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Beryciformes
Taxonomy
Beryciformes was named by Patterson (1964). It is extant.
It was assigned to Osteichthyes by Sepkoski (2002); to Euacanthopterygii by Forey et al. (2003); to Acanthopterygii by Nelson (2006); to Percomorpha by Nolf and Dockery (1990), Long (2011), Bannikov (2014); to Berycida by Nelson et al. (2016); and to Berycimorphaceae by Betancur et al. (2013), van der Laan (2019).
It was assigned to Osteichthyes by Sepkoski (2002); to Euacanthopterygii by Forey et al. (2003); to Acanthopterygii by Nelson (2006); to Percomorpha by Nolf and Dockery (1990), Long (2011), Bannikov (2014); to Berycida by Nelson et al. (2016); and to Berycimorphaceae by Betancur et al. (2013), van der Laan (2019).
Subtaxa
Africentrum, Anoplogastridae, Berycidae, Berycoidei, Berycomorus, Beryx dalmaticus (syn. Beryx lesinensis), Beryx subovatus, Brazosiella, Cryptoberyx, Ctenoberyx, Dinopterygoidei, Diretmidae, Holocentrites, Holocentroidea, Holocentroidei, Inocentrus, Kryptophanaron, Melamphaidae, Monocentridae, Omosoma, Pattersonoberyx, Plectrypops, Plesioberyx, Polymixioidei, Poromitra, Pseudholocentrum, Pseudomonocentrididae, Sargocentron, Scianenuropsis, Stephanoberycoidei, Trachichthyoidei
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1863 | Beryx dalmaticus Steindachner |
| 1880 | Beryx lesinensis Bassani |
| 1882 | Beryx subovatus Bassani p. 226 figs. Pl 8, fig 4 |
| 1964 | Beryciformes Patterson p. 217 |
| 1990 | Beryciformes Nolf and Dockery III p. 8 |
| 2002 | Beryciformes Sepkoski |
| 2003 | Beryciformes Forey et al. p. 229 |
| 2006 | Beryciformes Nelson |
| 2011 | Beryciformes Long p. 244 |
| 2013 | Beryciformes Betancur et al. |
| 2014 | Beryciformes Bannikov p. 26 |
| 2016 | Beryciformes Nelson et al. p. 308 |
| 2019 | Beryciformes van der Laan p. 74 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Or. Beryciformes Patterson 1964
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G. †Africentrum White and Moy-Thomas 1941
Fm. Berycidae Lowe 1843
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G. Beryx Cuvier 1829
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†Beryx altus Ciobanu 1977
†Beryx eogenus David 1946
†Beryx kcli David 1946
†Beryx longus Ciobanu 1977
†Beryx sanjoaquinensis David 1946
G. Centroberyx Gill 1862
+
†Centroberyx anguinicauda Schwarzhans and Bratischko 2011
†Centroberyx fragilis Schwarzhans 2003
†Centroberyx ingens Koken 1884
†Centroberyx integer Koken 1885
†Centroberyx worthyi Schwarzhans et al. 2017
Subor. Berycoidei Patterson 1964
+
G. †Trachichthodes Gilchrist 1903
+
†Trachichthodes circularis Shepherd 1916
†Trachichthodes integer Koken 1885
†Trachichthodes pulcher Schwarzhans 1980
†Trachichthodes salebrosus Stinton 1957
†Trachichthodes subrotundus Koken 1884
Invalid names: Trachichthodidae Boulenger 1902 [empty]
G. †Berycomorus Arambourg 1967
G. †Brazosiella Frizzell and Dante 1965
+
†Brazosiella kokeni Frizzell and Dante 1965
†Brazosiella moseleyi Frizzell and Dante 1965
G. †Cryptoberyx Gaudant 1978
G. Ctenoberyx
Subor. †Dinopterygoidei Patterson 1964
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G. †Holocentrites Conrad 1941
Superfm. †Holocentroidea Richardson 1846
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Subor. †Holocentroidei Johnson and Patterson 1993
