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Grylloblattodea (ice crawler)
Taxonomy
Grylloblattodea was named by Walker (1914). It is extant.
It was reranked as the suborder Grylloblattodea by Arillo and Engel (2006) and Misof et al. (2014).
It was assigned to Blattopteroidea by Sharov (1968); to Polyorthoptera by Engel and Grimaldi (2004); to Blattoneoptera by Kukalová-Peck and Lawrence (2004); to Polyneoptera by Béthoux and Beckemeyer (2007); to Notoptera by Arillo and Engel (2006) and Misof et al. (2014); and to Xenonomia by Terry and Whiting (2005) and Wipfler et al. (2019).
It was reranked as the suborder Grylloblattodea by Arillo and Engel (2006) and Misof et al. (2014).
It was assigned to Blattopteroidea by Sharov (1968); to Polyorthoptera by Engel and Grimaldi (2004); to Blattoneoptera by Kukalová-Peck and Lawrence (2004); to Polyneoptera by Béthoux and Beckemeyer (2007); to Notoptera by Arillo and Engel (2006) and Misof et al. (2014); and to Xenonomia by Terry and Whiting (2005) and Wipfler et al. (2019).
Subtaxa
Anaidelia, Angaroptera, Aristoviidae, Blattogryllopterida, Blattogryllulus, Cabarzopterum, Cacurgoidea, Chalcorychus, Chorolimnobia, Dyadozoarium, Euchauliodidae, Grylloblattina, Haplopterum, Ischnoneuroidea, Issadonympha, Kaltanopterodes, Kargalopteridae, Kenguronympha, Kerbia, Kolvidelia, Lemmatophoridae (syn. Germanopriscidae, Lecoriidae, Ortaidae), Lemmatophorina, Liomopteridae (syn. Khosaridae, Climaconeuridae, Kargalellidae), Liomoptoides, Lodevoisadia, Madygenidelia, Mitinovia, Neoliomopterum, Palaeocixius (syn. Fabrecia), Permobaharellus, Protoperla boltoni, Protoperlina, Pseudoshurabia, Rachimentomon, Scalicidae, Sinonamuropteridae, Sukhonoptera, Sunopterites, Sylvamicropteron, Termoides, Uralotermes, Uralotermidae
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1914 | Grylloblattida Walker |
| 1914 | Grylloblattodea Walker |
| 1917 | Protoperla boltoni Lameere p. 164 |
| 1922 | Protoperla boltoni Handlirsch p. 109 |
| 1939 | Uralotermidae Zalessky p. 53 |
| 1965 | Uralotermitidae Emerson p. 11 |
| 1968 | Grylloblattodea Sharov |
| 2001 | Grylloblattida Wappler p. 96 |
| 2004 | Grylloblattodea Engel and Grimaldi |
| 2004 | Grylloblattodea Kukalová-Peck and Lawrence |
| 2005 | Grylloblattodea Terry and Whiting |
| 2006 | Grylloblattodea Arillo and Engel p. 3 |
| 2007 | Grylloblattida Béthoux |
| 2007 | Grylloblattodea Béthoux and Beckemeyer |
| 2009 | Grylloblattida Aristov p. 643 |
| 2014 | Grylloblattodea Misof et al. |
| 2015 | Grylloblattida Prokop et al. p. 30 |
| 2019 | Grylloblattodea Wipfler et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Grylloblattodea Walker 1914 [ice crawler]
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Fm. †Aristoviidae Storozhenko and Gröhn 2023
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Fm. †Euchauliodidae Riek 1974
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Invalid names: Grylloblattida Walker 1914 [replaced]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Westphalian A to the top of the Early/Lower Cenomanian or 318.70000 to 93.50000 Ma
Minimum age of oldest fossil (stem group age): 316.9 Ma
Minimum age of oldest fossil (stem group age): 316.9 Ma
Collections (138 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Westphalian A | France (Nord-Pas-de-Calais) | Climaconeura reumauxi (151687) | |
| Westphalian B - Westphalian C | China (Ningxia) | Sinonamuropteris ningxiaensis (139576) Sinonamuropteris ningxiaensis, Separatonerva longa, Separatonerva qilianshanensis, Combonerva granulata, Combonerva huangheensis, Tectonerva longovata, Sinonamuropteris zhongweiensis, Sinonamuropteris rugoverrucosa, Sinonamuropteris xiaheyanensis (102981) | |
