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Warthia
Taxonomy
Warthia was named by Waagen (1880) [Sepkoski's age data: C Step-l P Dora Sepkoski's reference number: 1130,1258]. Its type is Warthia brevisinuata.
It was reranked as Bellerophon (Warthia) by Popov (1957).
It was assigned to Bellerophontidae by Ulrich and Scofield (1897); to Bellerophon by Popov (1957); to Bellerophontida by Sepkoski (2002); to Euphemitidae by Horný (1963), Mazaev (2015), Nützel and Ketwetsuriya (2016) and Foster et al. (2017); and to Euphemitinae by Knight et al. (1960) and Ketwetsuriya et al. (2020).
It was reranked as Bellerophon (Warthia) by Popov (1957).
It was assigned to Bellerophontidae by Ulrich and Scofield (1897); to Bellerophon by Popov (1957); to Bellerophontida by Sepkoski (2002); to Euphemitidae by Horný (1963), Mazaev (2015), Nützel and Ketwetsuriya (2016) and Foster et al. (2017); and to Euphemitinae by Knight et al. (1960) and Ketwetsuriya et al. (2020).
Species
Stachella bicarinata, Stachella micra, Stachella pseudohelix (syn. Stachella comelicana), W. americana, W. angustior, W. attalicus, W. borealis, W. brevisinuata (type species), W. brevizonata, W. carinata, W. crassus, W. fissus (syn. Bellerophon pitkorskyi), W. hisakatsui, W. huayingshanensis, W. imtatschanensis, W. inflata, W. intermedia, W. kingi, W. lata, W. micromphala (syn. W. undulata), W. peregrinus, W. perspecta, W. polita, W. saundersi, W. semiaurita, W. stricta, W. vaceki, W. waageni, W. welleri, W. zakharovi
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1880 | Warthia Waagen p. 158 |
| 1880 | Stachella Waagen p. 171 |
| 1897 | Stachella Ulrich and Scofield |
| 1897 | Warthia Ulrich and Scofield |
| 1957 | Bellerophon (Warthia) Popov |
| 1960 | Warthia Knight et al. p. 178 |
| 1960 | Stachella Knight et al. p. 179 |
| 1963 | Stachella Horný |
| 1963 | Warthia Horný |
| 2002 | Stachella Sepkoski |
| 2002 | Warthia Sepkoski |
| 2015 | Warthia Mazaev p. 884 |
| 2016 | Warthia Nützel and Ketwetsuriya p. 485 |
| 2017 | Warthia Foster et al. |
| 2020 | Warthia Ketwetsuriya et al. p. 55 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Warthia Waagen 1880
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†Warthia americana Girty 1909
†Warthia angustior Yochelson 1960
†Warthia attalicus Enderle 1901
†Warthia borealis Spath 1930
†Warthia brevisinuata Waagen 1880
†Warthia brevizonata Wang 1980
†Warthia carinata Dickins 1963
†Warthia crassus Yochelson 1960
†Warthia hisakatsui Murata 1981
†Warthia huayingshanensis Yu and Xi 1974
†Warthia imtatschanensis Popov 1957
†Warthia inflata Yin 1932
†Warthia intermedia Dickins 1963
†Warthia kingi Moore 1941
†Warthia lata Waagen 1880
†Warthia peregrinus Laube 1868
†Warthia perspecta Fletcher 1958
†Warthia polita Waagen 1880
†Warthia saundersi Yochelson 1960
†Warthia semiaurita Waagen 1880
†Warthia stricta Dana 1847
†Warthia vaceki Bittner 1899
†Warthia waageni Yochelson 1960
†Warthia welleri Yochelson 1960
†Warthia zakharovi Kaim 2009
Invalid names: Stachella Waagen 1880 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| W. J. Foster et al. 2017 | Deeply involute planispirally coiled bellerophont gastropod, distinguished by absence of spiral ornament. Sinus is broad, usually with a short slit. Selenizone obscured by inductural layer. Ornament of faint growth lines and growth wrinkles. Umbilicus generally filled. |
Measurements
| part | N | mean | minimum | maximum | median | s.d. | |
|---|---|---|---|---|---|---|---|
| shell height | 89 | 10.7 | 2.63 | 44.0 | 11.2 | 6.46 |
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| Source: f = family, o = order, c = class | |||||
| Reference: Kiessling 2004 | |||||
Age range: base of the Tournaisian to the top of the Smithian or 358.90000 to 247.20000 Ma
Collections (190 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Tournaisian | USA (Iowa) | W. sp. (3309) | |
| Desmoinesian | USA (New Mexico) | W. kingi (46536) | |
| Virgilian | USA (Kansas) | W. kingi (111225) | |
| Gzhelian - Asselian | China (Hopei) | Stachella inflata (149177) | |
| Sakmarian | Australia (New South Wales) | Bellerophon undulatus (119250) W. sp. (133744 152058) | |
| Sakmarian | India (Kashmir) | W. micromphala (86542) | |
| Sakmarian | Australia (Western Australia) | W. carinata (83188) W. carinata, W. intermedia (85021) | |
| Artinskian | Australia (Western Australia) | W. intermedia (85854) | |
| Artinskian | USA (Texas) | W. crassus (90007) | |
