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Dryininae
Taxonomy
Dryininae was named by Haliday (1833). It is extant. Its type is Dryinus.
It was assigned to Proctotrypidae by Ashmead (1893), Kieffer (1905); and to Dryinidae by Rasnitsyn (1975), Ponomarenko (1981), Olmi and Bechly (2001), Engel (2005), Olmi and Guglielmino (2012), Olmi and Virla (2014), Olmi et al. (2014), Tribull (2015), Martynova et al. (2019), Perkovsky et al. (2019), Olmi et al. (2021).
It was assigned to Proctotrypidae by Ashmead (1893), Kieffer (1905); and to Dryinidae by Rasnitsyn (1975), Ponomarenko (1981), Olmi and Bechly (2001), Engel (2005), Olmi and Guglielmino (2012), Olmi and Virla (2014), Olmi et al. (2014), Tribull (2015), Martynova et al. (2019), Perkovsky et al. (2019), Olmi et al. (2021).
Subtaxa
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1833 | Dryininae Haliday |
| 1833 | Dryinini Haliday |
| 1893 | Dryininae Ashmead p. 80 |
| 1905 | Dryininae Kieffer p. 52 |
| 1975 | Dryininae Rasnitsyn p. 104 |
| 1975 | Dryinini Rasnitsyn p. 104 |
| 1981 | Dryininae Ponomarenko |
| 1981 | Dryinini Ponomarenko p. 119 |
| 2001 | Dryininae Olmi and Bechly p. 42 |
| 2005 | Dryininae Engel p. 486 |
| 2012 | Dryininae Olmi and Guglielmino p. 21 |
| 2014 | Dryininae Olmi and Virla p. 213 |
| 2014 | Dryininae Olmi et al. p. 705 |
| 2015 | Dryininae Tribull |
| 2019 | Dryininae Martynova et al. p. 3 |
| 2019 | Dryininae Perkovsky et al. p. 170 |
| 2021 | Dryininae Olmi et al. p. 417 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. Dryininae Haliday 1833
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G. Dryinus Latreille 1804
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†Dryinus acutifrons Martynova et al. 2019
†Dryinus alamellatus Olmi and Guglielmino 2011
†Dryinus antiquus Ponomarenko 1981
†Dryinus arakanesis Martynova et al. 2019
†Dryinus balticus Olmi 1984
†Dryinus bamar Martynova et al. 2019
†Dryinus brachytarsis Martynova et al. 2019
†Dryinus bruesi Olmi 1984
†Dryinus burmensis Martynova et al. 2019
†Dryinus canadensis Ponomarenko 1981
†Dryinus carsteni Olmi et al. 2022
†Dryinus chin Martynova et al. 2019
†Dryinus grimaldii Olmi 1995
†Dryinus hymenaeaphilus Olmi 1995
†Dryinus janzeni Olmi 2000
†Dryinus katyae Martynova et al. 2019
†Dryinus laminatus Martynova et al. 2019
†Dryinus latitarsis Martynova et al. 2019
†Dryinus maderai Guglielmino et al. 2018
†Dryinus mortuorum Brues 1933
†Dryinus muenchi Olmi and Bechly 2001
†Dryinus nadezhdae Martynova et al. 2019
†Dryinus palaeodominicanus Currado and Olmi 1983
†Dryinus palaeomexicanus Olmi 1995
†Dryinus parisei Martynova et al. 2019
†Dryinus poinari Olmi 1998
†Dryinus pristinus Olmi 1998
†Dryinus rasnitsyni Olmi and Guglielmino 2011
†Dryinus reifi Olmi and Bechly 2001
†Dryinus taron Martynova et al. 2019
†Dryinus velteni Olmi 2011
†Dryinus vetus Brues 1933
†Dryinus vetustus Olmi 1995
†Dryinus wunderlichi Olmi and Bechly 2001
Invalid names: Avodryinus Ponomarenko 1981 [synonym], Richardsidryinus Moczar 1965 [synonym]
G. †Harpactosphecion Haupt 1944
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†Harpactosphecion deletum Brues 1933
†Harpactosphecion filicorne Brues 1923
†Harpactosphecion filicornis Brues 1923
†Harpactosphecion gracile Brues 1933
†Harpactosphecion scheveni Olmi 2005
†Harpactosphecion sucinum Olmi 1987
G. †Hybristodryinus Engel 2005
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†Hybristodryinus anomalus Perkovsky et al. 2019
†Hybristodryinus castaneus Olmi et al. 2021
†Hybristodryinus concavifrons Perkovsky et al. 2019
†Hybristodryinus cretacicus Perkovsky et al. 2019
†Hybristodryinus karen Perkovsky et al. 2019
†Hybristodryinus kayin Perkovsky et al. 2019
†Hybristodryinus konbaung Perkovsky et al. 2019
