PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Elongatoolithus

Osteichthyes - Elongatoolithidae

Taxonomy

Elongatoolithus was named by Zhao (1975). It is considered to be a form taxon.

It was assigned to Oviraptoridae by Zelenitsky and Currie (2004); and to Elongatoolithidae by Zhao (1975), Zeng and Zhang (1979), Zhao (1979), Mikhailov (1991), Zhao et al. (1991), Mikhailov (1994), Mikhailov et al. (1994), Zhao (1994), Mikhailov (1997), Mikhailov (2000), Zelenitsky et al. (2000), Amo Sanjuán et al. (2000), Cousin (2002), Fang et al. (2003), Watabe (2004), Mohabey (2005), Liang et al. (2009), Wang et al. (2010), Zhao et al. (2013), Wang et al. (2013), Huh et al. (2014), Zhang et al. (2017), Simon et al. (2019), Jin et al. (2020), Tanaka et al. (2023), Wang and Wang (2024).

Species

E. andrewsi, E. chichengshanensis, E. chimeiensis, E. elongatus, E. excellens, E. frustrabilis, E. jianchangensis, E. laijiaensis, E. magnus, E. sigillarius, E. subtitectorius, E. taipinghuensis, E. yangjiagouensis

Synonymy list

Year	Name and author
1975	Elongatoolithus Zhao p. 108
1979	Elongatoolithus Zeng and Zhang p. 133
1979	Elongatoolithus Zhao p. 336
1991	Elongatoolithus Mikhailov p. 221
1991	Elongatoolithus Zhao et al. p. 5
1994	Elongatoolithus Mikhailov p. 111
1994	Elongatoolithus Mikhailov et al. p. 103
1994	Elongatoolithus Zhao p. 187
1997	Elongatoolithus Mikhailov p. 62
2000	Elongatoolithus Amo Sanjuán et al. p. 9
2000	Elongatoolithus Mikhailov p. 569
2000	Elongatoolithus Zelenitsky et al. p. 130
2002	Elongatoolithus Cousin p. 43
2003	Elongatoolithus Fang et al. p. 517
2004	Elongatoolithus Watabe p. 17
2004	Elongatoolithus Zelenitsky and Currie p. 134A
2005	Elongatoolithus Mohabey p. 473
2009	Elongatoolithus Liang et al. p. 1590
2010	Elongatoolithus Wang et al. p. 114
2013	Elongatoolithus Wang et al. p. 199
2013	Elongatoolithus Zhao et al. p. 4665
2014	Elongatoolithus Huh et al. p. 220
2017	Elongatoolithus Zhang et al. p. 191
2019	Elongatoolithus Simon et al. p. 7
2020	Elongatoolithus Jin et al. p. 9
2023	Elongatoolithus Tanaka et al. p. 202
2024	Elongatoolithus Wang and Wang p. 40

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
class	Osteichthyes
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida

Rank	Name	Author
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Saurischia	()
—	Theropoda	()
family	Elongatoolithidae
genus	Elongatoolithus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Elongatoolithus Zhao 1975

show all | hide all

†Elongatoolithus andrewsi Zhao 1975

†Elongatoolithus chichengshanensis Fang et al. 2003

†Elongatoolithus chimeiensis Fang et al. 2007

†Elongatoolithus elongatus Young 1954

†Elongatoolithus excellens Mikhailov 1994

†Elongatoolithus frustrabilis Mikhailov 1994

†Elongatoolithus jianchangensis Fang et al. 2007

†Elongatoolithus laijiaensis Fang et al. 2003

†Elongatoolithus magnus Zeng and Zhang 1979

†Elongatoolithus sigillarius Mikhailov 1994

†Elongatoolithus subtitectorius Mikhailov 1994

†Elongatoolithus taipinghuensis Yu 1998

†Elongatoolithus yangjiagouensis Fang et al. 2007

Diagnosis

Reference	Diagnosis
K. E. Mikhailov 1994	Elongated-ellipsoidal eggs of medium size (up to 170 mm long) with "fine" ornamentation on surface (a = 8–11) and typically elongatoolithid microstructure of shell. Ratio of spongiose to mammilar layer ranging from 2:1 to 3:1.
K. E. Mikhailov 1997	Eggs of medium size (long diameter < 170 mm) with a fine linearituberculate ornamentation (8-1 1 lines of ridges per 10 mm range of egg short diameter). Elongation of egg = 2.0-2.2. Thickness ratio between continuous and mammillary layers varies from 2 : 1 to 3 : 1.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

solitaryo

Environment:

terrestrialo

Locomotion:

actively mobileo

Life habit:

ground dwellingo

Diet:

carnivoreo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2004-12-14 12:50:08