| Westphalian C | China (Ningxia) | Sinonamuropteris ningxiaensis (179078) | |
| Westphalian D | USA (Pennsylvania) | Fabreciella pennsylvanica, Fabreciella allegheniensis (124504) | |
| Westphalian D | USA (Illinois) | Epideigma elegans (112630) | |
| Kasimovian - Gzhelian | Argentina (San Luis) | Paganzophlebia polyclada (127043) | |
| Gzhelian | Brazil (São Paulo) | Proedischia mezzalirai (117285) | |
| Gzhelian | France | Grylloblattida indet., Stenoneura maximi, Protoperla westwoodi, Palaeocixius fayoli, Palaeocixius antiquus (103635) Protoperla boltoni, Polyetes elegantissima, Fayoliella elongata, Protoblattina giardi (121979) | |
| Stephanian C | Germany | Liomopterites germanicus (133665) | |
| Asselian | Germany (Thüringer Wald) | Pictoborella clara, Liomopterum fuscatum, Liomopterites sperbersbachensis (128478) | |
| Asselian | Germany | Liomopteridae indet., Oborella sp., Oborella brauckmanni, Pictoborella germanica, Oborella monsjovisensis, Oborella moschelensis, Artinska glanensis (112604) Oborella sp. (234925) Pictoborella clara, Liomopterum fuscatum, Uralioma thuringiensis, Cabarzopterum magnificus (192084) Pictoborella germanica (222678) | |
| Asselian | USA (New Mexico) | Grylloblattida indet., Liomopteridae indet., Probnis fossor, Tapopterum populus, Depressopterum generator (124386) Liomopteridae indet. (124385) Liomopteridae indet., Probnis fossor, Tapopterum populus, Liomopterites annosus (124387) | |
| Asselian | Germany (Rhineland Palatinate) | Pictoborella germanica (233069) | |
| Asselian | Germany (Rheinland-Pfalz) | Oborella germanica, Oborella brauckmanni (130150) | |
| Asselian | Czech Republic | Jablonia aestiva, Turbopterum finum, Depressopterum senior, Depressopterum fragile, Depressopterum mancum, Drahania avia, Donopterum sp., Donopterum carpenteri, Donopterum nocturnum, Lioma moravica, Fumopterum largum, Cerasopterum oborianum, Cerasopterum longum, Cerasopterum extendum, Cerasopterum subtile, Tyrannopterum minimum, Havlatia annae, Zephyropterum lentum, Oborella matura, Oborella rusticana, Oborella inexpectata, Blania rotunda, Blania oviformis, Blania falsa, Lioma sp. (114186) | |
| Asselian | Germany (Rhineland-Palatinate) | Pictoborella germanica (156248) | |
| Sakmarian | Germany (Thüringer Wald) | Germanoprisca zimmermanni (160735) | |
| Artinskian | USA (Kansas) | Artinska sp., Lemmatophora sp., Probnis speciosa, Stemma sp., Estadia tripunctata (151412) Lemmatophora typa, Lisca minuta, Artinska clara, Artinska ovata, Lecorium elongatum (114152) Lemmatophora typa, Lisca minuta, Artinska clara, Artinska ovata, Lecorium elongatum, Paraprisca grandis (114151) Lemmatophora typa, Lisca minuta, Artinska clara, Probnis speciosa, Liomopterum ornatum, Artinska ovata, Tapopterum celsum, Chelopterum peregrinum (111319) Lemmatophora typa, Lisca minuta, Artinska clara, Probnis speciosa, Liomopterum ornatum, Orta ovata, Prisca fragilis, Lemmatophora delicosa, Lemmatophora hirsuta, Lemmatophora elongata, Lemmatophora anomala, Artinska medialis, Artinska gracilis, Artinska pecta, Artinska major, Artinska extensa, Estadia elongata, Estadia arcuata, Estadia tenuis, Lectrum anomalum, Lecorium elongatum, Stemma elegans, Stemma extensa, Probnis coriacea, Espira obscura, Stoichus elegans, Stoichus arcuatus, Stoichus minor, Stoichus tenuis, Stinus brevicubitalis, Liomopterum extensum, Horates elongatus, Prosaites