Its type is Mosasaurus. It was considered monophyletic by Lee and Caldwell (2000), Lindgren and Siverson (2002), Cuthbertson et al. (2007), Buchy et al. (2007) and Konishi et al. (2012) and Karl and Nyhuis (2012).

It was assigned to Pythonomorpha by Cope (1869), Cope (1870), Cope (1871) and Cope (1875); to Pythonomorpha by Zittel (1890) and Hay (1902); to Mosasauriformes by Hay (1930); to Platynota by McDowell and Bogert (1954); to Lacertilia by Osborn (1899), Osborn (1899) and Thurmond and Jones (1981); to Platynota by Russell (1964), Miller (1967), Wiffen (1980), Wiffen (1990) and Lingham-Soliar (1995); to Varanoidea by Baur (1890), Antunes (1964), Kuhn (1966), Welles and Gregg (1971), Seiffert (1975), Carroll (1988) and Páramo-Fonseca (2000); to Pythonomorpha by Lee and Caldwell (2000); to Lacertilia by Nicholls and Meckert (2002); to Mosasauria by Kuhn (1946) and Reeder et al. (2015); to Mosasauroidea by Schulp et al. (2004), Bardet et al. (2005), Bell and Polcyn (2005), Schulp et al. (2008), Bell et al. (2013), Frey et al. (2016) and Hornung et al. (2018); to Lepidosauromorpha by Agnolín et al. (2023); to Squamata by Wright and Shannon (1988), Lindgren and Siverson (2002), Cuthbertson et al. (2007), Buchy et al. (2007), Polcyn and Everhart (2008), Komishi and Caldwell (2009), Konishi et al. (2010), Konishi et al. (2011), Konishi and Caldwell (2011), Polcyn et al. (2012), Konishi et al. (2012), Karl and Nyhuis (2012), Fernández and Talevi (2015), Cuthbertson and Holmes (2015), González Ruiz et al. (2019), Strong et al. (2020), Otero (2021), AbdelGawad et al. (2021), Zietlow et al. (2023), O’Gorman et al. (2023) and Garzon et al. (2024); and to Squamata by Bardet et al. (2004), Martin and Crame (2006), Schulp et al. (2009), Otero et al. (2012), Otero et al. (2017), Lively (2018), Novas et al. (2019), Grigoriev and Grabovskiy (2020), Lively (2020), Kiernan and Ebersole (2023), Shaker et al. (2023), Plasse et al. (2024) and Rivera-Sylva et al. (2024).

Subtaxa

Amphorosteus, Amphorosteus brumbyi, Clidastes iguanavus, Compressidens, Globidentinae, Haasiasaurus (syn. Haasia), Halisaurinae, Halisauromorpha, Liodon congrops, Mosasaurinae, Mosasaurus crassidens, Mosasaurus mitchillii, Mosasaurus platyodon, Platecarpinae, Prognathosaurus, Russellosaurina, Tylosaurinae, Tylosaurus capensis, Yaguarasaurinae

Synonyms

Mosasaurus platyodon was named by Sauvage (1873). Its type specimen is Ã‰cole des Mines, Paris, a tooth, and it is a 3D body fossil.

It was considered a nomen dubium by Buffetaut and Bardet (2012).
Clidastes iguanavus was named by Cope (1868). Its type specimen is Peabody Museum of Natural History no. 1601, a set of vertebrae (single anterior thoracic vertebra), and it is a 3D body fossil. Its type locality is Marl pit near Swedesboro, which is in a Campanian/Campanian marine marl in the Marshalltown Formation of New Jersey. It is the type species of Clidastes.

It was considered a nomen dubium by Kiernan (1992).
Clidastidae was named by Cope (1869). Its type is Clidastes.

It was synonymized subjectively with Mosasauridae by Russell (1967).

It was assigned to Pythonomorpha by Cope (1869) and Cope (1871).
Geosaurus mitchilli was named by DeKay (1830).

