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Osmyloidea

Insecta - Neuroptera

Taxonomy
Osmyloidea was named by Leach (1815). It is extant. Its type is Osmylus.

It was assigned to Neuroptera by Martins-Neto (2002); to Hemerobiiformia by Engel and Grimaldi (2008); and to Euneuroptera by Engel et al. (2018).

Synonymy list
YearName and author
1815Osmyloidea Leach
2002Osmyloidea Martins-Neto p. 38
2008Osmyloidea Engel and Grimaldi
2018Osmyloidea Engel et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
superorderNeuropterida
—Metaxyneuroptera
—Eidoneuroptera
orderNeuroptera
—Euneuroptera
superfamilyOsmyloideaLeach 1815
superfamilyOsmyloideaLeach 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Osmyloidea Leach 1815
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Fm. †Archeosmylidae Riek 1953
+
G. †Archeosmylus Riek 1953
+
†Archeosmylus costalis Riek 1955
G. †Babykamenia Ponomarenko and Shcherbakov 2004
+
†Babykamenia eskovi Ponomarenko and Shcherbakov 2004
G. †Lithosmylidia Riek 1955
+
†Lithosmylidia baronne Lambkin 1988
†Lithosmylidia lineata Riek 1955
†Lithosmylidia parvula Riek 1955
G. †Madygoneura Khramov 2024
+
†Madygoneura elongata Khramov 2024
G. †Osmylotriasia Khramov 2024
+
†Osmylotriasia superba Khramov 2024
G. †Triasella Khramov 2024
+
†Triasella ovata Khramov 2024
G. †Momonokus Khramov et al. 2023
+
†Momonokus obscurus Khramov et al. 2023
Fm. Nevrorthidae Nakahara 1958
+
G. †Balticoneurorthus Wichard 2016
+
†Balticoneurorthus elegans Wichard 2016
G. †Cretarophalis Wichard 2017
+
G. †Electroneurorthus Wichard et al. 2010
+
†Electroneurorthus malickyi Wichard et al. 2010
G. †Girafficervix Du et al. 2023
+
†Girafficervix baii Du et al. 2023
Invalid names: Palaeoneurorthus Du et al. 2023 [replaced]
G. †Palaeoneurorthus Wichard 2009
+
†Palaeoneurorthus eocaenus Wichard 2016
†Palaeoneurorthus groehni Wichard et al. 2010
G. †Proberotha Krüger 1923
+
†Proberotha dichotoma Wichard 2016
†Proberotha prisca Krüger 1923
G. †Rophalis Pictet 1854
+
†Rophalis relicta Pictet 1854
G. †Sisyroneurorthus Nakamine et al. 2023
+
†Sisyroneurorthus aspoeckorum Nakamine et al. 2023
Fm. Osmylidae Leach 1815 [lance lacewing]
+
G. †Archaeosmylidia Makarkin et al. 2014
+
†Archaeosmylidia fusca Makarkin et al. 2014
G. †Dimidiosmylus Khramov 2014
+
†Dimidiosmylus ramosus Khramov 2014
G. †Dzhidosmylus Khramov 2020
+
†Dzhidosmylus solus Khramov 2020
Subfm. Eidoporisminae Esben-Petersen 1917
+
G. †Stenosmylina Jepson et al. 2009
+
†Stenosmylina medialis Jepson et al. 2009
G. †Erlikosmylus Khramov 2014
+
†Erlikosmylus obscurus Khramov 2014
Subfm. Gumillinae Navas 1912
+
G. †Allotriosmylus Yang et al. 2010
+
†Allotriosmylus uniramosus Yang et al. 2010
G. †Enodinympha Ren and Engel 2007
+
†Enodinympha imperfecta Ren and Engel 2007
†Enodinympha translucida Ren and Engel 2007
G. †Epiosmylus Panfilov 1980
+
†Epiosmylus longicornis Panfilov 1980
†Epiosmylus longus Khramov 2017
†Epiosmylus panfilovi Ren and Yin 2002