compactus, Prosaites secundus, Lemmatophoridae indet., Estadia sp. (21979) Paraprisca fragilis, Artinska clara, Probnis speciosa, Liomopterum ornatum, Artinska ovata, Demopterum gracile, Lemmatophora typica, Lemmatophora minuta, Lemmatophora reducta, Artinska sellardsi, Lecorium elongatum, Sellardsia kansasensis, Sellardsia lecorioides, Paraprisca grandis, Paraprisca stigmatica, Liomopterum elongatum, Artinska inversa, Artinska pentamera, Artinska tillyardi, Artinska nova, Artinska simplex, Estadia altera, Estadia maculipennis, Estadia reducta, Estadia kansasia, Estadia aucta, Estadia banneria, Estadia confusa, Orta quadrifida, Orta reducta, Metalecorium confusum, Sellardsia separanda, Sellardsia abnormis, Liomopterum sellardsi, Semopterum venosum, Chelopterum peregrinum, Metalecorium anomalum, Lemmatophora cubitalis, Lemmatophora longipennis (111219) Paraprisca fragilis, Lemmatophora typa, Lisca minuta, Artinska clara, Liomopterum ornatum, Artinska ovata, Demopterum gracile, Lecorium elongatum, Liomopterum sellardsi, Semopterum venosum, Chelopterum peregrinum (111320) Paraprisca fragilis, Lemmatophora typa, Lisca minuta, Artinska clara, Liomopterum ornatum, Artinska ovata, Lecorium elongatum, Paraprisca grandis (111217) | |
| Artinskian | USA (Oklahoma) | Lemmatophora typa, Lisca minuta, Artinska clara, Probnis speciosa, Liomopterum ornatum (104826) Liomopterella pediaecetae (110869) Lisca minuta, Artinska ovata (104820) Midcopterum evidens (107405) Paraprisca fragilis, Lemmatophora typa, Artinska clara, Liomopterella pediaecetae (104828) Raaschia oklahomensis, Lisca minuta (104816) | |
| Artinskian | France (Languedoc) | Liomopteridae indet. A, Permobaharellus salagousensis (105044) | |
| Artinskian | Russian Federation (Perm) | Vilvaptera permyakovae (113348) | |
| Artinskian - Kungurian | Russian Federation (Kemerovo) | Liomopterella insignis (124599) | |
| Kungurian | Russian Federation (Kemerovo) | Parapermula sibirica (104951) Parapermula variabilis (124613) Sarbalopterum ignorabile (112708) | |
| Kungurian | Russian Federation (Perm) | Paraprisca solikamskensis, Artinska ufimica, Sylvaprisca tyulkinensis, Ufimoprisca mica, Ufimoprisca kamensis, Sylvaella ovalis, Tyulkinia bashkuevi (103639) | |
| Kungurian | China (Shanxi) | Sunopterites hejinensis (140456) | |
| Kungurian | Russian Federation (Komi republic) | Kaltanympha vorkutensis (122860) Liomopterum minor (127056) | |
| Kungurian | Russian Federation (Komi) | Sylvaprisca alba (105063) | |
| Kungurian | Russian Federation (Suksun) | Gurianovella silphidoides (122861) Khosarophlebia sylvaensis (113379) Paraprisca causaria (116652) Sylvaella paurovenosa, Parasylviodes tetracladus (116641) Sylvaprisca focaleata, Culiciforma formosa, Artinska infigurabilis, Euremisca splendens, Sylvafossor forcipatus, Khosarophlebia sylvaensis, Uralioma maxima, Ornaticosta novokshonovi, Parasylvaella umbra, Parasylvaella minor, Kungurolioma cancellata, Cerasopterum impurum, Tshekardites lientericus, Tshekardites calcomessor, Tshekardites curculiuculus, Tshekardites gracilis, Sylvamicropteron harpax (122753) Sylvaprisca gravis, Sylvaprisca forta, Culiciforma formosa, Artinska larisae, Artinska infigurabilis, Euremisca elegans, Euremisca splendens, Sylvaella vitiosa, Khosarophlebia sylvaensis, Uralioma variabilis, Sylvictor major, Parasylvaella umbra, Parasylvaella minor, Liomopterella kungurica, Tshekardites gracilis, Sylvaclinicus echinatus, Uraloprisca uralica, Tshekardeigma rasnitsyni, Chelopterum ultimum (108206) | |