| Artinskian | USA (New Mexico) | W. crassus (101771 101795 121509) | |
| Artinskian | Australia (Queensland) | W. micromphala (39937) W. sp. (103444) | |
| Kungurian | USA (Arizona) | W. crassus (121506) W. sp. (120746) | |
| Kungurian | China (Sichuan) | Stachella gigantea (116222) | |
| Kungurian | USA (Wyoming) | W. fissus (61953 62878) | |
| Kungurian | USA (Texas) | W. crassus (121370) W. waageni (75883 81630 81711) W. waageni, W. angustior (75884) W. welleri (43491 43515 43517 121485) | |
| Kungurian | New Zealand (Southland) | W. perspecta (41776) | |
| Kungurian | Mexico (Sonora) | W. sp. (52099 52100) | |
| Roadian | Australia (Western Australia) | Goniatites micromphalus (131549) | |
| Roadian | Russian Federation (Kirov) | W. fissus, W. waageni (126335) | |
| Roadian | Mexico (Chiapas) | W. waageni, W. welleri (220046) | |
| Roadian | Australia (New South Wales) | Agathiceras micromphalus (161842) Goniatites micromphalus (108756) | |
| Roadian | Thailand | W. sp. 1, W. sp. 2, W. welleri (103565) | |
| Roadian | Russian Federation (Tatarstan) | W. fissus (174705 174706 174709) | |
| Roadian | USA (Texas) | W. americana (108822) W. fissus, W. saundersi (81693 101767) W. fissus, W. welleri (43512) W. welleri (81640) | |
| Roadian | USA (Wyoming) | W. fissus (61938) | |
| Roadian | USA (Arizona) | W. sp. (40001) | |
| Roadian - Capitanian | USA (Texas) | W. americana (108835) | |
| Roadian - Capitanian | Russian Federation | W. imtatschanensis (110300) | |
| Wordian | Thailand | Bellerophon sp. 1 (167687 167690 167693) W. sp., W. brevisinuata (181783) | |
| Wordian | Pakistan | Stachella sp. (125851) W. brevisinuata (119691 125849) | |
| Wordian | Australia (New South Wales) | Bellerophon micromphalus (133342 133346 134048) Bellerophon micromphalus, Bellerophon strictus (133337) | |
| Wordian | USA (Montana) | W. sp. (11276) | |
| Wordian | USA (Wyoming) | W. sp. (11225) | |
| Wordian | Thailand (Ratchaburi) | W. saundersi (40954) | |
| Wordian | Mexico (Sonora) | Stachella sp. (40812) W. sp. A (40842) | |
| Wordian | New Zealand (Southland) | W. micromphala (41767 41784) W. micromphala, W. perspecta (120416) | |
| Wordian | New Zealand (Nelson) | W. sp. (41797) | |
| Wordian - Capitanian | Australia (Queensland) | W. stricta (104742) | |
| Capitanian | Japan | W. sp. (109667) | |
| Capitanian | Pakistan | Stachella sp. (127033) W. lata (119692) W. polita (126277) | |
| Capitanian | New Zealand (Southland) | W. micromphala (41788) W. micromphala, W. perspecta (41777) W. micromphala, W. stricta, W. perspecta (41778) | |
| Wuchiapingian | USA (Utah) | W. sp. (11234) | |
| Wuchiapingian | China (Chongqing) | W. huayingshanensis (98065) | |
| Wuchiapingian | USA (Wyoming) | W. fissus (11227) W. fissus, W. waageni (61951) W. waageni (61936) | |
| Wuchiapingian | Australia (Queensland) | Goniatites micromphalus (123258) Goniatites sp. (123259) W. micromphala (103924 120335 120336 120341 120539 120667) W. perspecta (85054) W. sp. (97270) | |
| Wuchiapingian | Australia (Tasmania) | W. micromphala (51314 105434) | |
| Wuchiapingian | China (Xizang) | Stachella micra (116129) | |
| Wuchiapingian | Japan | W. sp. (113878 113880 113881 113884 113885) | |
| Wuchiapingian | Pakistan | Stachella semiaurita (126738 126941) W. polita (126003 126939) W. sp. (125926) | |
| Wuchiapingian - Changhsingian | Pakistan | W. polita (126759 126762 127017) W. polita, W. brevisinuata (126741) | |
| Changhsingian | New Zealand | W. perspecta (98149) | |
| Changhsingian | China (Guangxi) | W. sp. (50024) | |
| Changhsingian | Italy (South Tyrol) | Bellerophon peregrinus, Bellerophon comelicanus (204983) | |
| Changhsingian | China (Guizhou) | Stachella bicarinata (111993) W. brevizonata (112094) | |
| Changhsingian | Italy | Bellerophon sp. (97947 101914 101915 101917 101921) Bellerophon vaceki (44450 74938 74939 87192 101922 101923 101924 101925 101926 101927 101936 101937) | |
| Changhsingian | Pakistan | Stachella semiaurita (90251) W. polita (89061 126887) | |
| Induan | Norway | Bellerophon borealis (31367) | |
| Griesbachian - Dienerian | Svalbard and Jan Mayen | Bellerophon borealis (211950) | |
| Griesbachian | Russian Federation | Bellerophon borealis (214748) W. zakharovi (90201 90205 221137) | |
| Griesbachian | India (Kashmir) | W. hisakatsui (48914) | |
| Griesbachian | China (Guizhou) | Stachella mica (112112) W. vaceki (200984 200988 201006 201013) | |
| