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Panendymenoptera

Insecta

Taxonomy
Panendymenoptera was named by Engel (2022). It is extant.

It was assigned to Euamphiesmenoptera by Engel (2022).

Subtaxa
Endymenoptera, Necrotrichoptera
Sister taxa
Endymenoptera

Synonymy list
YearName and author
2022Panendymenoptera Engel p. 4

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
RankNameAuthor
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
superorderAmphiesmenopteraKiriakoff 1948
—Metamphiesmenoptera
—Euamphiesmenoptera
—Panendymenoptera
—Panendymenoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Panendymenoptera Engel 2022
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Unr. Endymenoptera Engel 2022
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Unr. Stelloptera Engel 2022
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Or. Lepidoptera Linnaeus 1758 [moth]
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Superfm. Agathiphagoidea Kristensen 1967
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Fm. Agathiphagidae Kristensen 1967 [kauri moth]
Unr. Angiospermivora Regier et al. 2015
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Subor. Glossata Fabricius 1775
Fm. †Archaeolepiidae Whalley 1985
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G. †Archaeolepis Whalley 1985
Fm. †Ascololepidopterigidae Zhang et al. 2013
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G. †Ascololepidopterix Zhang et al. 2013
G. †Pegolepidopteron Zhang et al. 2013
G. †Trionolepidopteron Zhang et al. 2013
G. †Bombycites Heer 1849
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†Bombycites buchii Heer 1864
G. †Chionaemopsis Cockerell and Le Veque 1931
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†Chionaemopsis quadrifasciatus Cockerell and Le Veque 1931
G. Cyphanta Walker 1865
Unr. Diurni Latreille 1802
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Invalid names: Papilionides Latreille 1802 [empty]
Subor. †Eolepidopterigina Rasnitsyn 1983
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Superfm. †Eolepidopterigoidea Rasnitsyn 1983
G. Eucosmia Stephens 1829
G. †Gallerites Kernbach 1967
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†Gallerites keleri Kernbach 1967
Subor. Heterobathmiina Kristensen and Nielsen 1983
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Superfm. Heterobathmioidea Kristensen and Nielsen 1979
G. †Karataunia Kozlov 1989
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†Karataunia lapidaria Kozlov 1989
Superfm. Micropterigoidea Herrich-Schaeffer 1855
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Fm. Micropterigidae Cotes 1889
Unr. Nocturni Latreille 1802
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Invalid names: Bombycinae Latreille 1802 [empty], Phalaenites Latreille 1802 [empty], Pterophorii Latreille 1802 [empty], Tortrices Latreille 1802 [empty]
G. †Paleolepidopterites Kozlov 2018
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Unr. †Paralepidoptera Engel 2022
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Fm. †Mesokristenseniidae Huang et al. 2010
G. †Petisca Martins-Neto 1998
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†Petisca dryellina Martins-Neto 1998
G. †Phylledestes Cockerell 1907
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†Phylledestes vorax Cockerell 1907
G. †Pyralites Heer 1856
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†Pyralites obscurus Heer 1856
†Pyralites preecei Jarzembowski 1980
G. Schaefferia Houlbert 1918
G. †Sphingidites Kernbach 1967
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†Sphingidites weidneri Kernbach 1967
Invalid names: Aglossata Speidel 1977 [invalid subgroup], Monotrysia Borner 1939 [empty], Papilionida Laicharting 1781 [synonym], Phalaena Linnaeus 1758 [invalid subgroup], Zalepidoptera Engel 2022 [empty], Zeugloptera Chapman 1917 [empty]
Or. Trichoptera Kirby 1815 [caddisfly]
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G. †Amechanites Haupt 1956
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†Amechanites costatus Haupt 1956
G. †Archotaulius Handlirsch 1906
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†Archotaulius bavaricus Handlirsch 1906
G. †Axiomaldia Melnitsky and Ivanov 2020
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†Axiomaldia flinti Melnitsky and Ivanov 2020
G. †Calamodontus Botosaneanu and Wichard 1983
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†Calamodontus grandaevus Botosaneanu and Wichard 1983
Fm. †Cladochoristidae Riek 1953
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G. †Cladochorista Tillyard 1926
G. †Cladochoristella Riek 1955
G. †Meloclada Melnitsky and Ivanov 2020
G. †Conchindusia Vyalov and Sakacheva 1976
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†Conchindusia longaeva Sukacheva 2000