G. Gumilla Navas 1912
G. †Kolbasinella Khramov 2014
+
†Kolbasinella elongata Khramov 2014
G. †Nilionympha Ren and Engel 2007
+
†Nilionympha pulchella Ren and Engel 2007
†Nilionympha shantouensis Li et al. 2018
G. †Nuddsia Menon and Makarkin 2008
+
†Nuddsia longiantennata Menon and Makarkin 2008
†Nuddsia magnifica Myskowiak et al. 2016
†Nuddsia repatriata Martins-Neto 2010
†Nuddsia simplex Li et al. 2024
Invalid names: Burmaleon Myskowiak et al. 2016 [synonym]
G. †Osmylochrysa Jepson et al. 2012
+
†Osmylochrysa anomala Jepson et al. 2012
†Osmylochrysa fragilis Jepson et al. 2012
†Osmylochrysa navasia Khramov 2020
G. †Tenuosmylus Wang et al. 2009
+
†Tenuosmylus brevineurus Wang et al. 2009
Invalid names: Epiosmylinae Panfilov 1980 [synonym]
G. †Karaosmylus Makarkin 1990
+
†Karaosmylus latus Panfilov 1980
Invalid names: Parosmylus Panfilov 1980 [replaced]
Subfm. Kempyninae Carpenter 1943
+
G. †Arbusella Khramov 2014
+
†Arbusella bella Khramov 2014
†Arbusella magna Khramov et al. 2017
†Arbusella platyptera Ma and Wang 2022
G. †Cretosmylus Makarkin 1990
+
†Cretosmylus sibiricus Makarkin 1990
G. †Euporismites Tillyard 1916
+
†Euporismites balli Tillyard 1916
G. †Jurakempynus Wang et al. 2011
+
†Jurakempynus arcanus Khramov 2014
†Jurakempynus bellatulus Wang et al. 2011
†Jurakempynus epunctatus Wang et al. 2011
†Jurakempynus loculosus Ma et al. 2020
†Jurakempynus sinensis Wang et al. 2011
†Jurakempynus sublimis Khramov 2014
G. †Kempynosmylus Makarkin 2014
+
†Kempynosmylus zherikhini Makarkin 2014
G. †Mirokempynus Ma et al. 2020
+
G. †Ponomarenkius Khramov et al. 2017
+
†Ponomarenkius excellens Khramov et al. 2017
G. †Sauktangida Khramov 2014
+
†Sauktangida aenigmatica Khramov 2014
G. †Kubekius Khramov 2017
+
†Kubekius multiramosus Khramov 2017
G. †Mesosmylidus Jepson et al. 2012
+
†Mesosmylidus vulgaris Jepson et al. 2012
Subfm. †Mesosmylininae Bode 1953
+
G. †Mesosmylina Bode 1953
+
†Mesosmylina angusta Khramov 2014
†Mesosmylina exornata Bode 1953
†Mesosmylina falcifera Ansorge 1996
†Mesosmylina mongolica Ponomarenko 1984
†Mesosmylina shurabica Khramov 2014
†Mesosmylina sibirica Ponomarenko 1985
Subfm. Osmylinae Leach 1815
+
G. †Lithosmylus Carpenter 1943
+
†Lithosmylus columbianus Cockerell 1908
G. Osmylus Latreille 1802
G. †Vetosmylus Ma et al. 2020
+
†Vetosmylus maculosus Ma et al. 2020
†Vetosmylus tentus Ma et al. 2020
G. †Osmylopsis Handlirsch 1906
+
†Osmylopsis duplicata Giebel 1856
+
Invalid names: Pterinoblattina binneyi Scudder 1886 [objective synonym]
G. †Palaeothyridosmylus Wang et al. 2009
+
G. †Petrushevskia Martynova 1958
+
†Petrushevskia borisi Martynova 1958
G. †Pronymphites Panfilov 1980
+
†Pronymphites elegans Panfilov 1980
Subfm. Protosmylinae Krüger 1913
+
G. †Juraheterosmylus Wang et al. 2010
+
†Juraheterosmylus antiquatus Wang et al. 2010
†Juraheterosmylus astictus Wang et al. 2010
†Juraheterosmylus minor Wang et al. 2010
G. †Jurosmylus Makarkin and Archibald 2005
+
