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Exophytica

Insecta - Odonata

Taxonomy
Exophytica was named by Bechly (1996). It is extant.

It was assigned to Euanisoptera by Bechly et al. (1998), Ren et al. (2003), Bechly (2007).

Subtaxa
Cavilabiata, Gomphides
Sister taxa
Cavilabiata, Gomphides

Synonymy list
YearName and author
1996Exophytica Bechly
1998Exophytica Bechly et al.
2003Exophytica Ren et al.
2007Exophytica Bechly

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
infraclassHydropalaeopteraRohdendorf 1969
—Euhydropalaeoptera
—OdonatopteraMartynov 1932
RankNameAuthor
—Palaeodonatoptera
—Plesiodonatoptera
—Apodonatoptera
—Paneodonatoptera
—Neodonatoptera
—Euodonatoptera
—OdonatocladaBechly 1999
—Nodialata
—Discoidalia
—Stigmoptera
—Lobodonata
—Panodonata
orderOdonata
—Epiproctophora
—EuepiproctophoraBechly 1998
—Anisopteromorpha
—Trigonoptera
—Pananisoptera
—NeoanisopteraBechly 1998
suborderAnisopteraSelys-Longchamps and Hagen 1854
—Euanisoptera
—Exophytica
—Exophytica

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Exophytica Bechly 1996
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Unr. Cavilabiata Bechly 1996
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Unr. Chlorogomphida Bechly 1996
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Superfm. †Chlorogomphoidea Needham 1903
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Fm. †Araripechlorogomphidae Bechly and Ueda 2002
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G. †Araripechlorogomphus Bechly and Ueda 2002
Fm. †Mesochlorogomphidae Fleck et al. 2008
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G. †Hispanochlorogomphus Fleck et al. 2008
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†Hispanochlorogomphus rossi Fleck et al. 2008
G. †Mesochlorogomphus Fleck et al. 2008
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†Mesochlorogomphus crabbi Fleck et al. 2008
Unr. Cordulegastrida Bechly 1996
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Invalid names: Cordulegastridae Hagen 1875 [empty]
Fm. Cordulephyidae Tillyard 1917
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Subfm. †Neophyinae Tillyard and Fraser 1940
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G. Neophya Selys 1881
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†Neophya legrandi Nel and Fleck 2014
G. †Palaeophya Petrulevičius and Nel 2009
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†Palaeophya argentina Petrulevičius and Nel 2009
Fm. †Cordulibellulidae Sroka et al. 2023
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G. †Cordulibellula Sroka et al. 2023
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†Cordulibellula inopinata Sroka et al. 2023
Unr. Cristotibiata Bechly 1996
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Unr. Brachystigmata Bechly 1996
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Fm. †Nannogomphidae Bechly 1996
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G. †Nannogomphus Handlirsch 1906
G. †Sinahemeroscopus Huang and Nel 2010
Unr. Paucipostnodalia Bechly and Ueda 2002
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Unr. Eubrachystigmata Bechly 1996
Fm. †Prohemeroscopidae Bechly and Ueda 2002
Invalid names: Neopetaliidae Tillyard and Fraser 1940 [empty]
Fm. †Gallophlebiidae Nel et al. 2015
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G. †Gallophlebia Nel et al. 2015
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†Gallophlebia magnifica Nel et al. 2015
Fm. †Libanocorduliidae Azar et al. 2019
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G. †Libanocordulia Azar et al. 2019
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†Libanocordulia debiei Azar et al. 2019
Fm. †Nodalulaidae Lin et al. 2007
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G. †Nodalula Lin et al. 2007
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†Nodalula dalinghensis Lin et al. 2007
Fm. †Urolibellulidae Zeiri et al. 2015
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G. †Kohlslibellula Nel 2020
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†Kohlslibellula lini Nel 2020
G. †Urolibellula Zeiri et al. 2015
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†Urolibellula eocenica Zeiri et al. 2015
Unr. Gomphides Bechly et al. 1998
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Fm. †Araripegomphidae Bechly 1996
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G. †Araripegomphus Nel and Paicheler 1994
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†Araripegomphus andreneli Bechly 1998
†Araripegomphus cretacicus Nel and Paicheler 1994
†Araripegomphus ismari Nel and Ribeiro 2022
†Araripegomphus shai Zheng et al. 2018
Unr. Desmoproctida Bechly 1996
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Unr. Oligophlebiata Bechly 2003
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Unr. Brevicubitalia Bechly 2003
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Unr. Gomphida Bechly 1996
Fm. †Burmagomphidae Zheng et al. 2017
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G. †Burmagomphides Zheng et al. 2017
G. †Satelitala Petrulevičius 2017
Fm. †Libanogomphidae Azar and Nel 2023
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G. †Libanogomphus Azar and Nel 2023
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†Libanogomphus lionelcavini Azar and Nel 2023
Fm. †Paraburmagomphidae Zheng et al. 2018
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G. †Paraburmagomphides Zheng et al. 2018
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†Paraburmagomphides zhaoi Zheng et al. 2018
Invalid names: Hagenioidea Tillyard and Fraser 1940 [empty], Lindeniidae Jacobson and Bianchi 1905 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitino
Form: roller-shapedo
Environment: freshwater, terrestrialo
Locomotion: fast-movingo
Life habit: volanto
Diet: carnivoreo
Diet 2: insectivoreo
Vision: well-developedo
Dispersal: watero
Created: 2011-03-07 03:41:10
Modified: 2011-03-06 10:41:10
Source: o = order
Reference: Ren and Guo 1996