†Conchindusia rasnitsyni Jarzembowski 1995
G. †Coprindusia Ivanov 2006
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†Coprindusia pallida Ivanov 2006
Unr. Eutrichoptera Engel 2022
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Subor. Annulipalpia Martynov 1924
Subor. Integripalpia Martynov 1924
G. †Folindusia Berry 1927
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Subg. †Folindusia (Acrindusia) Vialov 1973
Subg. †Folindusia (Echinindusia) Vialov and Sukatsheva 1976
Subg. †Folindusia (Folindusia) Berry 1927
Subg. †Folindusia (Spirindusia) Vialov and Sukacheva 1976
†Folindusia adumbrata Sukacheva 2000
†Folindusia digna Sukacheva 2000
†Folindusia manlaica Sukatsheva 1980
†Folindusia megasporae Sukacheva 2000
†Folindusia miocenica Berry 1928
†Folindusia pilarius Sukacheva 2000
†Folindusia wilcoxiana Berry 1927
Fm. †Kalophryganidae Haupt 1956
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G. †Kalophryganea Haupt 1956
G. †Litholimnephilops Robinson et al. 2018
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†Litholimnephilops yinani Robinson et al. 2018
G. †Ostracindusia Vyalov 1973
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†Ostracindusia martynovae Sukacheva 2000
†Ostracindusia modesta Sukacheva 1982
†Ostracindusia vyalovi Jarzembowski 1995
G. †Paleodicella Sukatsheva 2014
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†Paleodicella anomala Sukatsheva 2014
G. †Pelindusia Vyalov and Sakacheva 1976
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†Pelindusia percealleni Jarzembowski 1995
G. †Petropteron Cockerell 1912
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†Petropteron mirandum Cockerell 1912
G. †Phryganeidium Westwood 1854
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†Phryganeidium pytho Westwood 1854
G. †Piscindusia Jarzembowski 1995
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†Piscindusia complexa Heads 2006
†Piscindusia sukachevae Jarzembowski 1995
Fm. †Prorhyacophilidae Riek 1955
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G. †Prorhyacophila Riek 1955
Subor. †Protomeropina Tillyard 1926
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G. †Adoptysma Novokshonov 1993
Fm. †Terminoptysmatidae Melnitsky and Ivanov 2020
G. †Sanctipaulus Pinto 1956
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†Sanctipaulus mendesi Pinto 1956
G. †Secrindusia Vialov and Sukacheva 1976
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†Secrindusia expressa Sukacheva 2000
G. †Terrindusia Vialov 1973
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Subg. †Terrindusia (Terrindusia) Vialov 1973
†Terrindusia antiqua Heer 1849
†Terrindusia buriatica Sukatsheva and Vasilenko 2019
†Terrindusia fluvialis Sukatsheva 1980
†Terrindusia khasurtica Sukatsheva and Vasilenko 2019
†Terrindusia laxa Sukatsheva 1980
†Terrindusia micacea Frič 1869
G. †Trichopterella Cockerell 1924
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†Trichopterella torta Cockerell 1924
G. †Tuanwangica Zhang 1985
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†Tuanwangica aethoneura Zhang 1985
G. †Zygobasis Hong 1982
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†Zygobasis mesozoica Hong 1982
Invalid names: Aphelocheira Stephens 1836 [invalid subgroup], Aspatherium geinitzii Kolenati 1851 [nomen nudum], Bittacus validus Hagen 1856 [nomen dubium], Cyrnus subatomarius Kolenati 1851 [nomen nudum], Hallesus retusus Hagen 1856 [nomen dubium], Hydropsyche submaculata Kolenati 1851 [nomen nudum], Hydropsyche subvariabilis Kolenati 1851 [nomen nudum], Kalasiroptera Engel 2022 [empty], Nebritus willistoni Melander 1949 [nomen dubium], Phryganea arenacea Fritsch 1899 [nomen dubium], Phryganea kolbi Frič 1889 [nomen dubium], Phryganea solitaria Fritsch 1899 [nomen dubium], Phryganeina Sukatsheva 1968 [empty], Phryganoidea Handlirsch 1906 [synonym], Rhyacophila subumbrosa Kolenati 1851 [nomen nudum], Rhyacophila succinica Kolenati 1851 [nomen nudum], Rhyacophila succinica major Kolenati 1851 [nomen nudum], Rhyacophila succinica media Kolenati 1851 [nomen nudum], Rhyacophila succinica minor Kolenati 1851 [nomen nudum], Spicipalpia [invalid subgroup]
Or. †Tarachoptera Mey et al. 2017
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Fm. †Tarachocelidae Mey et al. 2017
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G. †Kinitocelis Mey et al. 2017
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†Kinitocelis brevicostata Mey et al. 2017
†Kinitocelis dashengi Wang et al. 2022
†Kinitocelis divisonotata Mey et al. 2017
†Kinitocelis hennigi Mey et al. 2017
†Kinitocelis macroptera Mey and Wichard 2020
†Kinitocelis patrickmuelleri Mey and Wichard 2023
†Kinitocelis sparsella Mey et al. 2018
G. †Retortocelis Mey et al. 2018
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†Retortocelis longella Mey et al. 2018