It was recombined as Mosasaurus mitchillii by Cope (1869); it was synonymized subjectively with Atlantochelys mortoni by Leidy (1873); it was recombined as Liodon mitchillii by Cope (1875); it was recombined as Macrosaurus mitchillii by Hay (1902) and Hay (1930); it was considered a nomen vanum by Russell (1967).
Amphorosteus brumbyi was named by Gibbes (1851).

It was recombined as Mosasaurus brumbyi by Cope (1869), Hay (1902) and Hay (1930); it was considered a nomen vanum by Russell (1967).
Edestosauridae was named by Marsh (1876).

It was synonymized subjectively with Mosasauridae by Russell (1967).
Globidensidae was named by Dollo (1924). Its type is Globidens.

It was corrected as Globidentidae by Hay (1930) and Kuhn (1946); it was synonymized subjectively with Mosasauridae by Russell (1967).

It was assigned to Mosasauriformes by Hay (1930); and to Mosasauria by Kuhn (1946).
Amphorosteus was named by Gibbes (1851) [Sepkoski's age data: K u].

It was synonymized subjectively with Mosasaurus by Cope (1869), Zittel (1890) and Hay (1902); it was considered a nomen vanum by Russell (1967).

It was assigned to Squamata by Sepkoski (2002).
Tylosaurus capensis was named by Broom (1912). It is a 3D body fossil. Its type locality is Southwest of the Mzamba River, Pondoland, which is in a Maastrichtian marine horizon in South Africa.

It was considered a nomen dubium by Woolley et al. (2022).
Mosasaurus crassidens was named by Marsh (1870).

It was considered a nomen dubium by Russell (1967).
Liodon congrops was named by Cope (1870).

It was recombined as Tylosaurus congrops by Hay (1902) and Hay (1930); it was considered a nomen vanum by Russell (1967).

