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Umenocoleidae
Taxonomy
Umenocoleidae was named by Chen and Tan (1973). Its type is Umenocoleus.
It was assigned to Coleoptera by Chen and Tan (1973); to Protelytroptera by Carpenter (1992); to Dictyoptera by Beutel et al. (2020); to Umenocoleoidea by Vršanský (2003), Bechly (2007), Nel et al. (2014), Lee (2016), Podstrelená and Sendi (2018), Luo et al. (2021); and to Alienoptera by Luo et al. (2022), Luo et al. (2022).
It was assigned to Coleoptera by Chen and Tan (1973); to Protelytroptera by Carpenter (1992); to Dictyoptera by Beutel et al. (2020); to Umenocoleoidea by Vršanský (2003), Bechly (2007), Nel et al. (2014), Lee (2016), Podstrelená and Sendi (2018), Luo et al. (2021); and to Alienoptera by Luo et al. (2022), Luo et al. (2022).
Subtaxa
Alienopterix, Alienopterixinae, Archaeospinapteryx, Blattapterix, Compunctiotypus, Cratovitisma, Elytropterix, Enervipraeala, Laticephalana, Maloval, Nigropterix, Petropterix, Poikiloprosopon, Ponopterix, Pseudoblattapterix, Pseudojantaropterix, Trapezionotum, Umenocoleus (type genus), Umenopterix, Umenotypus, Vitisminae, Vzrkadlenie
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1973 | Umenocoleidae Chen and Tan |
| 1992 | Umenocoleidae Carpenter p. 149 |
| 2003 | Umenocoleidae Vršanský p. 6 |
| 2007 | Umenocoleidae Bechly p. 242 |
| 2014 | Umenocoleidae Nel et al. |
| 2014 | Ponopterixidae Nel et al. p. 546 |
| 2016 | Umenocoleidae Lee p. 434 |
| 2018 | Umenocoleidae Podstrelená and Sendi p. 122 |
| 2020 | Umenocoleidae Beutel et al. |
| 2021 | Umenocoleidae Luo et al. p. 470 |
| 2021 | Alienopterixinae Vršanský p. 2210 |
| 2022 | Umenocoleidae Luo et al. |
| 2022 | Alienopterixinae Szabó et al. p. 1704 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Umenocoleidae Chen and Tan 1973
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G. †Alienopterix Mlynský et al. 2018
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†Alienopterix mlynskyi Sendi 2021
†Alienopterix ocularis Mlynský et al. 2018
†Alienopterix santonicus Szabó et al. 2022
†Alienopterix smidovae Hinkelman 2021
G. †Archaeospinapteryx Sendi and Cumming 2023
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†Archaeospinapteryx tartarensis Sendi and Cumming 2023
G. †Cratovitisma Bechly 2007
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†Cratovitisma bechlyi Podstrelená 2018
†Cratovitisma cortexi Sendi 2018
†Cratovitisma oldreadi Bechly 2007
†Cratovitisma spinosa Sendi et al. 2023
G. †Petropterix Vršanský 2003
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†Petropterix alexeevi Vršanský 2003
†Petropterix fukuiensis Oyama et al. 2022
†Petropterix koreaensis Lee et al. 2024
†Petropterix kukalovae Vršanský 2003
†Petropterix mirabilis Vršanský 2003
†Petropterix sibirix Vršanský 2003
G. †Ponopterix Vršanský and Grimaldi 1999
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†Ponopterix axelrodi Vršanský and Grimaldi 1999
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Invalid names: Ponopterix maxima Bechly 2007 [synonym]
G. †Umenocoleus Chen and Tan 1973
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†Umenocoleus minimus Lee et al. 2024
†Umenocoleus nervosus Zhang 1997
†Umenocoleus sinuatus Chen and Tan 1973
Subfm. †Vitisminae Vršanský and Ansorge 2001
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G. †Perspicuus Koubová 2020
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†Perspicuus csincsii Szabó et al. 2024
†Perspicuus pilosus Koubová 2020
†Perspicuus vrsanskyi Mlynský 2020
G. †Vitisma Vršanský 1999
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†Vitisma coriacea Sendi 2024
†Vitisma diffusa Vršanský 2003
†Vitisma occidentalis Vršanský and Ansorge 2001
†Vitisma okata Kováčová 2022
†Vitisma orientalis Vršanský 2005
