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Ellipsocephaloidea

Artiopoda - Redlichiida

Synonymy list
YearName and author
1990Ellipsocephaloidea Lin
2008Ellipsocephaloidea Landing et al. p. 893
2011Ellipsocephaloidea Adrain p. 104
2022Ellipsocephaloidea Cederström et al. p. 31
2024Ellipsocephaloidea El Albani et al. p. S7

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
RankNameAuthor
classArtiopodaHou and Bergstrom
classTrilobitaWalch 1771
orderRedlichiidaRichter 1932
suborderRedlichiinaRichter 1932
superfamilyEllipsocephaloideaLin 1990
superfamilyEllipsocephaloideaLin 1990

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. †Ellipsocephaloidea Lin 1990
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Fm. †Agraulidae Raymond 1913
+
G. †Agaso Cobbold 1934
G. †Agraulos Hawle and Corda 1847
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†Agraulos ceticephalus Barrande 1846
+
Invalid names: Agraulos longicephalus Hicks 1872 [synonym]
†Agraulos globosus Walcott 1884
†Agraulos quadrangularis Whitfield 1884
†Agraulos selcupicus Rozova 1964
†Agraulos strenuus Billings 1874
†Agraulos transversus Yang et al. 1993
†Agraulos woosteri Whitfield 1878
Invalid names: Agrauloides Howell 1937 [synonym], Arion Barrande 1846 [replaced]
G. †Batenioides Repina 1960
G. †Chittidilla King 1941
+
†Chittidilla nanjiangensis Zhang et al. 1980
†Chittidilla oblonga Zhang et al. 1980
†Chittidilla similis Zhang et al. 1980
†Chittidilla yunnanensis Zhang et al. 1980
Invalid names: Diandongaspidella Yuan et al. 1997 [synonym]
G. †Chondroparia Lorenz 1906
G. †Litavkaspis Fatka et al. 1987
+
†Litavkaspis rejkovicensis Falka et al. 1987
G. †Metagraulos Kobayashi 1935
+
†Metagraulos dolon Walcott 1905
†Metagraulos sampoensis Kobayashi 1961
†Metagraulos truncatus Zhang and Yuan 1981
G. †Parachittidilla Zhang et al. 1980
+
†Parachittidilla obscura Zhang et al. 1980
†Parachittidilla xiaolinghouensis Zhang et al. 1980
Invalid names: Amurticephalus Shah et al. 1991 [synonym]
G. †Paragraulos Lu 1941
+
†Paragraulos convexus Yuan and Li 1999
G. †Plesiagraulos Chang 1963
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†Plesiagraulos transversus Yuan and Li 1999
G. †Poriagraulos Chang 1963
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†Poriagraulos abrota Walcott 1905
†Poriagraulos dactylogrammacus Zhang and Yuan 1981
+
Invalid names: Poriagraulos nanum Dames 1883 [synonym]
G. †Proampyx Frech 1897
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†Proampyx aculeatus Angelin 1851
†Proampyx acuminatus Angelin 1851
†Proampyx difformis Angelin 1851
G. †Protochittidilla Qiu 1980
G. †Pseudoeteraspis Chernysheva 1950
G. †Pseudoplesiagraulos Lu et al. 1988
+
†Pseudoplesiagraulos planus Lu et al. 1988
G. †Qiannanagraulos Yuan et al. 1997
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†Qiannanagraulos costatus Yuan et al. 1997
†Qiannanagraulos orientalis Yuan et al. 1997
G. †Skreiaspis Ruzicka 1946
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†Skreiaspis spinosus Jahn 1895
G. †Tangshihlingia Chu 1959
+
G. †Tholus Pegel 1986
Fm. †Bigotinidae Hupé 1953
+
G. †Bigotina Cobbold 1935
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†Bigotina angulata Surorova 1960