| Kungurian | France (Languedoc) | Grylloblattida indet. (105053) Liomopteridae indet. B (105045) Oborella lodevensis (174412) | |
| Kungurian | South Africa (Gauteng) | Thaumatophora pronotalis (122863) | |
| Kungurian | Russian Federation | Artinska larisae, Uralioma variabilis, Sylvonympha tshekardensis (114207) Depressopterum bardum (124283) Grylloblattida indet., Liomopteridae indet., Liomopterites svetlanae (115214) Kolvidelia curta (173161) Paraprisca uralica, Euremisca splendens, Rachimentomon reticulatum, Termoides productus, Gurianovella silphidoides, Tshekardites comitialis, Rachimentomon affine (102504) Rachimentomon reticulatum (155825) Rachimentomon reticulatum, Angaroptera nicolaswachtleri (196228) Sylvaprisca gravis, Megorkhosa vernacula, Artinska larisae, Uralioma variabilis, Ornaticosta tchekardensis (107787) Tshekardites bardensis (122857) Uralotermes permianus (122754) Vorkutopterum monstrosum (127050) | |
| Kungurian | France | Depressopterum minutus (103637) Lodevoisadia coheni (228665) Lodevopterum angustus (103636) | |
| Kungurian | Russian Federation (Komi Republic) | Artinska vorkutensis (127046) Lemmatonympha gracilissima (122862) Liomopterella parakitiakensis (127052) Liomopterum minor (127054 127057) | |
| Roadian | Russian Federation (Kemerovo) | Abashevia suchovi (124598) Archiprobnis repens (122581) Climaconeurites asiaticus, Liomopterella vulgaris (122569) Kaltanympha thysanuriformis, Permonympha gracile, Parapermula sibirica, Parapermula variabilis, Climaconeurites asiaticus, Liomopterella vulgaris, Liomopterites comans, Liomopterites festivus (122567) Liomopterella vulgaris (122571) Liomopterites gracilis, Archiprobnis repens, Archiprobnis similis, Archiprobnis conjuncta, Parapermula sibirica, Parapermula variabilis, Parapermula nitens, Climaconeurites asiaticus, Ornaticosta magna, Ornaticosta amoena, Ornaticosta media, Ornaticosta nana, Liomopterella vulgaris, Liomopterella bella, Kaltanella lata, Liomopterites expletus, Liomopterites comans, Liomopterites junctus, Liomopterites festivus, Liomopterites accolis, Sibirella paucinervis (104704) Permonympha arcuata, Permonympha gracile, Archiprobnis repens, Parapermula sibirica, Climaconeurites asiaticus, Ornaticosta magna, Liomopterella vulgaris, Liomopterites expletus, Liomopterites comans, Liomopterites junctus, Sibirella paucinervis (122568) | |
| Roadian | Russian Federation | Camptoneura reticulata, Haplopterum majus (116456) Camptoneurites soyanensis (96860) | |
| Roadian | Russian Federation (Kirov) | Liomopterella kirovensis, Protomia proteus, Liomopterella kitiakensis, Liomopterella acuminata, Artinska novella (105058) Protomia proteus (105061) | |
| Roadian | Russian Federation (Kazan) | Tataronympha kamensis, Kazanella rotundipennis, Kazanella compressa (116803) | |
| Roadian | Russian Federation (Arkangelsk) | Artinska sojanensis, Sylvaprisca colorata, Camptoneurites soyanensis, Euremisca kazanica, Uraloprisca torta (104716) Parasylviodes sojanensis, Uraloprisca lisca, Paraphenopterum unicolor, Parapermula sojanensis, Liomopterites argutus, Kazanella mendosa, Sibirella variegata, Analiomopterites maculosus, Ornaticostella limbata, Sylvaella martynovi (104992) | |
| Wordian | South Africa | Paraliomopterum sp., Liomopterum connexus, Liomopterum daenerys, Colubrosopterum karooensis (226396) | |
| Wordian | Russian Federation (Kemerovo) | Kaltanopterodes vanus (124600) Liomopterites proximus, Paraliomopterum paulum (122566) Mitinovia dubia (124601) | |
