PBDB Taxon

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Beryciformes

Actinopteri - Beryciformes

Taxonomy

Beryciformes was named by Patterson (1964). It is extant.

It was assigned to Osteichthyes by Sepkoski (2002); to Euacanthopterygii by Forey et al. (2003); to Acanthopterygii by Nelson (2006); to Berycimorphaceae by Betancur et al. (2013); to Percomorpha by Nolf and Dockery (1990), Long (2011) and Bannikov (2014); and to Berycida by Nelson et al. (2016).

Synonyms

Beryx dalmaticus was named by Steindachner (1863).

It was considered a nomen dubium by Patterson (1964).
Beryx lesinensis was named by Bassani (1880).

It was synonymized subjectively with Beryx dalmaticus by Patterson (1964).
Beryx subovatus was named by Bassani (1882). It is a 3D body fossil. Its type locality is Lesina (K.K. Geologischen Reichsanstalt GBA collection), which is in a Cenomanian lagoonal/restricted shallow subtidal lime mudstone in the Starigrad Limestone Formation of Croatia.

It was considered a nomen dubium by Patterson (1964).

Synonymy list

Year	Name and author
1863	Beryx dalmaticus Steindachner
1880	Beryx lesinensis Bassani
1882	Beryx subovatus Bassani p. 226 figs. Pl 8, fig 4
1964	Beryciformes Patterson p. 217
1990	Beryciformes Nolf and Dockery III p. 8
2002	Beryciformes Sepkoski
2003	Beryciformes Forey et al. p. 229
2006	Beryciformes Nelson
2011	Beryciformes Long p. 244
2013	Beryciformes Betancur et al.
2014	Beryciformes Bannikov p. 26
2016	Beryciformes Nelson et al. p. 308

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
superclass	Actinopterygii	()
class	Actinopteri	()
subclass	Neopterygii	(Regan 1923)
—	Teleosteomorpha

Rank	Name	Author
—	Teleostei	(Müller 1846)
—	Teleocephala	de Pinna 1996
—	Clupeocephala	Patterson and Rosen 1977
—	Euteleosteomorpha
—	Neoteleostei	Rosen 1973
—	Eurypterygia	(Rosen 1973)
—	Ctenosquamata
—	Acanthomorphata	(Rosen 1973)
—	Euacanthomorphacea
—	Berycimorphaceae
order	Beryciformes
order	Beryciformes

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Beryciformes Patterson 1964

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G. †Africentrum White and Moy-Thomas 1941