†Bigotina bivallata Cobbold 1935
†Bigotina inornata Egorova 1986
†Bigotina kelleri Geyer 2019
†Bigotina monningeri Geyer 2019
G. †Bigotinella Suvorova 1960
+
†Bigotinella malykanica Suvorova 1960
G. †Bigotinops Hupe 1952
+
†Bigotinops chouberti Geyer 2019
†Bigotinops dangeardi Hupe 1953
G. †Demuma Özdikmen 2009
+
†Demuma nicklesi Hupe 1952
Invalid names: Pruvostina Hupe 1952 [replaced]
G. †Eladiolinania Geyer 2019
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†Eladiolinania castor Geyer 2019
†Eladiolinania pollux Geyer 2019
G. †Hupetina Sdzuy 1978
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†Hupetina antiqua Sdzuy 1978
G. †Issendalenia Geyer 2019
+
G. †Ouijjania Hupe 1953
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G. †Pseudobigotina Geyer 2019
+
Subfm. †Serrinae Liñán et al. 2008
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G. †Sdzuyia Liñán and Gozalo 1999
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†Sdzuyia sanmamesi Liñán and Gozalo 1999
G. †Serrania Linan 1978
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†Serrania verae Linan 1978
G. †Suvorovaella Geyer 2019
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†Suvorovaella patria Suvorova 1960
†Suvorovaella priva Suvorova 1960
G. †Tioutella Geyer 2019
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†Tioutella floccofavosa Geyer 2019
Fm. †Ellipsocephalidae Matthew 1887
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G. †Acadolenus Sdzuy 1967
G. †Alueva Lotze 1958
Subfm. †Antatlasiinae Hupe 1953
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G. †Antatlasia Hupe 1952
G. †Berabichia Geyer 1990
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†Berabichia subditus Palmer and Gatehouse 1972
G. †Argunaspis Repina 1966
G. †Asiatella Repina 1964
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†Asiatella elegans Repina 1964
G. †Bergeroniaspis Lermontova 1951
G. †Bergeroniellus Lermontova 1940
G. †Bhargavia Peng et al. 2009
+
†Bhargavia prakritika Peng et al. 2009
G. †Blayacina Cobbold 1932
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Invalid names: Sardaspis Brasier 1976 [synonym]
G. †Brevitermierella Hupé 1953
+
Invalid names: Paratermierella Hupe 1952 [synonym]
G. †Cambrunicornia Geyer 1990
G. †Catadoxides Matthew 1899
G. †Culmenaspis Repina 1972
Subfm. †Ellipsocephalinae Mathew 1887
+
G. †Cambrosaurura Cederström et al. 2022
G. †Dellingia Cederström et al. 2022
G. †Ellipsocephalus Zenker 1833
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†Ellipsocephalus gripi Kautsky 1945
Invalid names: Germaropyge Snajdr 1957 [synonym]
G. †Ellipsostrenua Kautsky 1945
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†Ellipsostrenua sularpensis Ahlberg and Bergström 1978
†Ellipsostrenua troedssoni Cederström et al. 2022
G. †Kingaspidoides Hupe 1953
+
†Kingaspidoides alberti Geyer 2017
†Kingaspidoides meieri Geyer 2017
†Kingaspidoides usitata Geyer 1990
Invalid names: Elatius Orlowski 1985 [synonym]
G. †Ornamentaspis Geyer 1990
+
†Ornamentaspis hupei Orłowski 1964
G. †Geyerorodes Özdikmen 2009
+
†Geyerorodes howleyi Walcott 1890
†Geyerorodes schmitti Geyer 1990
Invalid names: Orodes Geyer 1990 [replaced]
G. †Glabrella Lermontova 1940