†Jurosmylus atalantus Panfilov 1980
†Jurosmylus parvulus Khramov 2014
Invalid names: Mesosmylus Panfilov 1980 [replaced]
G. †Myanmarosmylus Li et al. 2024
+
†Myanmarosmylus wintertoni Li et al. 2024
G. †Osmylidia Cockerell 1908
+
†Osmylidia donnae Makarkin et al. 2021
†Osmylidia glastrai Makarkin et al. 2021
†Osmylidia picta Hagen 1856
†Osmylidia requieta Scudder 1890
†Osmylidia taliae Makarkin et al. 2021
Invalid names: Oligosmylus Krüger 1913 [objective synonym], Protosmylus Krüger 1913 [synonym]
G. †Protosmylina Jepson et al. 2009
+
†Protosmylina bifasciata Jepson et al. 2009
G. †Pseudosmylidia Makarkin 2017
+
†Pseudosmylidia relicta Makarkin 2017
G. †Sogjuta Martynova 1958
+
†Sogjuta shartegica Khramov 2011
†Sogjuta speciosa Martynova 1958
G. †Scapoptera Panfilov 1980
+
†Scapoptera recta Panfilov 1980
Subfm. Spilosmylinae Kruger 1913
+
G. †Ensiosmylus Khramov 2014
+
†Ensiosmylus acutus Khramov 2014
G. †Imanosmylus Makarkin 1994
+
†Imanosmylus ussuriensis Makarkin 1994
G. †Zakamnosmylus Khramov 2020
+
†Zakamnosmylus elongatus Khramov 2020
G. †Stenochrysa Jepson et al. 2012
+
†Stenochrysa gradata Jepson et al. 2012
G. †Tengriosmylus Khramov 2014
+
†Tengriosmylus magnificus Khramov 2014
G. †Tetanoptilon Bode 1953
+
Fm. †Saucrosmylidae Ren and Yin 2003 [elegant lance lacewing]
+
G. †Bellinympha Wang et al. 2010
+
†Bellinympha dancei Wang et al. 2010
†Bellinympha filicifolia Wang et al. 2010
G. †Daohugosmylus Liu et al. 2014
+
†Daohugosmylus castus Liu et al. 2014
G. †Huiyingosmylus Liu et al. 2013
+
†Huiyingosmylus bellus Liu et al. 2013
G. †Laccosmylus Ren and Yin 2003
+
†Laccosmylus calophlebius Ren and Yin 2003
†Laccosmylus cicatricatus Fang et al. 2018
†Laccosmylus latizonus Fang et al. 2018
G. †Lebanosmylus Azar and Nel 2022
+
†Lebanosmylus leae Azar and Nel 2022
G. †Rudiosmylus Ren and Yin 2003
+
†Rudiosmylus ningchengensis Ren and Yin 2003
G. †Saucrosmylus Ren and Yin 2003
+
†Saucrosmylus sambneurus Ren and Yin 2003
G. †Thaumatomerobius Ponomarenko 1985
+
†Thaumatomerobius mirabilis Ponomarenko 1985
G. †Ulrikezza Fang et al. 2015
+
†Ulrikezza aspoeckae Fang et al. 2015
Fm. Sisyridae Banks 1905 [spongillafly]
+
Subfm. †Cretadilarinae Makarkin 2017
+
G. †Cretadilar Makarkin 2017
+
†Cretadilar olei Makarkin 2017
G. †Microsisyra Makarkin 2022
+
†Microsisyra carsteni Makarkin 2022
G. †Paleosisyra Nel et al. 2003
+
†Paleosisyra eocenica Nel et al. 2003
†Paleosisyra minor Wichard et al. 2016
Subfm. †Paradoxosisyrinae Makarkin 2016
+
G. †Buratina Khramov 2019
+
†Buratina lusitanica Termier et al. 1985
†Buratina truncata Khramov 2019
G. †Khobotun Khramov 2019
+
†Khobotun elephantinus Khramov 2019
G. †Paradoxosisyra Makarkin 2016
+
†Paradoxosisyra groehni Makarkin 2016
G. †Protosiphoniella Khramov 2019
+
G. †Sidorchukatia Khramov 2019
+
†Sidorchukatia gracilis Khramov 2019
G. †Prosisyrina Perkovsky and Makarkin 2015
+
†Prosisyrina sphinga Makarkin and Perkovsky 2016
†Prosisyrina sukachevae Perkovsky and Makarkin 2015