Synonymy list

Year	Name and author
1830	Geosaurus mitchilli DeKay
1851	Amphorosteus Gibbes p. 10
1851	Amphorosteus brumbyi Gibbes p. 11
1852	Mosasauridae Gervais p. 261
1868	Clidastes iguanavus Cope pp. 233
1869	Clidastes iguanavus Cope p. 258
1869	Clidastidae Cope p. 258
1869	Mosasauridae Cope p. 258
1869	Mosasaurus mitchillii Cope p. 262
1869	Mosasaurus brumbyi Cope p. 264
1870	Mosasauridae Cope p. 183
1870	Liodon congrops Cope p. 206
1870	Clidastes iguanavus Cope p. 220
1870	Liodon congrops Cope pp. 271-272
1870	Mosasaurus crassidens Marsh p. 2
1871	Clidastidae Cope p. 237
1871	Mosasauridae Cope p. 237
1873	Clidastes iguanavus Leidy p. 281
1873	Mosasaurus platyodon Sauvage p. 385
1875	Mosasauridae Cope p. 20
1875	Liodon mitchillii Cope p. 270
1876	Edestosauridae Marsh p. 59
1890	Mosasauridae Baur p. 262
1890	Mosasauridae Zittel p. 620
1899	Mosasauridae Osborn p. 188
1902	Mosasauridae Hay p. 464
1902	Mosasaurus brumbyi Hay p. 465
1902	Mosasaurus crassidens Hay p. 465
1902	Clidastes iguanavus Hay p. 467
1902	Macrosaurus mitchillii Hay p. 471
1902	Tylosaurus congrops Hay p. 472
1912	Tylosaurus capensis Broom p. 333
1924	Globidensidae Dollo p. 188
1930	Mosasauridae Hay p. 258
1930	Macrosaurus mitchillii Hay p. 259
1930	Tylosaurus congrops Hay p. 259
1930	Mosasaurus brumbyi Hay p. 262
1930	Mosasaurus crassidens Hay p. 262
1930	Clidastes iguanavus Hay p. 263
1930	Globidentidae Hay p. 264
1946	Globidentidae Kuhn p. 61
1946	Mosasauridae Kuhn p. 61
1954	Mosasauridae McDowell and Bogert p. 132
1964	Mosasauridae Antunes p. 137
1964	Mosasauridae Russell p. 167
1966	Mosasauridae Kuhn p. 59
1967	Mosasauridae Miller p. 227
1967	Clidastes iguanavus Russell p. 126
1971	Mosasauridae Welles and Gregg p. 49
1975	Mosasauridae Seiffert p. 10
1980	Mosasauridae Wiffen p. 508
1981	Mosasauridae Thurmond and Jones p. 148
1988	Mosasauridae Carroll
1988	Mosasauridae Wright and Shannon p. 102
1990	Mosasauridae Wiffen p. 67
1995	Mosasauridae Lingham-Soliar
2000	Mosasauridae Lee and Caldwell
2000	Mosasauridae Páramo-Fonseca p. 123
2002	Mosasauridae Lindgren and Siverson
2002	Mosasauridae Nicholls and Meckert p. 1593
2002	Amphorosteus Sepkoski
2004	Mosasauridae Bardet et al. p. 316
2004	Mosasauridae Schulp et al. p. 744
2005	Mosasauridae Bardet et al.
2005	Mosasauridae Bell and Polcyn p. 178
2006	Mosasauridae Martin and Crame p. 113
2007	Mosasauridae Buchy et al.
2007	Mosasauridae Cuthbertson et al.
2008	Mosasauridae Polcyn and Everhart p. 14
2008	Mosasauridae Schulp et al. p. 1
2009	Mosasauridae Komishi and Caldwell p. 418
2009	Mosasauridae Schulp et al. p. 162
2010	Mosasauridae Konishi et al.
2011	Mosasauridae Konishi et al. p. 1028
2011	Mosasauridae Konishi and Caldwell p. 756
2012	Mosasauridae Karl and Nyhuis
2012	Mosasauridae Konishi et al.
2012	Mosasauridae Otero et al. p. 130
2012	Mosasauridae Polcyn et al. p. 125
2013	Mosasauridae Bell Jr. et al. p. 574
2015	Mosasauridae Cuthbertson and Holmes
2015	Mosasauridae Fernández and Talevi p. 485
2015	Mosasauridae Reeder et al.
2016	Mosasauridae Frey et al. p. 20
2017	Mosasauridae Otero et al.
2018	Mosasauridae Hornung et al. p. 544
2018	Mosasauridae Lively p. 524
2019	Tylosaurus capensis Driscoll et al.
2019	Mosasauridae González Ruiz et al. p. 3
2019	Mosasauridae Novas et al. p. 227
2020	Mosasauridae Grigoriev and Grabovskiy p. 2
2020	Mosasauridae Lively p. 15
2020	Mosasauridae Strong et al. p. 8
2021	Mosasauridae AbdelGawad et al.
2021	Mosasauridae Otero p. 2
2023	Mosasauridae Agnolín et al.
2023	Mosasauridae Kiernan and Ebersole p. 4
2023	Mosasauridae O’Gorman et al.
2023	Mosasauridae Shaker et al. p. 4
2023	Mosasauridae Zietlow et al. p. 6
2024	Mosasauridae Garzon et al. p. 7
2024	Mosasauridae Plasse et al. p. 4
2024	Mosasauridae Rivera-Sylva et al. p. 4

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()

Rank	Name	Author
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Squamata	()
family	Mosasauridae
family	Mosasauridae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Mosasauridae Gervais 1852 [mosasaur]

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G. †Compressidens Dollo 1924

†Compressidens belgicus Woodward 1891

G. †Haasiasaurus Polcyn et al. 2003

†Haasiasaurus gittelmani Polcyn et al. 1999

Invalid names: Haasia Polcyn et al. 1999 [replaced]