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†Glabrella pitans Palmer and Gatehouse 1972
†Glabrella ventrosa Lermontova 1940
G. †Hamatolenus Hupe 1952
+
Subg. †Hamatolenus (Hamatolenus) Hupé 1952
+
†Hamatolenus (Hamatolenus) vincenti Geyer and Landing 2004
Subg. †Hamatolenus (Myopsolenus) Hupe 1952
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Invalid names: Collyrolenus Hupe 1952 [synonym]
G. †Issafeniella Geyer 1990
G. †Kadyella Pokrovskaya 1959
G. †Kameschkoviella Repina 1959
G. †Kijanella Repina 1958
G. †Kingaspis Kobayashi 1935
+
Subg. †Kingaspis (Kingaspidoides) Hupe 1953
+
†Kingaspis (Kingaspidoides) armatus Liñán et al. 2003
+
Invalid names: Kingaspis (Kingaspis) brevifrons Hupé 1953 [synonym]
Subg. †Kingaspis (Kingaspis) Hupé 1954
+
†Kingaspis avalonensis Geyer and Landing 2001
†Kingaspis convexus Palmer and Rowell 1995
Invalid names: Mesetaia Hupe 1953 [synonym]
G. †Krolina Repina 1960
G. †Kymataspis Geyer 1990
G. †Latikingaspis Geyer 1990
+
†Latikingaspis alatus Hupé 1954
G. †Latoucheia Hupe 1953
G. †Latuzella Repina et al. 1999
G. †Lermontovia Suvorova 1956
G. †Limataceps Repina 1969
G. †Mohicana Cobbold 1910
G. †Nelegeria Korobov 1966
+
†Nelegeria lata Korobov 1966
G. †Olekmaspis Suvorova 1956
G. †Ourikaia Geyer 1990
G. †Paramicmacca Lermontova 1951
G. †Paraprotolenella Zhu and Jiang 1981
+
†Paraprotolenella subcylindrica Zhu and Jiang 1981
G. †Planolimbus Geyer 1990
G. †Protagraulos Matthew 1895
G. †Protolenella Qian and Yao 1974
+
†Protolenella conica Qian and Yao 1974
†Protolenella lata Zhou 1974
Subfm. †Protoleninae Richter and Richter
+
G. †Anaxarete Geyer 2016
+
†Anaxarete senior Cobbold 1910
G. †Chorbusulina Lazarenko 1962
+
†Chorbusulina bella Lazarenko 1964
†Chorbusulina wilkesi Palmer and Gatehouse 1972
G. †Cuneoaxiella Cederström et al. 2022
+
†Cuneoaxiella grandis Ahlberg and Bergström 1978
G. †Protolenus Matthew 1892
+
†Protolenus elegans Matthew 1892
+
Invalid names: Protolenus paradoxoides Matthew 1892 [synonym]
†Protolenus pustulatus Cobbold 1910
Invalid names: Bergeronia Matthew 1895 [objective synonym]
G. †Pruvostinoides Hupe 1952
G. †Pseudokadyella Repina 1966
G. †Pseudolenus Hupe 1953
G. †Pseudoprotolenella Zhu and Lin 1983
+
G. †Ptychoparopsis Hupe 1953
G. †Rinconia Hupe 1953
G. †Sailycaspis Repina 1965
G. †Sectigena Geyer 1990
Subfm. †Strenuaevinae Cederström et al. 2022
+
G. †Epichalnipsus Geyer et al. 2004
G. †Strenuaeva Richter and Richter 1940
+
†Strenuaeva inflata Ahlberg and Bergström 1978
†Strenuaeva primaeva Brøgger 1878
†Strenuaeva spinosa Ahlberg and Bergström 1978
†Strenueva annio Cobbold 1910
Invalid names: Hindermeyeria Hupe 1953 [synonym]
G. †Strenuella Matthew 1887
+
Subg. †Strenuella (Comluella) Hupe 1953
†Strenuella calceola Cobbold 1931
†Strenuella cobboldi Richter 1919
†Strenuella platycephala Cobbold 1910
†Strenuella polonica Czarnocki 1926
†Strenuella strenua Billings 1874
G. †Tadakoustia Geyer 1990
G. †Termieraspis Repina 1960
G. †Termierella Hupe 1952
+
G. †Lusatiops Richter 1941
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†Lusatiops harveyi Walcott 1890
Invalid names: Jalonella Hupé 1953 [synonym]
†Termierella totanesensis Sepúlveda et al. 2024
G. †Thoralaspis Hupé 1953