†Vitisma rasnitsyni Vršanský 1999
G. †Vzrkadlenie Vršanský 2020
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†Vzrkadlenie karneri Sendi and Cumming 2023
†Vzrkadlenie miso Vršanský 2020
†Vzrkadlenie saintgermaini Sendi and Cumming 2023
Invalid names: Alienopterixinae Vršanský 2021 [invalid subgroup], Ponopterixidae Nel et al. 2014 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| P. Vršanský 2003 | Head more or less globular, antenna with wide segments and very long setae in transversal rows; pronotum mostly short and narrow, with reduced paranota; forewing heavily elytrised, with cup–like cells called "bunky" and reduced venation; hindwing venation generally of early polyphagoid groundplan (simplified venation with CuA without tertiary branches, CuP simple, A1 curved, with short branches), but with reduced venation of R (with few branches – possibly with pterostigma), M branched. Cerci with numerous long hair; females with short external ovipositor. |
Measurements
No measurements are available
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| Source: f = family | |||||
| Reference: Vršanský 2003 | |||||
Collections (51 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Callovian - Oxfordian | Kazakhstan (Chimkent) | Maloval hlavolam (114238) | |
| Barremian | Mongolia (Selenginskij) | Vitisma orientalis (117954) | |
| Early/Lower Barremian | Spain | Vitisma occidentalis (118913) | |
| Early/Lower Barremian | Lebanon | Blattulidae indet. (146535 153333) Cratovitisma cortexi (123416) Jantaropterix lebani (122359 123001) Pseudojantaropterix lebani, Cratovitisma cortexi (122471) | |
| Aptian | Russian Federation (Buryatia) | Petropterix sibirix (105536 118216) Petropterix sibirix, Vitisma rasnitsyni (122549) Vitisma rasnitsyni (106103 125412 125413 157030) | |
| Aptian | Japan | Vitisma sp., Petropterix fukuiensis (233240) | |
| Aptian | China (Jilin) | Umenocoleus nervosus (151876) | |
| Aptian | Mongolia | Elytropterix magnificiens (138924) Petropterix mirabilis (113355 121711 121713 125194 138922) Petropterix mirabilis, Petropterix alexeevi, Vitisma diffusa (113356) Petropterix mirabilis, Vitisma diffusa (113359) Vitisma diffusa (121714) Vitisma okata (103273) | |
| Late/Upper Aptian | Brazil (Ceará) | Cratovitisma oldreadi (122919 122921) Cratovitisma oldreadi, Ponopterix maxima, Ponopterix axelrodi (122557) Ponopterix axelrodi (122352 138921) Ponopterix burkhardi (122918) Ponopterix maxima (140371) | |
| Late/Upper Aptian | China (Gansu) | Blattapterix gansu (138919) | |
| Late/Upper Aptian | Brazil (Ceara) | Cratovitisma oldreadi, Ponopterix axelrodi (34899) | |
| Early/Lower Albian - Middle Albian | China (Gansu) | Petropterix kukalovae (138923) Ponopterix sp. 1, Ponopterix sp. 2, Umenocoleus sinuatus (224254) Umenocoleus sinuatus (138918) | |
| Albian | Jordan | Compunctiotypus hudai (124847) | |
| Albian | South Korea (Gyeongsangnam-do) | Umenocoleus minimus, Pseudoblattapterix weoni (238015) | |
| Albian | South Korea | Petropterix koreaensis (238014) | |
| Early/Lower Cenomanian | Myanmar | Alienopterix ocularis, Cratovitisma bechlyi, Enervipraeala nigra (161726) Cratovitisma bechlyi (123053) Cratovitisma bechlyi, Perspicuus pilosus, Perspicuus vrsanskyi, Vzrkadlenie miso, Alienopterix smidovae, Alienopterix mlynskyi, Archaeospinapteryx tartarensis, Trapezionotum vrsanskyi, Poikiloprosopon celiae, Vzrkadlenie karneri, Vzrkadlenie saintgermaini, Nigropterix cummingi, Vitisma coriacea (184414) Laticephalana liuyani (193537) | |
| Turonian | Kazakhstan | Nigropterix angustata, Umenotypus maculatus (103274) | |
| Turonian | Kazakhstan (Kzyl-Orda) | Nigropterix angustata, Umenotypus maculatus (113401) | |
| Santonian | Hungary | Alienopterix santonicus (230288) Perspicuus csincsii (224054) |