†Hybristodryinus ligulatus Perkovsky et al. 2019
†Hybristodryinus magnificus Perkovsky et al. 2019
†Hybristodryinus mon Perkovsky et al. 2019
†Hybristodryinus moutesoe Tribull et al. 2020
†Hybristodryinus nalae Perkovsky et al. 2019
†Hybristodryinus pyu Perkovsky et al. 2019
†Hybristodryinus resinicolus Engel 2005
†Hybristodryinus shan Perkovsky et al. 2019
†Hybristodryinus viriosus Wang et al. 2021
†Hybristodryinus zaifui Chen et al. 2021
G. Pseudodryinus Olmi 1991
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†Pseudodryinus burmensis Olmi et al. 2014
†Pseudodryinus parisiensis Peinado et al. 2006
Invalid names: Dryinini Haliday 1833 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Cenomanian to the top of the Early/Lower Miocene or 100.50000 to 15.98000 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Collections (33 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Cenomanian | Myanmar | Dryinus burmensis, Dryinus katyae (161726) Dryinus carsteni (172900) Dryinus maderai, Hybristodryinus anomalus, Hybristodryinus concavifrons, Hybristodryinus cretacicus, Hybristodryinus karen, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus ligulatus, Hybristodryinus magnificus, Hybristodryinus mon, Hybristodryinus nalae, Hybristodryinus pyu, Hybristodryinus shan, Dryinus acutifrons, Dryinus arakanesis, Dryinus bamar, Dryinus brachytarsis, Dryinus chin, Dryinus laminatus, Dryinus latitarsis, Dryinus nadezhdae, Dryinus parisei, Dryinus taron, Hybristodryinus castaneus, Rasnitsynum burmense (194884) Dryinus taron (206272) Hybristodryinus cretacicus, Dryinus taron, Hybristodryinus zaifui, Hybristodryinus viriosus (165561) Hybristodryinus moutesoe (111688) Hybristodryinus resinicolus (123116) Hybristodryinus resinicolus, Dryinus maderai, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus nalae, Hybristodryinus pyu, Dryinus bamar, Dryinus katyae, Dryinus parisei (184692) Pseudodryinus burmensis (123053) | |
| Late/Upper Cenomanian | Russian Federation | Laberius antiquus (35051) | |
| Late/Upper Santonian | Russian Federation | Cretodryinus zherichini (122590) | |
| Middle Campanian | Canada (Alberta) | Avodryinus canadensis (122384) | |
| Ypresian | France (Oise) | Pseudodryinus parisiensis (123850) | |
| Priabonian | Ukraine | Dryinus janzeni (196704) Dryinus reifi, Dryinus wunderlichi (207953) | |
| Priabonian | Russian Federation | Harpactosphecion gracile, Dryinus janzeni (123419) | |
| Priabonian | Russian Federation (Kaliningrad) | Dryinus mortuorum, Harpactosphecion filicornis (156497) Dryinus muenchi, Dryinus reifi, Dryinus wunderlichi, Harpactosphecion filicorne, Dryinus bruesi, Dryinus janzeni (120814) Dryinus sp., Dryinus janzeni, Harpactosphecion filicornis (154633) Harpactosphecion filicorne (123773 124318) Harpactosphecion filicornis (123757) Palaeodryinus groehni, Harpactosphecion gracile, Dryinus janzeni (110735) Thaumatodryinus gracilis, Thaumatodryinus filicornis, Dryinus filicornis, Thaumatodryinus deletus, Lestodryinus mortuorum, Lestodryinus vetus (123927) | |
| Priabonian | Denmark | Alphadryinus balticus, Alphadryinus bruesi (123762) | |
| Early/Lower Miocene | Mexico (Chiapas) | Dryinus palaeomexicanus (124928) | |
| Burdigalian - Langhian | Dominican Republic | Dryinus grimaldii, Dryinus hymenaeaphilus, Dryinus pristinus, Dryinus vetustus, Pseudodryinus sucinus (122982) Dryinus palaeodominicanus (154719 166828) Dryinus palaeodominicanus, Alphadryinus sucinus (157305) Dryinus palaeodominicanus, Harpactosphecion scheveni (124846) Dryinus rasnitsyni, Harpactosphecion sucinum, Dryinus alamellatus (122474) Dryinus sp., Dryinus grimaldii, Dryinus palaeodominicanus, Dryinus poinari, Harpactosphecion sucinum (122472) |