Fm. Berycidae Lowe 1843

G. †Argilloberyx Casier 1966

Argilloberyx prestwichae Casier 1966

G. Beryx Cuvier 1829

†Beryx altus Ciobanu 1977

†Beryx eogenus David 1946

†Beryx kcli David 1946

†Beryx longus Ciobanu 1977

†Beryx sanjoaquinensis David 1946

G. Centroberyx Gill 1862

†Centroberyx anguinicauda Schwarzhans and Bratischko 2011

†Centroberyx fragilis Schwarzhans 2003

†Centroberyx ingens Koken 1884

†Centroberyx integer Koken 1885

†Centroberyx worthyi Schwarzhans et al. 2017

G. †Nishiberyx Schwarzhans et al. 2017

†Nishiberyx nishimotoi Schwarzhans et al. 2017

Subor. Berycoidei Patterson 1964

G. †Austrocentris Schwarzhans 1980

†Austrocentris cavernosus Stinton 1957

G. †Lobopterus Gorjanović-Kramberger 1895

†Lobopterus pectinatus Gorjanović-Kramberger 1895

G. †Sagaberyx Schwarzhans et al. 2017

†Sagaberyx kishimaensis Schwarzhans et al. 2017

G. †Trachichthodes Gilchrist 1903

†Trachichthodes circularis Shepherd 1916

†Trachichthodes integer Koken 1885

†Trachichthodes pulcher Schwarzhans 1980

†Trachichthodes salebrosus Stinton 1957

†Trachichthodes subrotundus Koken 1884

Fm. Trachichthyidae Bleeker 1859

G. †Egregioberyx Schwarzhans 1980

†Egregioberyx erectus Schwarzhans 1980

†Egregioberyx sphaeroides Stinton 1957

G. †Eregioberyx Schwarzhans 1980

†Eregioberyx erectus Schwarzhans 1980

G. Gephyroberyx Boulenger 1902

Gephyroberyx darwini Johnson 1866

G. Hoplostethus Cuvier and Valenciennes 1829

Subg. †Hoplostethus (Hoplostethus) Cuvier 1839

†Hoplostethus (Hoplostethus) mediterraneus Cuvier 1839

†Hoplostethus bambergi Priem 1913

†Hoplostethus hexagonalis Leriche 1905

†Hoplostethus lacinatus Koken 1885

Hoplostethus lawleyi Koken 1891

Hoplostethus mediterraneus Cuvier 1829

†Hoplostethus pisanus Koken 1891

†Hoplostethus praemediterraneus Schubert 1905

†Hoplostethus sinaustralis Schwarzhans 1985

G. †Notoberyx Schwarzhans et al. 2017

†Notoberyx cionei Schwarzhans et al. 2017

G. Optivus Whitley 1947

†Optivus moko Schwarzhans et al. 2017

G. Paratrachichthys Waite 1899

G. †Trachichthyidarum Unknonw 1900

†Trachichthyidarum coffeesandensis Nolf and Dockery III 1990

Invalid names: Lissoberycinae Alvarado-Ortega and Than-Marchese 2013 [empty]

Invalid names: Hoplopterygidae [empty], Trachichthodidae Boulenger 1902 [empty]

G. †Berycomorus Arambourg 1967

G. †Berycopsis Agassiz 1850

Berycopsis elegans Dixon 1850

†Berycopsis germanus Agassiz 1837

Invalid names: Platycormus oblongus von der Marck 1863 [synonym]

†Berycopsis paucoradius Dietze 2009

†Berycopsis pulcher Bannikov and Bacchia 2005

Invalid names: Platycormus von der Marck 1858 [synonym]

G. †Brazosiella Frizzell and Dante 1965

†Brazosiella kokeni Frizzell and Dante 1965

†Brazosiella moseleyi Frizzell and Dante 1965

G. †Cryptoberyx Gaudant 1978

G. Ctenoberyx

Subor. †Dinopterygoidei Patterson 1964

Fm. †Dinopterygidae

G. †Dinopteryx Woodward 1901

†Dinopteryx spinosus Davis 1887

Fm. †Pharmacichthyidae Patterson 1964

G. †Pharmacichthys Woodward 1942

†Pharmacichthys judensis Gayet 1980

†Pharmacichthys numismalis Gayet 1980

†Pharmacichthys venenifer Woodward 1942

Fm. Diretmidae

G. Diretmus Johnson 1864

G. †Holocentrites Conrad 1941

Superfm. †Holocentroidea Richardson 1846

G. †Paracentrus Forey et al. 2003

†Paracentrus lebanonensis Forey et al. 2003

Subor. †Holocentroidei Johnson and Patterson 1993

G. †Adriacentrus Radovcic 1975

G. †Alloberyx Gaudant 1969

G. †Caproberyx Regan 1911

†Caproberyx pharsus Patterson 1967

†Caproberyx polydesmus Arambourg 1954

†Caproberyx superbus Dixon 1850

Invalid names: Berycopsis major Woodward 1902 [synonym]