Age range

Maximum range based only on fossils: base of the Bathonian to the top of the Chibanian or 168.20000 to 0.12900 Ma
Minimum age of oldest fossil (stem group age): 165.3 Ma

Collections (156 total)


Time interval Ma Country or state Original ID and collection number
Bathonian168.2 - 165.3United Kingdom (Oxfordshire) Libellula westwoodii (140495)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Juralibellula ningchengensis, Daohugoulibellula lini (113371)
Early/Lower Tithonian149.2 - 145.06Germany (Bavaria) Juracordulia schiemenzi, Nannogomphus bavaricus (122586) Nannogomphus bavaricus (139876)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Agrion gracile, Agrion vetustum (123985) Agrion vetustum (123906) Nannogomphus bavaricus (123144) Nannogomphus buergeri (139551) Prohemeroscopus jurassicus, Prohemeroscopus kuehnapfeli (152883) Prohemeroscopus kuehnapfeli (227117)
Early/Lower Tithonian149.2 - 145.06Germany Nannogomphus buergeri (122880)
Early/Lower Berriasian143.1 - 141.0United Kingdom (England) Araripelibellula britannica (104006)
Berriasian143.1 - 137.05United Kingdom (England) Homoptera indet. (127481)
Middle Berriasian141.0 - 139.1United Kingdom (England) Araripelibellula britannica (119561)
Late/Upper Berriasian139.1 - 137.05United Kingdom (England) Araripelibellula britannica (103833)
Late/Upper Hauterivian132.6 - 125.77China (Hebei) Jibeigomphus xinboensis (154962)
Late/Upper Hauterivian132.6 - 125.77United Kingdom (Surrey) Cretaneophya strevensi (74973)
Late/Upper Hauterivian132.6 - 125.77United Kingdom (England) Valdicordulia wellsorum (106281)
Barremian125.77 - 121.4China (Nei Mongol) Sinogomphus taushanense (154968)
Early/Lower Barremian125.77 - 121.4United Kingdom (England) Cretaneophya strevensi, Mesochlorogomphus crabbi (45193)
Early/Lower Barremian125.77 - 121.4Spain Hispanochlorogomphus rossi (123437) Ilerdaegomphus pallerolae (152691) Palaeaeschna pallerolae (152686)
Late/Upper Barremian125.77 - 121.4Spain (Cuenca) Ilerdaegomphus torcae (152992)
Late/Upper Barremian125.77 - 121.4China (Nei Mongol) Sinahemeroscopus magnificus (125655)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Gomphus sp., Liogomphus yixianensis, Gomphus biconvexus (148368) Guocordulia boreala (111725) Liogomphus yixianensis, Mesocordulia boreala (138436) Nodalula dalinghensis (103832) Rencordulia sinica (151524) Sopholibellula eleganta, Sopholibellula amoena (111722)
Aptian121.4 - 113.2China (Ningxia) Hemeroscopus baissicus (237978)
Aptian121.4 - 113.2China (Liaoning) Abrohemeroscopus mengi (105248)
Aptian121.4 - 113.2Russian Federation (Buryatia) Hemeroscopus baissicus (105536 105969)
Aptian121.4 - 113.2Mongolia (Kobdo aimak) Eocordulia cretacea (104054)
Late/Upper Aptian119.57 - 113.2China (Beijing Shi) Hemeroscopus baissicus (106965)
Late/Upper Aptian119.57 - 113.2Brazil (Ceará) Araripechlorogomphus muratai (228900) Araripechlorogomphus muratai, Araripegomphus andreneli, Araripephlebia mirabilis (122926) Araripegomphus andreneli (122918) Araripegomphus andreneli, Araripephlebia mirabilis (122922 122925) Araripegomphus andreneli, Araripephlebia mirabilis, Araripegomphus hanseggeri, Cratolindenia knuepfae (122557) Araripegomphus andreneli, Araripephlebia mirabilis, Cratocordulia borschukewitzi, Araripelibellula martinsnetoi (122919) Araripegomphus cretacicus (125509) Araripegomphus ismari (217013) Araripelibellula martinsnetoi (119562) Araripelibellula sennlaubi (215116) Cratocordulia borschukewitzi, Araripelibellula martinsnetoi (122553) Valdiphlebia sennlaubi (227841)