G. †Triangulaspis Lermontova 1940
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Invalid names: Acutaspis Repina 1976 [synonym], Angustaeva Hupé 1953 [synonym], Triangulina Repina 1965 [synonym]
G. †Yeshanaspis Zhu and Jiang 1981
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†Yeshanaspis subconicus Zhu and Jiang 1981
Fm. †Estaingiidae Opik 1970
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G. †Alanisia Hupe 1953
G. †Chulanolenus Zhang 1983
G. †Eomalungia Lee 1978
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†Eomalungia conica Li 1980
G. †Estaingia Pocock 1964
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†Estaingia bilobata Pocock 1964
†Estaingia cerastes Öpik 1975
†Estaingia fletcheri Öpik 1975
Invalid names: Hsuaspis Chang 1957 [synonym]
G. †Hupeia Kobayashi 1944
G. †Ichangia Chang 1965
+
†Ichangia ichangensis Chang 1965
G. †Mundocephalina Repina 1958
G. †Paraichangia Lee 1978
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†Paraichangia jiangsuensis Zhu and Jiang 1981
†Paraichangia spinosa Zhu and Jiang 1981
G. †Pararaia Kobayashi 1942
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†Pararaia bunyerooensis Bengtson et al. 1990
Invalid names: Proichangia Zhang et al. 1980 [synonym], Tannuolaspis [synonym]
G. †Shifangia Chien and Yao 1974
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†Shifangia yefanggouensis Yang et al. 1993
G. †Sichuanolenus Zhang et al. 1980
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†Sichuanolenus nanjiangensis Zhang et al. 1980
G. †Szechuanolenus Zhang and Zhu 1978
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†Szechuanolenus nanjiangensis Zhang and Zhu 1978
Fm. †Palaeolenidae Hupé 1953
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G. †Alataurus Repina 1964
G. †Bajangoliaspis Sivov 1960
G. †Enammocephalus Repina 1964
G. †Enixus Özdikmen 2009
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†Enixus enigmaticus Chernysheva 1956
†Enixus juvenis Chernysheva 1956
+
Invalid names: Wurmaspis rarus Sdzuy 1960 [synonym]
Invalid names: Schistocephalus Chernysheva 1956 [replaced]
G. †Ferralsia Cobbold 1935
G. †Gigoutella Hupe 1953
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†Gigoutella atlasensis Hupé 1953
†Gigoutella mauretanica Ortega-Hernández et al. 2017
G. †Habrocephalus Repina 1964
G. †Palaeolenella Repina 1960
G. †Palaeolenides Egorova 1967
G. †Palaeolenus Mansuy 1912
+
†Palaeolenus douvillei Mansuy 1912
†Palaeolenus lantenoisi Mansuy 1912
†Palaeolenus longispinus Zhang et al. 1980
†Palaeolenus tingi Lu 1941
G. †Resimopsis Repina 1958
G. †Sibirolenus Geyer 2016
+
†Sibirolenus enormis Repina 1965
G. †Validaspis Repina 1977
Fm. †Yunnanocephalidae Hupé 1953
+
G. †Elicicola Jel and Runnegar 1990
G. †Luaspis Hupe 1953
G. †Pinarella Özdikmen 2009
+
Invalid names: Pensacola Palmer and Gatehouse 1972 [replaced]
G. †Wangzishia Sun 1977
G. †Yunnanocephalus Kobayashi 1936
+
Invalid names: Pseudoptychoparia Ting 1940 [objective synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingo
Life habit: low-level epifaunalo
Diet: deposit feedero
Created: 2009-09-21 20:18:56
Modified: 2012-08-27 07:43:00
Source: o = order, c = class
References: Hendy et al. 2009, Aberhan et al. 2004