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G. †Adriacentrus Radovcic 1975
G. †Ctenocephalichthys Gaudant 1969
G. †Erugocentrus Radovcic 1975
G. †Parospinus Gayet 1982
G. †Stichoberyx Gaudant 1978
G. †Inocentrus
G. Kryptophanaron Silvester and Fowler 1927
Fm. Melamphaidae
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G. Melamphaes Guenther 1864
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†Melamphaes acanthifer Lin et al. 2017
†Melamphaes leeae Schwarzhans 2019
Melamphaes lugubris Gilbert 1890 [highsnout melamphid]
†Melamphaes ovalis Schwarzhans 1980
Melamphaes polylepis Ebeling 1962
G. Scopelogadus Vaillant 1888
+
Scopelogadus bispinosus Gilbert 1915 [twospine bigscale]
†Scopelogadus capistranensis Ebeling 1962
Fm. Monocentridae
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G. Cleidopus De Vis 1882
G. Monocentris Schneider 1801
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†Monocentris coxi Casier 1958
†Monocentris lemoinei Priem 1911
†Monocentris lemoinei gulatus Frost 1925
†Monocentris lerichei Schubert 1916
†Monocentris senni Casier 1958
G. †Omosoma Costa 1857
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†Omosoma garretti Bardack 1976
†Omosoma intermedium Woodward 1901
†Omosoma monasterii von der Marck 1885
†Omosoma pulchellum Davis 1887
†Omosoma sahelalmae Costa 1857
†Omosoma simum Arambourg 1954
G. †Pattersonoberyx Gayet 1980
G. Plectrypops Gill 1863
G. †Plesioberyx Gayet 1980
G. Poromitra Goode and Bean 1883
G. †Pseudholocentrum Arambourg 1967
Fm. †Pseudomonocentrididae González-Rodríguez et al. 2013
+
G. †Handuichthys González-Rodríguez et al. 2013
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†Handuichthys interopercularis González-Rodríguez et al. 2013
G. †Pseudomonocentris González-Rodríguez et al. 2013
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†Pseudomonocentris microspinosus González-Rodríguez et al. 2013
G. Sargocentron Fowler 1904
G. †Scianenuropsis
Subor. Stephanoberycoidei
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Invalid names: Barbourisiidae [empty], Cetomimidae [empty], Gibberichthyidae [empty], Hispidoberycidae [empty], Rondeletiidae [empty], Stephanoberycidae [empty]
Subor. †Trachichthyoidei Parr 1933
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G. †Acrogaster Agassiz 1839
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†Acrogaster anceps Arambourg 1954
†Acrogaster brevicostatus von der Marck 1863
†Acrogaster daviesi Davis 1887
†Acrogaster dayi Woodward 1942
†Acrogaster heckeli Pictet 1850
†Acrogaster minutus von der Marck 1863
†Acrogaster parvus Agassiz 1839
G. †Gigantokranion Riickert-Ulkiimen 1988
Fm. †Hoplopterygiidae Jordan 1923
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G. †Hoplopteryx Agassiz 1839
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†Hoplopteryx antiquus Agassiz 1839
†Hoplopteryx dakotaensis Grandstaff and Parris 2016
†Hoplopteryx gephyrognathus Patterson 1964
†Hoplopteryx gibbus von der Marck 1863
†Hoplopteryx insculptus Cope 1869
Hoplopteryx lewesiensis Mantell 1822
+
Invalid names: Beryx ornatus Agassiz 1835 [synonym], Beryx zippei Agassiz 1834 [synonym], Hoplopteryx brevis Bayer 1902 [synonym]
†Hoplopteryx lewisi Davis 1887
†Hoplopteryx macracanthus Patterson 1964