G. Sisyra Burmeister 1839
+
†Sisyra amissa Hagen 1856
G. †Stictosisyra Yang et al. 2018
+
†Stictosisyra pennyi Yang et al. 2018
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Changhsingian to the top of the Messinian or 254.17000 to 5.33300 Ma
Minimum age of oldest fossil (stem group age): 252.17 Ma

Collections (133 total)


Time interval Ma Country or state Original ID and collection number
Changhsingian254.17 - 252.17South Africa (Natal) Archeosmylidae indet. (124428)
Changhsingian254.17 - 252.17South Africa (Kwazulu-Natal) Archeosmylus sp. (14361)
Changhsingian254.17 - 252.17Australia (New South Wales) Archeosmylus pectinatus (122433)
Griesbachian251.902 - 250.5Russian Federation (Kemerovo) Babykamenia eskovi (104713)
Anisian247.2 - 242.0Australia (Queensland) Lithosmylidia baronne (154203)
Ladinian242.0 - 237.0Kyrgyzstan (Osh) Madygoneura elongata (105007) Osmylotriasia superba (106088) Triasella ovata (110763)
Carnian237.0 - 228.0Japan (Yamaguchi) Momonokus obscurus (106482)
Norian228.0 - 208.5Australia (Queensland) Archeosmylus sp. (111558) Lithosmylidia parvula (154204) Lithosmylidia sp. A, Archeosmylus stigmatus, Archeosmylus costalis, Lithosmylidia lineata, Lithosmylidia parvula (106484)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi, Sogjuta speciosa (105087)
Sinemurian199.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi (113412)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Mesosmylina falciferum (114993)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Mesosmylina exornata (105002) Tetanoptilon brunsvicense (107323)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Erlikosmylus obscurus (118023) Mesosmylina shurabica (105003) Tengriosmylus magnificus, Mesosmylina angusta (110817)
Toarcian - Aalenian182.7 - 170.3Kyrgyzstan (Osh) Sauktangida aenigmatica (156499)
Bajocian170.3 - 168.3France Neuropora sp. (34712 34714)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Osmylidae indet., Thaumatomerobius mirabilis, Mesosmylina sibirica, Kubekius multiramosus, Epiosmylus longus (118010)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Saucrosmylidae indet. (114615)
Bathonian168.3 - 166.1Romania (Central Dobrogea) Neuropora spinosa (38190)
Bathonian168.3 - 166.1Mongolia Mesosmylina mongolica (119621)
Late/Upper Bathonian167.7 - 164.7France Neuropora spinosa, Neuropora sp. erect form (37510)
Late/Upper Bathonian167.7 - 164.7India (Kachchh) Neuropora sp. (24050)
Late/Upper Bathonian167.7 - 164.7India (Gujarat) Neuropora sp. (93561)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Arbusella sp. (106279) Ensiosmylus acutus (113439) Jurosmylus parvulus, Jurakempynus arcanus, Arbusella sp. (105970) Jurosmylus parvulus, Kolbasinella elongata, Arbusella bella (105426) Mesosmylus atalantus, Epiosmylus longicornis (106101) Pronymphites elegans, Parosmylus latus (105971) Scapoptera recta, Jurosmylus parvulus (106102)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Allotriosmylus