Unr. †Halisaurinae Bardet et al. 2005

Unr. †Halisaurini Longrich et al. 2021

G. †Eonatator Bardet et al. 2005

†Eonatator coellensis Páramo-Fonseca 2013

†Eonatator sternbergii Wiman 1920

G. †Halisaurus Marsh 1869

†Halisaurus arambourgi Bardet et al. 2005

†Halisaurus hebae Shaker et al. 2023

†Halisaurus onchognathus Merriam 1894

†Halisaurus platyspondylus Marsh 1869

Invalid names: Baptosaurus Marsh 1870 [nomen dubium]

G. †Phosphorosaurus Dollo 1889

†Phosphorosaurus ortliebi Dollo 1889

†Phosphorosaurus ponpetelegans Konishi et al. 2016

Unr. †Pluridensini Longrich et al. 2021

G. †Pluridens Lingham-Soliar 1998

†Pluridens calabaria Longrich 2016

†Pluridens serpentis Longrich et al. 2021

†Pluridens walkeri Lingham-Soliar 1998

Subfm. †Mosasaurinae Gervais 1853

G. †Clidastes Cope 1868

†Clidastes cineriarum Cope 1870

†Clidastes intermedius Leidy 1870

†Clidastes liodontus Merriam 1894

†Clidastes moorevillensis Shannon 1975

†Clidastes propython Cope 1869

Invalid names: Clidastes medius Merriam 1894 [synonym], Clidastes pumilus Marsh 1871 [synonym], Clidastes westi Williston and Case 1892 [synonym], Clidastes wymani Marsh 1871 [synonym], Edestosaurus dispar Marsh 1871 [synonym], Edestosaurus rex Marsh 1872 [synonym], Edestosaurus stenops Cope 1871 [synonym], Edestosaurus tortor Cope 1871 [synonym], Edestosaurus velox Marsh 1871 [synonym]

Invalid names: Edestosaurus Marsh 1871 [synonym]

G. †Dallasaurus Bell and Polcyn 2005

†Dallasaurus turneri Bell and Polcyn 2005

Tr. †Globidensini Russell 1967

G. †Carinodens Thurmond 1969

†Carinodens belgicus Woodward 1891

†Carinodens fraasi Dollo 1913

†Carinodens minalmamar Schulp et al. 2009

G. †Globidens Gilmore 1912

†Globidens alabamensis Gilmore 1912

†Globidens dakotensis Russell 1975

†Globidens phosphaticus Bardet et al. 2005

†Globidens schurmanni Martin 2007

†Globidens simplex LeBlanc et al. 2019

G. †Prognathodon Dollo 1889

†Prognathodon compressidens Gaudry 1892

†Prognathodon currii Christiansen and Bonde 2002

†Prognathodon giganteus Dollo 1889

†Prognathodon kianda Schulp et al. 2008

†Prognathodon lutugini Yakovlev 1905

†Prognathodon mosasauroides Gaudry 1892

†Prognathodon overtoni Williston 1897

†Prognathodon rapax Hay 1902

Invalid names: Ancylocentrum hungerfordi Chaffee 1939 [synonym], Liodon validus Cope 1868 [synonym]

†Prognathodon saturator Dortangs et al. 2002

†Prognathodon sectorius Cope 1871

†Prognathodon solvayi Dollo 1889

†Prognathodon waiparaensis Welles and Gregg 1971

Invalid names: Ancylocentrum Schmidt 1927 [synonym], Brachysaurana Strand 1926 [synonym], Brachysaurus Williston 1897 [synonym], Dollosaurus Yakovlev 1901 [synonym], Liodon Owen 1842 [nomen dubium]