Albian113.2 - 100.5South Korea Hemeroscopus baissicus (184781)
Albian113.2 - 100.5South Korea (Gyeongsangnam-do) Hemeroscopus baissicus (164493)
Middle Albian110.1 - 106.3China (Gansu) Hemeroscopus baissicus (164492)
Early/Lower Cenomanian100.5 - 93.9Myanmar Burmagomphides electronica, Araripegomphus shai (161726) Burmalindenia imperfecta (123053) Paraburmagomphides zhaoi (196204)
Early/Lower Cenomanian100.5 - 93.9France Gallophlebia magnifica (162748)
Late/Upper Cenomanian100.5 - 93.9Lebanon Libanocordulia debiei (196672) Libanogomphus lionelcavini (229466)
Turonian93.9 - 89.8Kazakhstan Palaeolibellula zherikhini (119502)
Late/Upper Campanian83.6 - 72.2Canada (Alberta) Cordualadensa acorni (241744)
Late/Upper Paleocene59.24 - 56.0Argentina (Jujuy) Curviarculia lamasi (114155) Jujusia maizgorda, Curviarculia delicata, Palaeomacromia multicellulata (113004) Palaeophya argentina, Austrolibellula noroestenia (106995)
Ypresian56.0 - 48.07Denmark Molercordulia karinae (105202) Molertrum eburneum (157040)
Ypresian56.0 - 48.07France (Oise) Eurypalpida indet. (123850)
Ypresian56.0 - 48.07Italy Bolcathemis nervosa, Bolcacordulia paradoxa (122230)
Ypresian56.0 - 48.07Canada (British Columbia) Auroradraco eos (103205)
Ypresian56.0 - 48.07Argentina (Chubut) Satelitala soberana (133066)
Bridgerian50.5 - 46.2USA (Wyoming) Libellulidae indet. (121322)
Bridgerian50.5 - 46.2USA (Colorado) Kohlslibellula lini (192072) Stenogomphus carletoni (114681)
Bridgerian50.5 - 46.2USA (Utah) Cordulibellula inopinata (230563)
MP 1148.07 - 41.03Germany (Hessen) Corduliidae indet. (120798)
Uintan46.2 - 39.7USA (Colorado) Libellulidae indet. (156492) Urolibellula eocenica (173817)
Priabonian37.71 - 33.9Poland Gomphoides occulta (120811)
Priabonian37.71 - 33.9United Kingdom (England) Corduliidae indet. A, Neophya legrandi, Eomacrodiplax incompleta (123960) Corduliidae indet. B, Corduliidae indet. C, Eomacrodiplax incompleta (21976) Neophya legrandi (113829) Pantalinae indet. (207259)
Priabonian37.71 - 33.9United Kingdom (Isle of Wight) Corduliidae indet. A (157667)
Early/Lower Oligocene33.9 - 27.3USA (Oregon) Somatochlora oregonica (117157)
Early/Lower Oligocene33.9 - 27.3Germany Petalura acutipennis (113964)
MP 2333.9 - 27.3France Tauriphila cerestensis (123913)
Rupelian33.9 - 27.3France (Alpes-de-Haute) Gomphinae indet. A, Paleotauriphila sp. (114016) Paleotauriphila marquesi (155208)
Rupelian33.9 - 27.3France (Auvergne) Libellula minuscula (127630)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Libellulidae sp. A, Palaeotramea cellulosa, Paleotramea cellulosa (122010)
Chattian27.3 - 23.04France (Provence-Alpes-Côte d'Azur) Libellulidae larva, Libellula martini (122487)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Libellula requieniana, Libellula pourqueryi (153412) Libellulidae indet. (157194) Libellulidae larva (113712 122501) Lindeniidae indet., Caussanelia papaziani, Palaeotramea aquisextana, Sympetrum bigoti (127415) Paleotramea aquisextana (177171)
Late/Upper Oligocene27.3 - 23.04France (Aix-en-Provence) Jeanlegrandia oligocenica (114117)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Libellulidae indet., Ictinogomphus engelorum (71942) Lindeniinae indet. (211302)
MP 3027.3 - 23.04Germany Libellula cellulosa (123945)
MP 3027.3 - 23.04Germany (Nordrhein-Westfalen) Libellula cellulosa (138696)
Chattian - Aquitanian27.3 - 20.45Germany (Sachsen) Ictinogomphus sp. (26765)
Aquitanian23.04 - 20.45Germany Libellula sp., Trameobasileus moguntiacus (123872) Libellulidae indet. (237236)