uniramosus, Archaeosmylidia fusca (113365) Huiyingosmylus bellus, Daohugosmylus castus, Ponomarenkius sp., Ponomarenkius excellens, Arbusella magna, Palaeoneurorthus baii (113371) Jurakempynus sinensis, Jurakempynus bellatulus, Jurakempynus epunctatus, Bellinympha filicifolia, Bellinympha dancei, Juraheterosmylus antiquatus, Juraheterosmylus astictus, Juraheterosmylus minor (113366) Laccosmylus calophlebius, Laccosmylus cicatricatus, Laccosmylus latizonus (196979) Mirokempynus profundobifurcus, Jurakempynus loculosus, Arbusella platyptera (219840) Nilionympha shantouensis (122589) Palaeoneurorthus baii (229433) Saucrosmylus sambneurus, Rudiosmylus ningchengensis, Laccosmylus calophlebius, Enodinympha translucida, Nilionympha pulchella, Nilionympha imperfecta, Palaeothyridosmylus sp., Palaeothyridosmylus septemaculatus, Epiosmylus panfilovi, Tenuosmylus brevineurus (113367) Ulrikezza sp., Ulrikezza aspoeckae (172659) Vetosmylus tentus, Vetosmylus maculosus (210302)
Early/Lower Callovian164.7 - 161.2India Neuropora sp. (67608)
Late/Upper Callovian164.7 - 161.2India Neuropora sp. (68291)
Middle Callovian - Early/Lower Kimmeridgian164.7 - 150.8Iran (Tabas Area) Neuropora sp. (41417)
Late/Upper Oxfordian161.2 - 155.7Germany (Bavaria) Neuropora sp. (25411)
Late/Upper Oxfordian161.2 - 155.7Portugal Neuropora lusitanica (68246)
Late/Upper Oxfordian161.2 - 155.7Romania Neuropora sp., Neuropora spinulosa (35199)
Late/Upper Oxfordian161.2 - 155.7Poland Neuropora sp. (60853 60854 60855)
Late/Upper Oxfordian161.2 - 155.7France Neuropora spinosa (67975)
Middle Oxfordian161.2 - 155.7France Neuropora spinosa (68642 163585)
Middle Oxfordian161.2 - 155.7Germany (Niedersachsen) Neuropora sp., Neuropora lusitanica (31435)
Late/Upper Oxfordian - Kimmeridgian161.2 - 152.1Portugal Neuropora lusitanica (68245)
Late/Upper Oxfordian - Valanginian161.2 - 132.9China (Xizang) Neuropora sp. (61414)
Kimmeridgian157.3 - 152.1Portugal (Centro) Neuropora spinosa, Neuropora lusitanica (68220)
Kimmeridgian157.3 - 152.1Portugal Neuropora lusitanica (68784 68787 68819) Neuropora sp. (68212) Neuropora spinosa (68207 68242)
Kimmeridgian - Early/Lower Tithonian157.3 - 145.0Romania Neuropora lusitanica (201710 201711)
Late/Upper Kimmeridgian155.7 - 150.8Austria Neuropora lusitanica (36223)
Tithonian152.1 - 145.0Mongolia Jurakempynus sublimis (125978) Osmylidae indet., Sogjuta shartegica, Dimidiosmylus ramosus (104748)
Tithonian152.1 - 145.0Mongolia (Dund-Gov’ Aymag) Saucrosmylidae indet. (122321)
Tithonian152.1 - 145.0Russian Federation (Chita) Mesosmylina sp. (125407)
Late/Upper Tithonian - Berriasian150.8 - 139.8Austria Neuropora lusitanica (144461)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Orthoptera indet. (105966)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Mesosmylidus vulgaris, Osmylochrysa anomala, Osmylochrysa fragilis, Stenochrysa gradata (122909)