†Retortocelis minimella Mey et al. 2018
†Retortocelis spicipalpia Mey and Wichard 2020
†Retortocelis tyloptera Mey et al. 2018
G. †Tarachocelis Mey et al. 2017
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†Tarachocelis emmarossae Mey and Wichard 2023
Or. †Necrotrichoptera Engel 2022
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Fm. †Necrotauliidae Handlirsch 1906
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G. †Acisarcuatus Liu et al. 2014
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†Acisarcuatus locellatus Zhang et al. 2017
†Acisarcuatus variradius Liu et al. 2014
G. †Anecrotaulius Engel 2022
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G. †Archiptilia Handlirsch 1939
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†Archiptilia ovata Handlirsch 1939
G. †Austaulius Kelly et al. 2018
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†Austaulius furcatus Giebel 1856
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Invalid names: Necrotaulius apicalis Tillyard 1933 [synonym], Necrotaulius stigmaticus Tillyard 1933 [synonym], Necrotaulius westwoodi Tillyard 1933 [synonym], Orthophlebia liasina Giebel 1856 [synonym]
†Austaulius haustrum Kelly et al. 2018
G. †Cretotaulius Sukacheva 1982
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†Cretotaulius ultimus Sukacheva 1982
G. †Epididontus Handlirsch 1939
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†Epididontus geinitzianus Handlirsch 1939
G. †Karataulius Sukacheva 1968
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†Karataulius aeternus Sukacheva 1968
†Karataulius martae Sukatsheva and Sinitshenkova 2023
G. †Mesotrichopteridium Handlirsch 1906
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Invalid names: Mesotrichopteridium pusillum Handlirsch 1906 [synonym], Necrotaulius ellipticus Handlirsch 1939 [synonym], Necrotaulius maculatus Handlirsch 1920 [synonym], Necrotaulius megapolitanus Handlirsch 1939 [synonym], Necrotaulius pullus Handlirsch 1939 [synonym], Necrotaulius similis Handlirsch 1906 [synonym]
G. †Metarchitaulius Handlirsch 1939
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†Metarchitaulius longus Handlirsch 1939
G. †Nannotrichopteron Handlirsch 1906
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†Nannotrichopteron gracile Handlirsch 1906
G. †Necrotaulius Handlirsch 1906
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†Necrotaulius communis Sukatsheva and Sinitshenkova 2023
†Necrotaulius kritus Lin 1986
†Necrotaulius kubekovi Sukatsheva 1985
†Necrotaulius mantellorum Jarzembowski 1991
†Necrotaulius minutissimus Sukacheva 1990
†Necrotaulius minutus Sukacheva 1968
†Necrotaulius parvulus Geinitz 1884
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Invalid names: Necrotaulius bodei Handlirsch 1939 [synonym], Necrotaulius dobbertinensis Handlirsch 1906 [synonym], Necrotaulius minimus Handlirsch 1939 [synonym], Necrotaulius nanus Handlirsch 1906 [synonym], Necrotaulius obtusior Bode 1953 [synonym], Necrotaulius pygmaeus Tillyard 1933 [synonym], Necrotaulius pygmaeus Handlirsch 1939 [synonym], Necrotaulius ulmeri Fischer 1962 [synonym], Orthophlebia marginata Bode 1905 [synonym]
†Necrotaulius tener Sukacheva 1990
Invalid names: Karatauliodes Sukacheva 1968 [synonym]
G. †Palaeotaulius Handlirsch 1939
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†Palaeotaulius vicinus Handlirsch 1939
G. †Paranecrotaulius Engel 2022
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†Paranecrotaulius proximus Sukatsheva 1973
G. †Pararchitaulius Handlirsch 1939
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†Pararchitaulius ovalis Handlirsch 1939
G. †Parataulius Handlirsch 1939
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†Parataulius jurassicus Handlirsch 1939
G. †Paratrichopteridium Handlirsch 1906
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†Paratrichopteridium costale Handlirsch 1939
†Paratrichopteridium efossum Handlirsch 1939
Invalid names: Architaulius Handlirsch 1939 [synonym]
G. †Pseudorthophlebia Handlirsch 1906
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†Pseudorthophlebia brodiei Handlirsch 1906
†Pseudorthophlebia platyptera Handlirsch 1906
G. †Pteromixanum Sukatsheva and Jarzembowski 2001
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†Pteromixanum inviolatum Sukatsheva and Jarzembowski 2001
†Pteromixanum poxwellense Sukatsheva and Jarzembowski 2001
†Pteromixanum purbeckianum Handlirsch 1906
†Pteromixanum ruderatum Sukatsheva and Jarzembowski 2001
G. †Scyphindusia Sukatsheva 1985
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†Scyphindusia hydroptiliformis Sukatsheva 1985
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Westphalian C to the top of the Holocene or 314.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 309.8 Ma