†Hoplopteryx oblongus Davis 1887
Hoplopteryx simus Woodward 1902
†Hoplopteryx spinulosus Woodward 1942
†Hoplopteryx stachei Gorjanović-Kramberger 1895
†Hoplopteryx syriacus Pictet and Humbert 1866
G. †Libanoberyx Gayet 1980
G. †Stichopteryx Gaudant 1969
Invalid names: Anoplogastridae [empty], Beryx dalmaticus Steindachner 1863 [nomen dubium], Beryx subovatus Bassani 1882 [nomen dubium], Polymixioidei Patterson 1964 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: o = order, subp = subphylum | |||||
| References: Carroll 1988, Kiessling 2004, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Early/Lower Cenomanian to the top of the Chibanian or 100.50000 to 0.12900 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Minimum age of oldest fossil (stem group age): 93.9 Ma
Collections (170 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Albian - Cenomanian | Mexico (Hidalgo) | Handuichthys interopercularis, Pseudomonocentris microspinosus (135815) | |
| Cenomanian | United Kingdom (England) | Hoplopteryx lewesiensis (221383) | |
| Early/Lower Cenomanian | Palestine | Judeoberyx princeps (11804) | |
| Late/Upper Cenomanian | Croatia | Beryx subovatus (123565) | |
| Late/Upper Cenomanian | Italy (Padova) | Beryciformes indet., Hoplopteryx sp. (214718) | |
| Late/Upper Cenomanian | Slovenia | Lobopterus pectinatus, Beryx stachei (221237) | |
| Late/Upper Cenomanian | Lebanon | Acrogaster dayi (221233) Pycnosterinx lewisii (216596) Quaesitoberyx minutus (69119 220882 221061) | |
| Late/Upper Cenomanian | Italy | Beryciformes indet., Hoplopteryx sp. (214600) | |
| Middle Cenomanian | Lebanon | Paracentrus lebanonensis (124426) | |
| Middle Cenomanian - Late/Upper Cenomanian | United Kingdom (England) | Hoplopteryx lewesiensis (59055 63042) Hoplopteryx simus (221373 222437 222438) Trachichthyoides ornatus (222451) | |
| Cenomanian - Turonian | United Kingdom (England) | Hoplopteryx lewesiensis, Hoplopteryx simus, Trachichthyoides ornatus, Hoplopteryx macracanthus, Hoplopteryx gephyrognathus (238510) | |
| Early/Lower Turonian | Czechia | Hoplopteryx brevis (242202) | |
| Middle Turonian | United Kingdom (England) | Hoplopteryx lewesiensis (221378) | |
| Turonian | USA (South Dakota) | Hoplopteryx dakotaensis (226543) | |
| Turonian | United Kingdom (England) | Hoplopteryx simus (222439) | |
| Late/Upper Turonian - Coniacian | United Kingdom (England) | Hoplopteryx lewesiensis (221382) Hoplopteryx simus (222440) | |
| Coniacian | United Kingdom (England) | Hoplopteryx lewesiensis (133319 221381) | |
| Coniacian - Santonian | Italy (Gorizia) | Beryciformes indet. (103811 116133) | |
| Late/Upper Santonian | Lebanon | Beryx syriacus, Pycnosterinx heckelii (221226) Gnathoberyx stigmosus (92241) Hoplopteryx spinulosus (221232) Microdon pulchellus, Hoplopteryx spinosus, Pycnosterinx heckelii, Hoplopteryx oblongus, Omosoma sahelalmae (170327) Omosoma sachelalmae (221191) Pycnosterinx daviesii (221157) Pycnosterinx heckelii (221187) | |
| Santonian | USA (Kansas) | Kansius sternbergi (221066 221077 221103) Omosoma garretti (220909 221063 221064) | |
| Late/Upper Santonian - Early/Lower Campanian | USA (Alabama) | Hoplopteryx sp. (81932 82049) | |
| Campanian | Italy | Beryciformes indet. (35332) | |
| Early/Lower Campanian | USA (Texas) | Hoplopteryx lewesiensis (221085) | |
| Late/Upper Campanian | USA (Alaska) | Beryciformes indet. (14495 231243 231244 240913) | |
| Late/Upper Campanian | Germany | Acrogaster brevicostatus (221714) Acrogaster parvus, Omosoma monasterii, Sphenocephalus cataphractus, Hoplopteryx gibbus, Acrogaster minutus (221717) Hoplopteryx antiquus, Acrogaster parvus, Sphenocephalus cataphractus, Hoplopteryx gibbus (221711) | |
| Campanian - Early/Lower Maastrichtian | USA (South Dakota) | Beryx sp. (167918) | |
| Early/Lower Maastrichtian | USA (New Jersey) | Beryx insculptus (275664) | |
| Late/Upper Maastrichtian | USA (Arkansas) | Beryciformes indet. (242204) | |
| Late/Upper Maastrichtian | Antarctica | Antarctiberyx seymouri (221133) | |
| Late/Upper Maastrichtian | USA (New Jersey) | Beryx insculptus (88034) | |
| Danian | Greenland | Centroberyx fragilis (22943) | |
| Selandian | Denmark | Centroberyx fragilis (159792 159795) Centroberyx fragilis, Centroberyx integer (159793 159794) | |
| Selandian | USA (Alabama) | Centroberyx sp. (87817) | |
| Selandian - Thanetian | Ukraine | Centroberyx sp., Centroberyx anguinicauda, Centroberyx fragilis, Centroberyx integer (159509) | |
| Tiffanian | USA (California) | Beryciformes indet. (169439) | |
| Waipawan - Mangaorapan | New Zealand (South Island) | Centroberyx sp. (219497) | |
| Ypresian | USA (Alabama) | Centroberyx sp. (237215 237217) | |
| Ypresian | USA (Texas) | Lactarius kokeni (237222) | |
| Ypresian | USA (Mississippi) | Centroberyx sp. (237219) | |
| Ypresian | Antarctica | Centroberyx sp. (100414) | |
| Ypresian | United Kingdom (England) | Argilloberyx prestwichae (13299) | |
| Eocene | USA (Texas) | Brazosiella kokeni, Brazosiella moseleyi (84145) | |
| Eocene | Australia (Victoria) | Monocentris lemoinei (205844) | |
| Eocene | France | Trachichthodes circularis (220174) | |
| Mangaorapan | New Zealand (South Island) | Centroberyx pulcher (219499) | |
| Mangaorapan - Heretaungan | New Zealand (South Island) | Centroberyx pulcher (219500 219501) | |
| Heretaungan | New Zealand (South Island) | Centroberyx sp., Centroberyx pulcher (219498) | |
| Lutetian | France | Beryx sp., Centroberyx eocenicus, Melamphaes acanthifer (188957) Centroberyx eocenicus, Centroberyx sp., Diretmus semicompletus, Melamphaes acanthifer (188958) Melamphaes acanthifer (188962) | |
| Lutetian | USA (Texas) | Brazosiella kokeni, Brazosiella moseleyi (84146) Lactarius kokeni (209370 209371 237116 237130) | |
| Lutetian | Belgium | Trachichthodes circularis (123069) | |
| Lutetian | Germany (Lower Saxony,) | Centroberyx sp., Centroberyx ingens, Epigonidarum lerichei (188954) | |
| Lutetian | United Kingdom (England) | Monocentris cristata (41992 41996 41998) | |
| Lutetian | USA (Mississippi) | Lactarius kokeni (237142) | |
| Lutetian | USA (California) | Beryx sp., Brazosiella sp. (200654) | |
| Lutetian | USA (Alabama) | Beryciformes indet. (209368) | |
| Lutetian | Barbados | Monocentris coxi, Monocentris senni (221164) | |
| Lutetian | USA (Virginia) | Lactarius kokeni (237135) | |
| Lutetian - Bartonian | Antarctica (Seymour Island) | Beryciformes indet. (53666) | |
| Middle Eocene | USA (Texas) | Brazosiella kokeni, Brazosiella moseleyi (84143 84144 84147) | |
| Middle Eocene | USA (Alabama) | Brazosiella kokeni, Brazosiella moseleyi (84149) | |
| Middle Eocene - Late/Upper Eocene | USA (California) | Beryx sanjoaquinensis, Beryx eogenus (191015) | |
| Porangan | New Zealand (South Island) | Centroberyx pulcher (219502) | |
| Bortonian | New Zealand (South Island) | Centroberyx pulcher (219504 219509) Trachichthodes pulcher (205332 219322) | |
| Bartonian | USA (Mississippi) | Lactarius kokeni (237141) | |
| Bartonian | USA (Alabama) | Lactarius kokeni (237117) | |
| Bartonian | United Kingdom (England) | Otolithus lemoinei (42636) | |
| Bartonian | USA (Texas) | Brazosiella moselevi, Brazosiella kokeni (7437 7438 7439) Lactarius kokeni (237131 237143) | |
| Kaiatan | New Zealand (South Island) | Centroberyx pulcher (219512) Trachichthodes pulcher (219333) | |
| Rupelian | Japan (Saga) | Nishiberyx nishimotoi, Sagaberyx kishimaensis (204551) | |
| Rupelian | Romania | Beryx altus, Beryx longus (113962) | |
| Rupelian | Germany | Plectrypops insignis, Centroberyx ingens (95898) | |
| Duntroonian | New Zealand (South Island) | Centroberyx worthyi (219615 219618) | |
| Chattian | Australia (Victoria) | Cleidopus carvernosus, Trachichthodes salebrosus (48647) Cleidopus cavernosus, Austrocentris cavernosus (174152) | |
| Chattian | France | Centroberyx sp. (219194) | |
| Late/Upper Oligocene | USA (Louisiana) | Centroberyx sp. (44521) | |
| Waitakian | New Zealand (South Island) | Centroberyx worthyi (219336 219579) | |
| Aquitanian | France | Poromitra sp., Sargocentron sp. (219195) | |
| Early/Lower Miocene | USA (California) | Beryx kcli (191012) | |
| Neogene | Indonesia | Monocentris lemoinei (223638) | |
| Neogene | Indonesia (Aceh) | Monocentris lemoinei (223637) | |
| Otaian | New Zealand | Melamphaes leeae (152323) | |
| Otaian | New Zealand (North Island) | Melamphaes leeae (219522) Scopelogadus sp., Centroberyx worthyi, Melamphaes leeae (219517) | |
| Otaian | New Zealand (South Island) | Centroberyx worthyi (219591) | |
| Altonian | New Zealand (North Shore) | Centroberyx worthyi (219524 219525 219526) | |
| Altonian | New Zealand (South Island) | Centroberyx worthyi (219600 219622) | |
| Langhian | France | Sargocentron ostialis (219197) | |
| Langhian | Australia (Victoria) | Trachichthodes salebrosus (46165 219137 219142) | |
| Middle Miocene | Australia (Victoria) | Trachichthodes salebrosus (51397 122349) | |
| Clifdenian | New Zealand (South Island) | Centroberyx worthyi (219623) | |
| Tortonian | Italy | Kryptophanaron sp., Scopelogadus sp. (178120) Melamphaes sp., Scopelogadus sp. (187503) Scopelogadus sp. (187504 187506 187596) | |
| Tortonian | Italy (Asti) | Centroberyx sp. (237147) | |
| Tortonian | Greece (Crete) | Scopelogadus sp. (207584 207586 207587 207588) | |
| Late/Upper Miocene | USA (California) | Scopelogadus capistranensis (85204) Scopelogadus mizolepis (91023) | |
| Tongaporutuan | New Zealand (North Island) | Melamphaes ovalis, Scopeloberyx siooides (219269) Scopeloberyx siooides, Melamphaes ovalis (219553) | |
| Messinian | Greece (Crete) | Scopelogadus sp. (207591) | |
| Zanclean | Italy | Sargocentron rubrum (210256) | |
| Piacenzian - Gelasian | Italy | Sargocentron rubrum (57058) | |
| Gelasian | Italy | Scopelogadus beanii (159096) | |
| Chibanian | USA (California) | Melamphaes lugubris, Scopelogadus bispinosus (276728) |