Modified:

2004-12-14 14:50:08

Source: o = order

References: Kiessling 2004, Marsh 1875

Age range: base of the Early/Lower Barremian to the top of the Maastrichtian or 125.77000 to 66.00000 Ma

Collections (75 total)

Time interval	Ma	Country or state	Original ID and collection number
Early/Lower Cretaceous	143.1 - 100.5	China (Henan)	E. yangjiagouensis (275532)
Early/Lower Barremian	125.77 - 121.4	Spain (Aragón)	Elongatoolithidae indet. (61409)
Early/Lower Albian	113.2 - 110.1	Japan (Hyogo)	E. sp. (117980)
Late/Upper Cenomanian - Turonian	100.5 - 89.8	China (Zhejiang)	E. chichengshanensis (204721) E. laijiaensis (204718)
Cenomanian - Santonian	100.5 - 83.6	China (Guangdong)	E. sp. (188428)
Late/Upper Cretaceous	100.5 - 66.0	China (Xinjiang)	Oolithes elongatus (97613)
Late/Upper Cretaceous	100.5 - 66.0	China (Hunan)	E. sp., E. magnus (71664)
Late/Upper Coniacian	89.8 - 85.7	China (Henan)	E. sp. (275545 275546 275547)
Coniacian - Santonian	89.8 - 83.6	China (Henan)	E. chimeiensis (275535) E. jianchangensis (275513 275517 275524 275525 275526)
Late/Upper Coniacian - Late/Upper Santonian	89.8 - 83.6	China (Henan)	E. sp. (275542)
Campanian	83.6 - 72.2	Mongolia (Omnogov)	E. frustrabilis (37211)
Campanian	83.6 - 72.2	China (Guangdong)	E. andrewsi (275468 275473 275475 275477 275479 275480 275482 275483) E. andrewsi, E. elongates (275474 275481 275491) E. elongatus (275469 275478 275484 275485 275488 275492 275497) E. elongatus, E. andrewsi (275489 275496)
Campanian	83.6 - 72.2	China (Shandong)	Oolithes elongatus (80114 80115 80117) Oolithes elongatus, Reptilia indet. (24757)
Campanian	83.6 - 72.2	Mongolia (Ömnögovi Aimag)	E. subtitectorius (130798)
Campanian	83.6 - 72.2	China (Nei Mongol)	Elongatoolithidae indet. (46904) E. sp. (205652)
Late/Upper Campanian	83.6 - 72.2	Mongolia (Omnogov)	Elongatoolithidae indet. (53229 65344) E. sp. (118259 127668) Protoceratops andrewsi (46152)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	Mongolia (Dornogov)	Elongatoolithidae indet. (45331)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	Mongolia (Omnogov)	E. sp. (174156)
Campanian - Maastrichtian	83.6 - 66.0	China (Xinjiang)	Oolithes elongatus (159589) Oolithes sp. (24753)
Campanian - Maastrichtian	83.6 - 66.0	China (Guangdong)	E. elongatus (275422)
Campanian - Maastrichtian	83.6 - 66.0	China (Hunan)	Oolithes elongatus (79741)
Maastrichtian	72.2 - 66.0	Mongolia (Omnogov)	Elongatoolithidae indet. (55257 65335 165580)
Maastrichtian	72.2 - 66.0	China (Guangdong)	Dinosauria indet. (71855) E. andrewsi (275461) E. elongatus, E. andrewsi (24752 68149 68396 68397 68398 68399 275464 275465 275466) Oolithes elongatus (71854 71859 71867)
Maastrichtian	72.2 - 66.0	China (Jiangxi)	E. elongatus (220036)