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Bethylidae (flat wasp)
Taxonomy
Bethylidae was named by Forster (1856). It is extant.
It was assigned to Apocrita by Carpenter (1992); and to Chrysidoidea by Brothers (1999), Sharkey (2007), McKellar and Engel (2014), Antropov (2014), Engel et al. (2016), Cockx et al. (2016), Brazidec and Perrichot (2022).
It was assigned to Apocrita by Carpenter (1992); and to Chrysidoidea by Brothers (1999), Sharkey (2007), McKellar and Engel (2014), Antropov (2014), Engel et al. (2016), Cockx et al. (2016), Brazidec and Perrichot (2022).
Synonyms
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Synonymy list
Year | Name and author |
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1856 | Bethylidae Forster p. 95 |
1914 | Epyrinae Kieffer |
1988 | Epyrinae Sorg p. 72 |
1992 | Bethylidae Carpenter |
1999 | Bethylidae Brothers p. 243 |
2002 | Fushunochrysidae Hong p. 314 |
2007 | Bethylidae Sharkey |
2014 | Bethylidae Antropov p. 394 |
2014 | Bethylidae McKellar and Engel p. 436 |
2016 | Bethylidae Cockx et al. p. 4 |
2016 | Bethylidae Engel et al. p. 4 |
2022 | Bethylidae Brazidec and Perrichot p. 700 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Bethylidae Forster 1856 [flat wasp]
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Subfm. Bethylinae Halliday 1840
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G. Bethylus Latreille 1802
G. Eupsenella Westwood 1874
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†Eupsenella aulax Ramos and Azevedo 2014
†Eupsenella eocenica De Ploëg and Nel 2004
†Eupsenella klesoviana Ramos and Azevedo 2014
†Eupsenella rossica Ramos and Azevedo 2014
†Eupsenella yantarnica Ramos and Azevedo 2014
Invalid names: Eupsenella neoeocenica Ramos and Azevedo 2018 [nomen nudum], Eupsenella neohongi Ramos and Azevedo 2018 [nomen nudum], Fushunochrysites Hong 2002 [nomen nudum], Protobethylus De Ploëg and Nel 2004 [synonym], Sinibethylus Hong 2002 [nomen nudum]
G. Goniozus Förster 1856
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†Goniozus antiquus Olmi 2005
†Goniozus contractus Brues 1933
†Goniozus copalinus Meunier 1916
†Goniozus cotyi Brazidec and Perrichot 2022
†Goniozus definitus Ramos and Azevedo 2014
†Goniozus respectus Sorg 1988
Invalid names: Messoria Meunier 1916 [synonym], Perisierola Kieffer 1914 [synonym], Pterosclerogibba Olmi 2005 [synonym]
G. Lytopsenella Kieffer 1911
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†Lytopsenella baltica Ramos and Azevedo 2014
†Lytopsenella crastina Brues 1923
†Lytopsenella kerneggeri Ohl 1995
†Lytopsenella maritima Ramos and Azevedo 2014
†Lytopsenella setigera Brues 1923
†Lytopsenella simplex Brues 1923
G. Sierola Cameron 1881
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†Sierola colomboi Brazidec and Perrichot 2022
†Sierola hastata Sorg 1988
†Sierola rovniana Ramos and Azevedo 2014
Subfm. †Cretabythinae Brazidec et al. 2023
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Subfm. †Elektroepyrinae Colombo and Azevedo 2020
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Subfm. Epyrinae Kieffer 1914
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G. Chlorepyris Kieffer 1913
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†Chlorepyris concaptus Brues 1933
†Chlorepyris deploegi Colombo and Azevedo 2021
†Chlorepyris engeli Colombo and Azevedo 2021
†Chlorepyris gallicus Perrichot and Nel 2008
†Chlorepyris hopei Colombo and Azevedo 2021
†Chlorepyris invelatus Brues 1933
†Chlorepyris jouaulti Colombo and Azevedo 2021
†Chlorepyris mckellari Colombo and Azevedo 2021
†Chlorepyris meunieri Colombo and Azevedo 2021
†Chlorepyris neli Colombo and Azevedo 2021
†Chlorepyris perrichoti Colombo and Azevedo 2021
†Chlorepyris setosus Brues 1933