Aquitanian23.04 - 20.45Türkiye Libellulidae indet., Lindeniinae indet., Trameini indet., Paleotramea aquisextana, Paleotramea fasciata, Paleotramea acutibucula (123307)
Early/Lower Miocene23.04 - 15.98Spain Lindeniidae indet. (155648)
Early/Lower Miocene23.04 - 15.98Mexico Erpetogomphus shii (200059)
Early/Lower Miocene23.04 - 15.98Spain (Castellon) Libellulidae indet. (140796)
Burdigalian20.45 - 15.98Czechia Onychothemis rihai (181765)
Burdigalian20.45 - 15.98China (Shandong) Miopetalura shanwangica (128551)
Burdigalian20.45 - 15.98Austria Libellulidae indet. (139942)
Orleanian20.0 - 15.98Germany (Baden-Wurttemberg) Parabrachydiplax miocenica (106002)
Hemingfordian18.5 - 16.3USA (California) Orthemis ferruginea (106431) Orthemis sp. (123022 123023 123025 123042)
MN 416.9 - 15.98Czechia Libellulidae indet., Libellulidae indet. larvae, Libellula knetti (157202) Onychothemis rihai (105501) Prorhyothemis bubiki (105500)
Langhian15.98 - 13.82USA (Washington) Miocordulia latipennis (140652)
MN 515.98 - 13.82Germany (Baden-Württemberg) Randecktrum ebi (215189)
Middle Miocene15.98 - 11.63Japan Gomphidae indet. (153619)
Middle Miocene15.98 - 11.63Australia (New South Wales) Corduliidae indet., Austrogomphus sp. (224044)
Middle Miocene15.98 - 11.63China (Hebei) Lanthus wuluogongensis (128554)
Middle Miocene15.98 - 11.63Russian Federation Pantalinae indet., Sympetrum sp. (105571)
Badenian13.82 - 12.8Austria Gomphidae indet. (131964) Ictinogomphus hassleri (184614)
Sarmatian12.8 - 11.63Croatia Libellula platyptera (117246) Lithemis lejeunecarpentieri, Oligocaemia imperfecta (145135) Protopaltothemis hageni (113698)
Sarmatian12.8 - 11.63Slovenia Sloveniatrum robici (103103)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Cordulia sp. (115305) Libellula oeningensis (151877) Lindenia heeri (117252)
Vallesian11.63 - 8.725Spain Trithemis pseudodistanti (114010)
Late/Upper Miocene11.63 - 5.333Poland Libellula sieboldiana, Libellula kieseli, Libellula pannewitziana (138936)
Late/Upper Miocene11.63 - 5.333Ukraine (Kerch) Libellula ukrainensis (127948)
Vallesian - Turolian11.63 - 5.09Spain (Lleida) Libellulidae indet. (154246)
Turolian8.725 - 5.09France (Ardèche) Gomphidae indet., Corduliinae indet., Somatochlora brisaci (113893) Libellulidae indet., Miorhodopygia andancensis, Cellulocordulia fasciata (156164)
Messinian7.246 - 5.333France (Cantal) Gomphidae indet., Celithemis cantalensis, Sympetrum cantalensis (122195)
Messinian7.246 - 5.333Italy Corduliinae indet., Epitheca miocenica, Epitheca annae (113886) Libellula sp., Sympetrum italicum, Sympetrum krzeminskii, Sympetrum elongatum, Libellula mediterranea, Tramea miocenica, Pisaurum coloratum, Deielia sarae, Trapezostigma sp., Trapezostigma barbaresii, Libellula adriatica, Celithemis zavattinii, Sympetrum marinum (157306) Libellulidae indet. (157203)
Messinian7.246 - 5.333Italy (Piedmont) Oryctodiplax gypsorum (124409) Oryctodiplax sp., Oryctodiplax gypsorum (113998)
Messinian7.246 - 5.333Italy (Tuscany) Libellulidae indet., Diplax sp. (113751)
Pliocene5.333 - 2.58India (Jharkhand) Libellulidae indet. (214996)
Blancan4.7 - 1.4USA (Texas) Libellulidae indet. (154909)
Piacenzian3.6 - 2.58USA (California) Gomphidae indet. (162797) Libellulidae indet. (162798)
Piacenzian3.6 - 2.58Germany (Niedersachsen) Gomphidae indet., Priscogomphus subhercynicus (109196)
Gelasian2.58 - 1.8France Libellula eusebioi (113996)
Chibanian0.774 - 0.129Japan Sieboldius albardae (211426)