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Bethylinae

Insecta - Hymenoptera - Bethylidae

Synonymy list
YearName and author
1840Bethylinae Halliday
1874Bethyllides Westwood p. 156
1893Bethylinae Ashmead p. 27
1905Bethylinae Kieffer p. 227
1938Bethylinae Statz p. 107
1988Bethylinae Sorg p. 71
2004Bethylinae De Ploëg and Nel p. 75
2012Bethylinae Azevedo and Azar p. 216
2014Bethylinae Ramos et al. p. 205
2016Bethylinae Cockx et al. p. 4
2018Bethylinae Azevedo et al. p. 25
2020Bethylinae Colombo et al.
2022Bethylinae Brazidec and Perrichot p. 2
2023Bethylinae Barbosa and Melo p. 171
2023Bethylinae Brazidec et al. p. 365

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
Dicondylia
Paranotalia
subclassPterygota()
RankNameAuthor
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Hymenopterida
orderHymenoptera
Unicalcarida
suborderApocritaGerstaecker 1867
Aculeata()
superfamilyChrysidoidea
familyBethylidaeForster 1856
subfamilyBethylinaeHalliday 1840
subfamilyBethylinaeHalliday 1840

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Bethylinae Halliday 1840
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G. Bethylus Latreille 1802
G. †Crassibethylus Barbosa and Melo 2023
+
Crassibethylus dominicanus Barbosa and Melo 2023
G. †Cretobethylellus Rasnitsyn 1990
+
Cretobethylellus lucidus Rasnitsyn 1990
G. Eupsenella Westwood 1874
+
Eupsenella aulax Ramos and Azevedo 2014
Eupsenella eocenica De Ploëg and Nel 2004
Eupsenella klesoviana Ramos and Azevedo 2014
Eupsenella rossica Ramos and Azevedo 2014
Eupsenella yantarnica Ramos and Azevedo 2014
Invalid names: Eupsenella neoeocenica Ramos and Azevedo 2018 [nomen nudum], Eupsenella neohongi Ramos and Azevedo 2018 [nomen nudum], Fushunochrysites Hong 2002 [nomen nudum], Protobethylus De Ploëg and Nel 2004 [synonym], Sinibethylus Hong 2002 [nomen nudum]
G. Goniozus Förster 1856
+
Goniozus antiquus Olmi 2005
Goniozus contractus Brues 1933
Goniozus copalinus Meunier 1916
Goniozus cotyi Brazidec and Perrichot 2022
Goniozus definitus Ramos and Azevedo 2014
Goniozus respectus Sorg 1988
Invalid names: Messoria Meunier 1916 [synonym], Perisierola Kieffer 1914 [synonym], Pterosclerogibba Olmi 2005 [synonym]
G. Lytopsenella Kieffer 1911
+
Lytopsenella baltica Ramos and Azevedo 2014
Lytopsenella crastina Brues 1923
Lytopsenella maritima Ramos and Azevedo 2014
Lytopsenella setigera Brues 1923
Lytopsenella simplex Brues 1923
G. †Nucifrangibulum Cockx et al. 2016
+
Nucifrangibulum carentonensis Cockx et al. 2016
G. Prosierola Kieffer 1905
+
Prosierola submersa Brues 1933
G. Sierola Cameron 1881
+
Sierola colomboi Brazidec and Perrichot 2022
Sierola hastata Sorg 1988
Sierola rovniana Ramos and Azevedo 2014
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Aptian to the top of the Holocene or 125.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 113.0 Ma

Collections (22 total)


Time interval Ma Country or state Original ID and collection number
Aptian125.0 - 113.0Russian Federation Cretobethylellus lucidus (124811)
Early/Lower Cenomanian99.6 - 93.5France (Poitou-Charentes) Nucifrangibulum carentonensis (161514)
Ypresian56.0 - 47.8China (Liaoning) Fushunochrysites eocenicus (124530) Sinibethylus eocenicus (124820)
Ypresian56.0 - 47.8France (Oise) Protobethylus eocenicus (123850)
Bridgerian50.3 - 46.2USA (Colorado) Eupsenella sp. (229578)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Eupsenella aulax, Lytopsenella baltica, Lytopsenella maritima (160471) Eupsenella crastina (127170) Goniozus definitus (120610) Lytopsenella crastina (124318) Lytopsenella kerneggeri (138800) Lytopsenella kerneggeri, Eupsenella rossica, Eupsenella yantarnica (122591) Sierola simplex, Sierola setigera, Sierola crastina, Eupsenella setigera, Eupsenella simplex, Eupsenella crastina, Prosierola submersa, Perisierola contracta (123927)
Priabonian38.0 - 33.9Russian Federation Lytopsenella crastina (173264)
Priabonian38.0 - 33.9Denmark Sierola hastata (123762)
Priabonian38.0 - 33.9Ukraine Lytopsenella kerneggeri, Eupsenella aulax, Eupsenella klesoviana, Sierola rovniana (122997)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Goniozus cotyi (225741)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Crassibethylus dominicanus (203120) Goniozus respectus (146557) Pterosclerogibba antiqua (122472)
Langhian15.97 - 13.82China (Fujian) Sierola colomboi (220273)
Holocene0.0117 - 0.0Madagascar Messoria copalina (151372)