G. †Ctenocephalichthys Gaudant 1969

G. †Erugocentrus Radovcic 1975

G. †Kansius Hussakof 1929

†Kansius sternbergi Hussakof 1929

G. †Parospinus Gayet 1982

G. †Stichoberyx Gaudant 1978

G. †Stichocentrus Patterson 1967

†Stichocentrus liratus Patterson 1967

G. †Trachichthyoides Woodward 1902

Trachichthyoides ornatus Woodward 1902

G. †Inocentrus

G. Kryptophanaron Silvester and Fowler 1927

Fm. Melamphaidae

G. Melamphaes Guenther 1864

Subg. Melamphaes (Plectromus) Gill 1883

Melamphaes (Plectromus) ovalis Schwarzhans 1980

†Melamphaes acanthifer Lin et al. 2017

†Melamphaes leeae Schwarzhans 2019

†Melamphaes ovalis Schwarzhans 1980

Melamphaes polylepis Ebeling 1962

G. Scopeloberyx Zugmayer 1911

†Scopeloberyx siooides Schwarzhans 1980

G. Scopelogadus Vaillant 1888

†Scopelogadus capistranensis Ebeling 1962

Fm. Monocentridae

G. Cleidopus De Vis 1882

G. Monocentris Schneider 1801

†Monocentris coxi Casier 1958

†Monocentris lemoinei Priem 1911

†Monocentris lemoinei gulatus Frost 1925

†Monocentris lerichei Schubert 1916

†Monocentris senni Casier 1958

G. †Omosoma Costa 1857

†Omosoma garretti Bardack 1976

†Omosoma intermedium Woodward 1901

†Omosoma monasterii von der Marck 1885

†Omosoma pulchellum Davis 1887

†Omosoma sahelalmae Costa 1857

†Omosoma simum Arambourg 1954

G. †Pattersonoberyx Gayet 1980

G. Plectrypops Gill 1863

G. †Plesioberyx Gayet 1980

G. Poromitra Goode and Bean 1883

G. †Pseudholocentrum Arambourg 1967

Fm. †Pseudomonocentrididae González-Rodríguez et al. 2013

G. †Handuichthys González-Rodríguez et al. 2013

†Handuichthys interopercularis González-Rodríguez et al. 2013

G. †Pseudomonocentris González-Rodríguez et al. 2013

†Pseudomonocentris microspinosus González-Rodríguez et al. 2013

Fm. †Quaesitoberycidae Bannikov and Sorbini 2005

G. †Microcapros Gayet 1980

G. †Quaesitoberyx Bannikov and Sorbini 2005

†Quaesitoberyx minutus Bannikov and Sorbini 2005

G. Sargocentron Fowler 1904

G. †Scianenuropsis

Subor. †Trachichthyoidei Parr 1933

G. †Acrogaster Agassiz 1839

†Acrogaster anceps Arambourg 1954

†Acrogaster brevicostatus von der Marck 1863

†Acrogaster daviesi Davis 1887

†Acrogaster dayi Woodward 1942

†Acrogaster heckeli Pictet 1850

†Acrogaster minutus von der Marck 1863

†Acrogaster parvus Agassiz 1839

G. †Antarctiberyx Grande and Chatterjee 1987

†Antarctiberyx seymouri Grande and Chatterjee 1987

G. †Gigantokranion Riickert-Ulkiimen 1988

G. †Gnathoberyx Patterson 1967

†Gnathoberyx stigmosus Patterson 1967

Fm. †Hoplopterygiidae Jordan 1923

G. †Hoplopteryx Agassiz 1839

†Hoplopteryx antiquus Agassiz 1839

†Hoplopteryx dakotaensis Grandstaff and Parris 2016

†Hoplopteryx gephyrognathus Patterson 1964

†Hoplopteryx gibbus von der Marck 1863

†Hoplopteryx insculptus Cope 1869

Hoplopteryx lewesiensis Mantell 1822

Invalid names: Beryx ornatus Agassiz 1835 [synonym], Beryx zippei Agassiz 1834 [synonym], Hoplopteryx brevis Bayer 1905 [synonym]

†Hoplopteryx lewisi Davis 1887

†Hoplopteryx macracanthus Patterson 1964

†Hoplopteryx oblongus Davis 1887

Hoplopteryx simus Woodward 1902

†Hoplopteryx spinulosus Woodward 1942

†Hoplopteryx stachei Gorjanović-Kramberger 1895

†Hoplopteryx syriacus Pictet and Humbert 1866

G. †Judeoberyx Gayet 1980

†Judeoberyx princeps Gayet 1980

G. †Libanoberyx Gayet 1980

G. †Lissoberyx Patterson 1967

†Lissoberyx dayi Patterson 1967

G. †Stichopteryx Gaudant 1969

G. †Tubantia Patterson 1964

†Tubantia cataphractus von der Marck 1863

Invalid names: Anoplogastridae [empty], Barbourisiidae [empty], Beryx dalmaticus Steindachner 1863 [nomen dubium], Beryx subovatus Bassani 1882 [nomen dubium], Cetomimidae [empty], Gibberichthyidae [empty], Hispidoberycidae [empty], Polymixioidei Patterson 1964 [empty], Rondeletiidae [empty], Stephanoberycidae [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

marinesubp

Locomotion:

actively mobileo

Life habit:

nektonico

Diet:

carnivoreo

Created:

2009-07-20 20:59:28

Modified:

2009-07-20 22:59:28

Source: o = order, subp = subphylum

References: Hendy et al. 2009, Carroll 1988

Age range

Maximum range based only on fossils: base of the Early/Lower Cenomanian to the top of the Ionian or 99.60000 to 0.12900 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma

Collections (193 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Albian - Cenomanian	113.0 - 93.9	Mexico (Hidalgo)	Handuichthys interopercularis, Pseudomonocentris microspinosus (135815)
Cenomanian	100.5 - 93.9	United Kingdom (England)	Hoplopteryx lewesiensis (221383)
Cenomanian - Coniacian	100.5 - 86.3	United Kingdom (England)	Hoplopteryx superbus (222435)
Early/Lower Cenomanian	99.6 - 93.5	Palestinian Territory	Pharmacichthys judensis, Judeoberyx princeps (11804)
Early/Lower Cenomanian - Middle Cenomanian	99.6 - 93.5	United Kingdom (England)	Berycopsis elegans (118407)
Late/Upper Cenomanian	99.6 - 93.5	Lebanon	Acrogaster dayi (221233) Berycopsis pulcher (221172) Lissoberyx dayi, Caproberyx pharsus (220886) Lissoberyx dayi, Stichocentrus liratus (220887) Pharmacichthys venenifer (221080) Pycnosterinx lewisii (216596) Quaesitoberyx minutus (220882 221061) Quaesitoberyx minutus, Berycopsis pulcher (69119)
Late/Upper Cenomanian	99.6 - 93.5	Italy (Padova)	Beryciformes indet., Hoplopteryx sp. (214718)
Late/Upper Cenomanian	99.6 - 93.5	Italy	Beryciformes indet., Hoplopteryx sp. (214600)
Late/Upper Cenomanian	99.6 - 93.5	Slovenia	Lobopterus pectinatus, Beryx stachei (221237)
Late/Upper Cenomanian	99.6 - 93.5	Croatia	Beryx subovatus (123565)
Middle Cenomanian	99.6 - 93.5	Lebanon	Paracentrus lebanonensis (124426) Pharmacichthys numismalis (221634)
Middle Cenomanian - Late/Upper Cenomanian	99.6 - 93.5	United Kingdom (England)	Berycopsis elegans, Hoplopteryx simus (221373) Hoplopteryx lewesiensis (59055 63042) Hoplopteryx simus (222437 222438) Trachichthyoides ornatus (222451)
Turonian	93.9 - 89.8	USA (South Dakota)	Hoplopteryx dakotaensis (226543)
Turonian	93.9 - 89.8	United Kingdom (England)	Beryx superbus (222434) Hoplopteryx simus (222439)
Early/Lower Turonian	93.5 - 89.3	United Kingdom (England)	Berycopsis elegans (221374) Hoplopteryx superbus (222436)
Middle Turonian	93.5 - 89.3	United Kingdom (England)	Hoplopteryx lewesiensis (221378)
Middle Turonian - Late/Upper Turonian	93.5 - 89.3	United Kingdom (England)	Berycopsis elegans (221375) Berycopsis elegans, Berycopsis major (221376)
Late/Upper Turonian - Coniacian	93.5 - 86.3	United Kingdom (England)	Hoplopteryx lewesiensis (221382) Hoplopteryx simus (222440)
Coniacian	89.8 - 86.3	United Kingdom (England)	Hoplopteryx lewesiensis (133319 221381)
Coniacian - Santonian	89.8 - 83.6	Italy (Gorizia)	Beryciformes indet. (103811 116133)