G. Epyris Westwood 1832
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†Epyris deletus Brues 1910
†Epyris kiefferi Brues 1933
†Epyris moulyi Falieres and Nel 2018
†Epyris rectinervis Cockerell 1921
†Epyris tenellus Statz 1938
Invalid names: Isobrachium Forster 1856 [synonym], Rhabdepyris Kieffer 1904 [synonym]
G. Holepyris Kieffer 1904
G. Laelius Ashmead 1893
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†Laelius pallidus Brues 1933
†Laelius preteritus Barbosa and Azevedo 2013
†Laelius rovnensis Barbosa and Azevedo 2013
G. Rysepyris Kieffer 1906
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†Rysepyris minor Brues 1939
†Rysepyris planiceps Brues 1933
†Rysepyris precursor Brues 1933
†Rysepyris rasnitsyni Colombo and Azevedo 2021
†Rysepyris robustus Brues 1933
†Rysepyris terayamai Colombo and Azevedo 2021
Invalid names: Misepyris Kieffer 1913 [synonym]
Subfm. †Lancepyrinae Azevedo and Azar 2012
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G. †Burmapyris Jouault et al. 2021
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†Burmapyris azevedoi Jouault et al. 2021
†Burmapyris ohmkuhnlei Brazidec et al. 2023
Subfm. Pristocerinae Mocsary 1881
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G. Apenesia Westwood 1874
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†Apenesia miki Terayama 2004
Invalid names: Archaeopristocera Terayama 2004 [synonym]
G. Cleistepyris Kieffer 1910
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†Cleistepyris allosaurus Colombo et al. 2020
†Cleistepyris baryonyx Colombo et al. 2020
G. Eleganesia Alencar and Azevedo 2018
G. Pristocera Klug 1808
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†Pristocera alaini Falières and Nel 2019
†Pristocera skwarrae Brues 1933
Invalid names: Parapristocera Brues 1933 [synonym]
G. Pseudisobrachium Kieffer 1904
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†Pseudisobrachium elatus Brues 1933
†Pseudisobrachium inhabilis Brues 1923
†Pseudisobrachium megalosaurus Colombo et al. 2020
†Pseudisobrachium stegosaurus Colombo et al. 2020
Subfm. †Protopristocerinae Nagy 1974
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Subfm. Scleroderminae Kieffer 1914
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G. †Celonophamia Evans 1973
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†Celonophamia granama McKellar and Engel 2014
†Celonophamia taimyria Evans 1973
G. †Mael Colombo and Azevedo 2021
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G. Sclerodermus Latreille 1809
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†Sclerodermus quadridentatum Cockerell 1917
†Sclerodermus quadridentatus Cockerell 1917
Invalid names: Epyrinae Kieffer 1914 [invalid subgroup], Fushunochrysidae Hong 2002 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Early/Lower Barremian to the top of the Holocene or 125.77000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 121.4 Ma
Minimum age of oldest fossil (stem group age): 121.4 Ma
Collections (54 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Early/Lower Barremian | Lebanon | Lancepyris opertus (138801) | |
Early/Lower Barremian | Spain | Bethylidae indet. (104052) | |
Aptian | Russian Federation | Cretobethylellus lucidus (124811) | |
Early/Lower Albian | Myanmar (Sagaing) | Azepyris delamarrei (229858) Paralanceis chotardi, Yunbayin rossei (215115) | |
Albian | Jordan | Bethylidae indet. (124847) | |
Late/Upper Albian | Spain (Álava) | Lancepyris alavaensis, Liztor pilosus, Cretepyris martini (108769) | |
Early/Lower Cenomanian | Myanmar | Apenesia electriphila, Bethylitella cylindrella, Epyris atavellus, Scleroderma quadridentatum (123020) Burmapyris azevedoi, Protopyris myanmarensis, Cretapristocera longiscapa, Ramageoptera platycephala, Burmapyris ohmkuhnlei, Yunbayin rossei (179227) Gwesped groehni (172900) Holopsenelliscus pankowskiorum (123116) Hukawngepyris setosus (184841) Megalopsenella pouilloni (212258) Nadezhdabythus burmensis (161726) | |