G. †Gnathomortis Lively 2020

†Gnathomortis stadtmani Kass 1999

G. †Igdamanosaurus Lingham-Soliar 1991

†Igdamanosaurus aegyptiacus Zdansky 1935

G. †Jormungandr Zietlow et al. 2023

†Jormungandr walhallaensis Zietlow et al. 2023

G. †Kourisodon Nicholls and Meckert 2002

†Kourisodon puntledgensis Nicholls and Meckert 2002

G. †Leiodon Owen 1842

†Leiodon anceps Owen 1841

†Leiodon lundgreni Schröder 1885

G. †Megapterygius Konishi et al. 2023

†Megapterygius wakayamaensis Konishi et al. 2023

G. †Moanasaurus Wiffen 1980

†Moanasaurus mangahouangae Wiffen 1980

Tr. †Mosasaurini Russell 1967

G. †Eremiasaurus LeBlanc et al. 2012

†Eremiasaurus heterodontus LeBlanc et al. 2012

G. †Plotosaurus Camp 1951

†Plotosaurus bennisoni Camp 1942

†Plotosaurus tuckeri Camp 1942

Invalid names: Garzasaurus Anderson 1943 [nomen nudum], Kolposaurus Camp 1942 [replaced]

Invalid names: Liodon asiaticum Répelin 1915 [nomen dubium]

G. †Mosasaurus Parkinson 1822

†Mosasaurus beaugei Arambourg 1952

†Mosasaurus caroliniensis Gibbes 1851

†Mosasaurus conodon Cope 1881

†Mosasaurus couperi Gibbes 1851

†Mosasaurus dekayi Bronn 1838

Invalid names: Mosasaurus major DeKay 1842 [synonym], Mosasaurus meirsii Marsh 1869 [synonym]

†Mosasaurus depressus Cope 1869

Invalid names: Halisaurus fraternus Marsh 1869 [synonym], Mosasaurus copeanus Marsh 1869 [synonym]

†Mosasaurus flemingi Wiffen 1990

†Mosasaurus gaudryi Thevenin 1896

†Mosasaurus gracilis Owen 1850

†Mosasaurus grosjeani Greppin 1870

†Mosasaurus hobetsuensis Suzuki 1985

†Mosasaurus hoffmannii Mantell 1829

Invalid names: Mosasaurus belgicus Holl 1829 [synonym], Mosasaurus camperi Meyer 1832 [synonym], Mosasaurus maximus Cope 1869 [synonym]

†Mosasaurus horridus Williston 1895

†Mosasaurus ivoensis Russell 1967

†Mosasaurus lemonnieri Dollo 1889

†Mosasaurus lonzeensis Dollo 1904

†Mosasaurus minor Gibbes 1851

†Mosasaurus missouriensis Harlan 1834

Invalid names: Mosasaurus maximilliani Goldfuss 1845 [synonym], Mosasaurus neovidii von Meyer 1845 [synonym]

†Mosasaurus mokoroa Welles and Gregg 1971

Invalid names: Leiodon haumuriensis Hector 1874 [synonym]

†Mosasaurus occidentalis Morton 1844

†Mosasaurus prismaticus Sakurai et al. 1999

†Mosasaurus scanicus Schröder 1885

Invalid names: Amphekepubis Mehl 1930 [synonym], Baseodon Leidy 1865 [synonym], Batrachiosaurus Harlan 1839 [synonym], Batrachotherium Harlan 1839 [synonym], Drepanodon Leidy 1861 [synonym], Lesticodus Leidy 1861 [synonym], Macrosaurus Owen 1849 [synonym], Mosasaurus impar Leidy 1856 [nomen dubium], Mosasaurus reversus Leidy 1865 [nomen dubium], Nectoportheus Cope 1868 [synonym], Pterycollosaurus Dollo 1882 [synonym]

G. †Plesiotylosaurus Camp 1942

†Plesiotylosaurus crassidens Camp 1942

G. †Polygonodon Leidy 1856

Unr. †Prognathodontini Russell 1967

G. †Thalassotitan Longrich et al. 2022

†Thalassotitan atrox Longrich et al. 2022

G. †Rikisaurus Wiffen 1990

†Rikisaurus tehoensis Wiffen 1990

G. †Stelladens Longrich et al. 2023

†Stelladens mysteriosus Longrich et al. 2023

G. †Xenodens Longrich et al. 2021

†Xenodens calminechari Longrich et al. 2021

Invalid names: Globindensini Bell 1997 [empty], Leiodontini Lingham-Soliar 1995 [empty], Plotosaurini Russell 1967 [empty]