Collections (495 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Moscovian315.2 - 307.0Czech Republic Phryganea kolbi, Phryganea arenacea (125662)
Westphalian C314.6 - 309.8Czech Republic Phryganea solitaria (155300)
Kungurian279.3 - 272.3Russian Federation (Komi) Adoptysma anomalanala (97621)
Wuchiapingian259.9 - 254.17Russian Federation (Vologda) Cladochorista issadica (104754)
Changhsingian254.17 - 252.17South Africa (Natal) Cladochoristidae indet. (124428)
Changhsingian254.17 - 252.17Russian Federation (Vladimir) Cladochoristella ryzhkovae, Prorhyacophila rasnitsyni (113351)
Changhsingian254.17 - 252.17Australia (New South Wales) Cladochorista belmontensis (122423 122433)
Ladinian242.0 - 237.0Brazil (Rio Grande do Sul) Sanctipaulus mendesi (116119)
Ladinian242.0 - 237.0Kyrgyzstan (Osh) Cladochorista curta, Cladochoristella sola, Prorhyacophila rara, Prorhyacophila batkenica (163987) Cladochorista multivenosa, Prophilopotamus asiaticus (106088) Necrotaulius proximus, Prorhyacophila furcata (110765)
Longobardian242.0 - 235.0China (Shaanxi) Folindusia sp. (183209)
Carnian237.0 - 228.0China (Xinjiang) Terrindusia sp. (196151)
Norian228.0 - 208.5Australia (Queensland) Cladochoristella bryani, Prorhyacophila colliveri (106484)
Rhaetian208.5 - 201.3United Kingdom (Somerset) Austaulius furcatus (193353)
Rhaetian208.5 - 201.3Germany Lepidoptera indet., Glossata indet. (191173)
Rhaetian208.5 - 201.3United Kingdom (England) Hemerobius sp. 1, Hemerobius sp. 2 (115246) Necrotaulius sp. (113957 114644) Necrotaulius stigmaticus, Necrotaulius apicalis, Necrotaulius sp. (115222)
Planorbis201.3 - 196.5United Kingdom (England) Necrotaulius furcatus (115368)
Planorbis201.3 - 196.5Germany (Niedersachsen) Lepidoptera indet., Glossata indet. (191174)
Turneri196.5 - 189.6United Kingdom (England) Archaeolepis mane (114645)
Turneri196.5 - 189.6United Kingdom (Dorset) Austaulius haustrum (189409)
Early/Lower Toarcian183.0 - 182.0Germany Epididontus geinitzianus, Paratrichopteridium efossum, Paratrichopteridium costale, Palaeotaulius vicinus, Necrotaulius intermedius, Necrotaulius megapolitanus, Necrotaulius ellipticus, Necrotaulius pullus, Necrotaulius maculatus, Mesotrichopteridium pusillum, Necrotaulius similis, Archiptilia ovata, Metarchitaulius longus, Pararchitaulius ovalis, Parataulius jurassicus, Nannotrichopteron gracile, Orthophlebia parvula, Necrotaulius nanus, Necrotaulius dobbertinensis, Necrotaulius pygmaeus, Necrotaulius minimus, Paratrichopteridium breve, Paratrichopteridium angustum, Liadotaulius acutipennis, Paratrichopteridium areatum, Orthophlebia furcata, Orthophlebia furcata (123987) Mesotrichopteridium intermedium, Necrotaulius parvulus, Liadotaulius maior (114991)