| Wordian | Russian Federation (Udmurtia) | Kostovatoprisca acuminata, Paraliomopterum rectum (104750) Liomopterites novissimus (103502) Liomopterites novissimus, Parapermula tatarica (103538) | |
| Capitanian | Russian Federation (Evenki Autonomous Okrug) | Kerbia aptera, Kaltanympha ornata (105059) | |
| Capitanian | Russian Federation (Orenburg) | Dyadozoarium pachypus, Chalcorychus walchiae (127931) Kargalella subcostalis (104994) Kargalella subcostalis, Kargalella gibbosa, Liomopterites amanakicis, Expartolioma urzhumica, Urzhumskalicia kargalensis, Kargaloptera connexa, Kargalia sakmarica (104057) Kargalodes incerta (116610) Khosara permiakovae (116607) | |
| Wuchiapingian | Mongolia | Liomopterites exoticus, Mongolopermula adunca (97602) | |
| Wuchiapingian | Russian Federation (Vologda) | Liomopterites suhonensis, Isadia bipectinata, Sukhonoptera insolita, Sylvaella semicolorata, Liomofrater circumcisus, Falseshurabia transitoria, Issadonympha oculea, Kenguronympha lenta (104754) | |
| Changhsingian | South Africa (Eastern Cape) | Mioloptera sp., Mioloptera stuckenbergi (222191) | |
| Changhsingian | South Africa (Kwazulu-Natal) | Liomopterites ulterior (14357) Liomoptoides similis, Liomopterites meridionalis (14361) Mioloptera stuckenbergi (14350) Neoliomopterum picturatum (14362) | |
| Changhsingian | Kazakhstan (Zaisan) | Karaungiroptera maculosa (117256) Paraliomopterum alium, Paraliomopterum karaungirense (105062) | |
| Changhsingian | South Africa (Natal) | Mioloptera stuckenbergi, Liomoptoides similis, Mioloptoides andrei (124428) | |
| Changhsingian | Russian Federation (Vologda) | Expartalioma hirta (105041) | |
| Griesbachian | Russian Federation (Krasnoyarsk) | Tunguskaptera eximia (103638) | |
| Griesbachian | Russian Federation (Kemerovo) | Triasseuryptilon acostai, Khosara ultima (104713) | |
| Illyrian | Netherlands | Grylloblattida indet. (186555) | |
| Ladinian | Kyrgyzstan (Osh) | Gorochovia individua, Gorochovia minuta, Gorochovia anomala, Gorochovia fecunda, Gorochoviella conjuncta, Madygenophlebia bella, Anaidelia extrema, Pseudoshurabia pallidula (110763) Gorochovia individua, Gorochovia minuta, Gorochovia bifurca, Gorochoviella conjuncta, Pseudoliomopterites obscurus, Madygenophlebia bella, Madygenophlebia nana, Pseudoshurabia pallidula (105007) Gorochovia individua, Gorochovia minuta, Gorochovia bifurca, Gorochoviella conjuncta, Pseudoliomopterites obscurus, Pseudoliomopterites lucidus, Madygenophlebia nana (106090) Gorochovia individua, Madygenophlebia bella, Madygenophlebia primitiva, Micromadygenophlebia obscura, Pseudoshurabia pallidula, Madygenidelia conjuncta (106088) | |
| Carnian | Lesotho | Grylloblattida indet. (23075) | |
| Carnian | South Africa (Eastern Cape) | Euchauliodes distinctus (23142) | |
| Carnian | Argentina | Delopterum acostai (123021) | |
| Norian | Ukraine (Kharkiv) | Triassoprobnis humilis (105057) | |
| Early/Lower Toarcian | Germany (Mecklenburg-Vorpommern) | Nele jurassica (114993) | |
| Toarcian - Aalenian | China (Hunan) | Chorolimnobia ostera (114577) | |
| Callovian - Oxfordian | China (Nei Mongol) | Aristoviblattogryllus minor (230242) Plesioblattogryllus aristovi (140015) Plesioblattogryllus magnificus (113371) Plesioblattogryllus minor (113362) Plesioblattogryllus minor, Duoduo qianae (117610) Sinonele fangi, Sinonele hei, Sinonele phasmoides, Sinonele mini (113366) | |
| Tithonian | Mongolia (Dund-Gov’ Aymag) | Blattogryllulus mongolicus, Blattogryllulus lucidus, Bajanzhargalana magna (122321) | |
| Early/Lower Cenomanian | Myanmar | Aristovia daniili (172900) Sylvalitoralis cheni (178121) |