G. †Prognathosaurus Dollo 1889

Unr. †Russellosaurina Bell 1997

G. †Angolasaurus Antunes 1964

†Angolasaurus bocagei Antunes 1964

Invalid names: Angolasaurus bocagei Antunes 1964 [valid]

Subfm. †Plioplatecarpinae Dollo 1884

G. †Angolasaurus Antunes 1964

†Angolasaurus bocagei Antunes 1964

Invalid names: Angolasaurus bocagei Antunes 1964 [valid]

G. †Ectenosaurus Russell 1967

†Ectenosaurus clidastoides Merriam 1894

Invalid names: Platecarpus oxyrhinus Merriam 1894 [synonym]

†Ectenosaurus shannoni Kiernan and Ebersole 2023

†Ectenosaurus tlemonectes Kiernan and Ebersole 2023

Unr. †Plioplatecarpini Russell 1967

G. †Latoplatecarpus Konishi and Caldwell 2011

†Latoplatecarpus nichollsae Cuthbertson et al. 2007

†Latoplatecarpus willistoni Konishi and Caldwell 2011

G. †Platecarpus Cope 1869

†Platecarpus crassartus Cope 1871

†Platecarpus curtirostris Cope 1871

†Platecarpus planifrons Cope 1874

†Platecarpus tympaniticus Cope 1869

Invalid names: Holcodus coryphaeus Cope 1871 [synonym], Lestosaurus felix Marsh 1872 [synonym], Lestosaurus gracilis Marsh 1872 [synonym], Lestosaurus latifrons Marsh 1872 [synonym], Lestosaurus simus Marsh 1872 [synonym], Liodon ictericus Cope 1870 [synonym], Liodon mudgei Cope 1870 [synonym]

Invalid names: Holcodus Gibbes 1851 [synonym], Holosaurus Marsh 1880 [synonym], Lestosaurus Marsh 1872 [synonym], Platecarpus affinis Leidy 1873 [nomen dubium], Platecarpus anguliferus Cope 1874 [nomen vanum], Platecarpus brachycephalus Loomis 1915 [nomen dubium], Platecarpus glandiferus Cope 1871 [nomen dubium], Platecarpus latispinus Cope 1871 [nomen dubium], Platecarpus tectulus Cope 1871 [nomen dubium], Sironectes Cope 1874 [synonym]

G. †Plesioplatecarpus Konishi and Caldwell 2011

G. †Plioplatecarpus Dollo 1882

†Plioplatecarpus crassartus Cope 1871

†Plioplatecarpus houzeaui Dollo 1889

†Plioplatecarpus marshi Dollo 1882

†Plioplatecarpus peckensis Cuthbertson and Holmes 2015

†Plioplatecarpus primaevus Russell 1967

Invalid names: Oterognathus Dollo 1889 [synonym]

G. †Sarabosaurus Polcyn et al. 2023

†Sarabosaurus dahli Polcyn et al. 2023

Unr. †Selmasaurini Longrich et al. 2024

G. †Gavialimimus Strong et al. 2020

†Gavialimimus almaghribensis Strong et al. 2020

G. †Goronyosaurus Azzaroli et al. 1972

†Goronyosaurus nigeriensis Swinton 1930

G. †Khinjaria Longrich et al. 2024

†Khinjaria acuta Longrich et al. 2024

G. †Selmasaurus Wright and Shannon 1988

†Selmasaurus johnsoni Polcyn and Everhart 2008

†Selmasaurus russelli Wright and Shannon 1988

G. †Yaguarasaurus Páramo 1994

†Yaguarasaurus columbianus Páramo 1994

†Yaguarasaurus regiomontanus Rivera-Sylva et al. 2024

Invalid names: Platecarpus ptychodon Arambourg 1952 [nomen dubium], Platecarpus somenensis Thevenin 1896 [nomen dubium]

Invalid names: Platecarpus ptychodus Arambourg 1952 [nomen dubium]