Age range: base of the Branchian to the top of the Trempealeauan or 516.20000 to 486.85000 Ma

Collections (363 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Caerfai538.8 - 506.5Canada (New Brunswick) Ellipsocephalus galeatus (34572 34597) Ellipsocephalus galeatus, Protolenus elegans (34571) Kingaspidoides obliquoculatus (34573) Protolenus articephalus, Protolenus elegans (34567 34751) Protolenus elegans (34569 34570 34595 34596 34752)
Early/Lower Cambrian538.8 - 506.5Spain (Aragon) Alueva undulata, Hamatolenus ibericus, Kingaspis campbelli, Sdzuyia sanmamesi, Protolenus pisidianus, Protolenus jilocanus (120550 120552)
Early/Lower Cambrian538.8 - 506.5Antarctica Glabrella pitans, Chorbusulina wilkesi, Pensacola isolata, Chorbusulina subdita (154030)
Early/Lower Cambrian538.8 - 506.5China (Yunnan) Yunnanocephalus yunnanensis (185938 185939)
Early/Lower Cambrian538.8 - 506.5Norway (Troms) Ellipsocephalus sp. (190927) Strenuaeva inflata (190925 190926)
Cambrian538.8 - 486.85Canada (British Columbia) Agraulos sp. (186)
Cambrian538.8 - 486.85USA (Wisconsin) Ellipsocephalus sp. (250)
Atdabanian521.0 - 516.0Spain Bigotinella sp. (153693 153695) Bigotinella sp., Bigotina bivallata (153696 153698) Lusatiops sp., Protolenus schneideri, Strenueva vigilans, Strenueva annio (225087) Serrania verae (55276 153167)
Atdabanian521.0 - 516.0United Kingdom (Wales) Hamatolenus douglasi (41750) Myopsolenus sp. (41747)
Atdabanian521.0 - 516.0Morocco (Ant-Atlas) Antatlasia hollardi (232863)

Youngest occurrences
Time interval Ma Country or state PBDB collection number
Marjumian504.5 - 497.0USA (South Carolina) Agraulos sp., Skreiaspis sp. (75062)
Wangcunian504.5 - 497.0China (Shanxi) Agraulos vicina (232857)
Undillan503.0 - 500.5Germany Enixus juvenis, Ornamentaspis crassilimbata, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186155) Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186152) Kingaspidoides frankenwaldensis (186149 186150 186153 186154 186159 186165 186166 186170) Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186167) Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186151 186157 186158) Kingaspidoides sp. (186162 197474) Latikingaspis alatus, Kingaspidoides meieri, Kingaspidoides frankenwaldensis (186169) Ornamentaspis crassilimbata, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186156 186163) Ornamentaspis crassilimbata, Kingaspidoides frankenwaldensis (186148 186160) Ornamentaspis crassilimbata, Kingaspidoides meieri, Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186168) Ornamentaspis sp., Kingaspidoides alberti, Kingaspidoides frankenwaldensis (186164) Skreiaspis sp. (155786)
Boomerangian500.5 - 499.0Australia (Queensland) Proampyx agra (225231)
Boomerangian500.5 - 499.0Australia (Queenslandd) Proampyx agra (225252)
Guzhangian500.5 - 497.0Sweden (Jämtland) Proampyx acuminatus (239967) Proampyx difformis, Proampyx acuminatus, Proampyx aculeatus (240068)
Guzhangian500.5 - 497.0China (Liaoning) Tangshihlingia subtriangulata (126903)
Guzhangian500.5 - 497.0Spain Proampyx difformis (182506)
Guzhangian500.5 - 497.0USA (Wisconsin) Cedaria woosteri (157538)
Trempealeauan490.2 - 486.85USA (New York) Agraulos saratogensis (74540 74541)