Late/Upper Hauterivian136.4 - 130.0United Kingdom (England) Stenosmylina medialis (106417)
Early/Lower Barremian130.0 - 125.45United Kingdom (England) Protosmylina bifasciata (118921)
Aptian125.0 - 113.0Russian Federation (Buryatia) Cretosmylus sibiricus (119498) Kempynosmylus zherikhini (106103) Osmylochrysa navasia, Zakamnosmylus elongatus, Dzhidosmylus solus (113350)
Late/Upper Aptian122.46 - 113.0Brazil (Ceara) Nuddsia repatriata (34901)
Late/Upper Aptian122.46 - 113.0Brazil (Ceará) Nuddsia longiantennata (122557)
Early/Lower Cenomanian99.6 - 93.5Myanmar Buratina truncata, Sidorchukatia gracilis, Protosiphoniella anthophila, Khobotun elephantinus (190961) Burmaleon magnificus, Nuddsia simplex (161726) Cretarophalis patrickmuelleri (123053 167760 178159) Nevrorthidae indet. (184692 184693) Nuddsia simplex (212006) Osmylidae indet. (111688) Paradoxosisyra groehni, Cretadilar olei, Microsisyra carsteni (172900) Sisyroneurorthus aspoeckorum, Nuddsia simplex, Myanmarosmylus wintertoni (196491) Stictosisyra pennyi (165561)
Late/Upper Cenomanian99.6 - 93.5Lebanon Lebanosmylus leae (229466)
Late/Upper Santonian85.8 - 83.5Russian Federation Prosisyrina sp. (35055) Prosisyrina sukachevae, Prosisyrina sphinga (35056)
Maastrichtian72.1 - 66.0Russian Federation (Primorskiy) Imanosmylus ussuriensis (119699)
Ypresian56.0 - 47.8France (Oise) Paleosisyra eocenica (123850)
Ypresian56.0 - 47.8Canada (British Columbia) Osmylidia donnae (31677) Osmylidia sp. (104793)
Ypresian56.0 - 47.8Australia (Queensland) Euporismites balli (111780 125666 139941)
Ypresian56.0 - 47.8USA (Washington) Osmylidia glastrai (103206)
Bridgerian50.3 - 46.2USA (Colorado) Osmylidia taliae (181561)
Priabonian38.0 - 33.9Ukraine Rophalis relicta (122997)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Electroneurorthus malickyi, Palaeoneurorthus groehni, Proberotha prisca, Paleosisyra electrobaltica, Palaeoneurorthus bifurcatus, Proberotha dichotoma, Osmylidia picta (110735) Palaeoneurorthus eocaenus, Balticoneurorthus elegans (120814) Palaeoneurorthus sp. (201682) Paleosisyra electrobaltica (140480) Paleosisyra minor (123928) Proberotha prisca (123766) Rophalis relicta (123439) Rophalis relicta, Electroneurorthus malickyi, Proberotha prisca, Paleosisyra electrobaltica, Palaeoneurorthus hoffeinsorum (124468) Rophalis relicta, Proberotha prisca (123420)
Priabonian38.0 - 33.9United Kingdom (England) Sisyridae sp.A (21976)
Priabonian38.0 - 33.9Poland Sisyra relicta, Osmylus pictus, Proberotha prisca, Sisyra amissa (120811) Sisyra relicta, Proberotha prisca (123438)
Priabonian38.0 - 33.9Russian Federation Palaeoneurorthus hoffeinsorum (123419)
Chadronian37.2 - 33.9USA (Colorado) Lithosmylus columbianus (151548) Osmylidia requieta (113749 124624) Osmylidia requietus (106379) Osmylus columbianus (106453) Osmylus requietus (106384) Pseudosmylidia relicta (106381)
Sarmatian12.7 - 11.608Romania Osmylus sp. (156756)
Messinian7.246 - 5.333France (Cantal) Sisyra sp. (122195)