Early/Lower Cenomanian | France (Poitou-Charentes) | Bethylidae indet., Nucifrangibulum carentonensis (161514) | |
Early/Lower Cenomanian | Myanmar (Kachin) | Uniceratops trex (221971) | |
Late/Upper Santonian | Russian Federation | Archaepyris minutus, Celonophamia taimyria, Ekaterina volgatitan (35056) Cretabythus sibiricus, Archaepyris minutus (122590) | |
Santonian | Hungary | Ajkanesia harmincipsziloni, Amissidigitus belae (224054) | |
Middle Campanian | Canada (Alberta) | Bethylidae indet., Celonophamia granama (122383) | |
Ypresian | France (Oise) | Protobethylus eocenicus, Epyrini indet., Rhabdepyris gallicus, Elektroepyris magnificus, Paleoscleroderma lamarrei, Pristocera alaini, Dissomphalus schubnelli, Epyris moulyi, Eopristocera sp., Eopristocera bilobata, Gynopteron inexpectatus (123850) | |
Ypresian | China (Liaoning) | Fushunochrysites eocenicus (124530) Sinibethylus eocenicus (124820) | |
Bridgerian | USA (Colorado) | Eupsenella sp. (229578) | |
MP 13 | Germany (Eifel) | Bethylidae indet. (135641) | |
Priabonian | United Kingdom (England) | Mesitius rectinervis (123959) | |
Priabonian | Denmark | Holepyris sp., Sclerodermini indet., Pristocerinae indet., Sierola hastata, Plastanoxus atrescens (123762) | |
Priabonian | Ukraine | Chlorepyris neli, Cleistepyris allosaurus (113621) Laelius rovnensis, Laelius preteritus, Lytopsenella kerneggeri, Eupsenella aulax, Eupsenella klesoviana, Sierola rovniana, Chlorepyris deploegi, Chlorepyris jouaulti, Chlorepyris mckellari, Holepyris terayamai, Pseudisobrachium megalosaurus, Pseudisobrachium stegosaurus (122997) Voronknoxus zerovae (216854) | |
Priabonian | Russian Federation | Lytopsenella crastina, Epyrinae indet. (173264) | |
Priabonian | Russian Federation (Kaliningrad) | Allobethylus bei (219707) Calyoza longiceps, Protopristocera sucini, Epyris inhabilis, Sierola simplex, Sierola setigera, Sierola crastina, Bethylopteron ambiguum, Parapristocera skwarrae, Eupsenella setigera, Eupsenella simplex, Eupsenella crastina, Prosierola submersa, Perisierola contracta, Artiepyris kiefferi, Misepyris robustus, Misepyris dubius, Holepyris planiceps, Holepyris precursor, Isobrachium concaptum, Isobrachium invelatum, Rhabdepyris setosus, Rhabdepyris elatus, Laelius pallidus (123927) Eupsenella aulax, Lytopsenella baltica, Lytopsenella maritima (160471) Eupsenella crastina (127170) Goniozus definitus (120610) Lytopsenella crastina (124318) Lytopsenella kerneggeri (138800) Lytopsenella kerneggeri, Eupsenella rossica, Eupsenella yantarnica, Chlorepyris engeli, Chlorepyris hopei, Chlorepyris meunieri, Chlorepyris perrichoti, Holepyris rasnitsyni, Archaeonoxus scintillatus, Glenosema electrum, Nothepyris pretiosus, Cleistepyris baryonyx (122591) Misepyris minor, Homoglenus bifossatus (124205) | |
Chadronian | USA (Colorado) | Bethylidae indet. (122976) Epyris deletus (123119) | |
Chattian | Germany (North Rhine-Westphalia) | Epyris tenellus (122010) | |
Early/Lower Miocene | Mexico (Chiapas) | Goniozus cotyi, Bakeriella nanyhelae, Solepyris electromexicanus (225741) | |
Burdigalian - Langhian | Dominican Republic | Alongatepyris pedrocai (229461) Archaeopristocera miki (151097) Crassibethylus dominicanus (203120) Goniozus respectus, Epyrinae indet., Anisepyris gradatus (146557) Pterosclerogibba antiqua (122472) | |
Langhian | China (Fujian) | Sierola colomboi, Parascleroderma palaeosinica (220273) | |
Holocene | Tanzania | Bethylidae indet. (234540) Calyoza ramosa (140788) Calyoza staphylinoides (165051) | |
Holocene | Madagascar | Messoria copalina (151372) |