Subfm. †Tylosaurinae Wiliston 1895

G. †Hainosaurus Dollo 1885

†Hainosaurus boubker Rempert et al. 2022

†Hainosaurus neumilleri Martin 2007

G. †Kaikaifilu Otero et al. 2017

†Kaikaifilu hervei Otero et al. 2017

G. †Taniwhasaurus Hector 1874

†Taniwhasaurus antarcticus Novas et al. 2002

†Taniwhasaurus mikasaensis Caldwell et al. 2008

†Taniwhasaurus oweni Hector 1874

Invalid names: Tylosaurus haumuriensis Hector 1874 [synonym]

Invalid names: Lakumasaurus Novas et al. 2002 [synonym]

G. †Tylosaurus Marsh 1872

†Tylosaurus bernardi Dollo 1885

†Tylosaurus iembeensis Antunes 1964

†Tylosaurus ivoensis Russell 1967

†Tylosaurus kansasensis Everhart 2005

†Tylosaurus laticaudus Marsh 1870

†Tylosaurus nepaeolicus Cope 1874

†Tylosaurus pembinensis Nicholls 1988

†Tylosaurus perlatus Cope 1869

†Tylosaurus proriger Cope 1869

Invalid names: Rhinosaurus micromus Marsh 1872 [synonym]

†Tylosaurus saskatchewanensis Jiménez-Huidobro et al. 2018

†Tylosaurus zangerli Russell 1970

Invalid names: Elliptonodon Emmons 1858 [nomen dubium], Elliptonodon compressus Emmons 1858 [nomen dubium], Mosasaurus alseni Stolley 1892 [nomen dubium], Rhamphosaurus Cope 1872 [replaced], Rhinosaurus Marsh 1872 [replaced], Tylosaurus dyspelor Cope 1870 [nomen vanum]

Invalid names: Hainosaurus lonzeensis Dollo 1904 [nomen dubium]

Subfm. †Yaguarasaurinae Palci et al. 2013

G. †Romeosaurus Palci et al. 2013

†Romeosaurus fumanensis Palci et al. 2013

†Romeosaurus sorbinii Palci et al. 2013

G. †Russellosaurus Polcyn and Bell 2005

†Russellosaurus coheni Polcyn and Bell 2005

Invalid names: Amphorosteus Gibbes 1851 [nomen vanum], Amphorosteus brumbyi Gibbes 1851 [nomen vanum], Clidastes iguanavus Cope 1868 [nomen dubium], Clidastidae Cope 1869 [synonym], Edestosauridae Marsh 1876 [synonym], Globidensidae Dollo 1924 [synonym], Globidentinae [empty], Halisauromorpha Bell and Polcyn 2005 [empty], Liodon congrops Cope 1870 [nomen vanum], Mosasaurus crassidens Marsh 1870 [nomen dubium], Mosasaurus mitchillii DeKay 1830 [nomen vanum], Mosasaurus platyodon Sauvage 1873 [nomen dubium], Platecarpinae Williston 1897 [empty], Tylosaurus capensis Broom 1912 [nomen dubium]

Diagnosis

Reference	Diagnosis
E. D. Cope 1869	The vertebrae without the zygosphen articulation; the pterygoids in contact on the median line; (?) a postparietal arch.
E. D. Cope 1869 (Clidastidae)	The vertebrae with the zygosphen articulation; the pterygoids free on the internal and external margins; no (?) postparietal arch
S. B. McDowell and C. M. Bogert 1954	Temporal arch present; frontals fused; descending processes of frontals meeting to underarch olfactory tracts bu tnot separating orbits; pineal foramen present; vertebral central articulations not oblique; hyperphalangy present; no sacrum; interclavicle absent or reduced; pelvis much reduced; cervical vertebrae seven; palatine toothless, pterygoid strongly toothed; quadrate massive, surrounding the columella auris of the stapes; muzzle long and pointed, the nasal fused to the premaxilla; teeth secondarily thecodont, lodged in an uncompartmented groove.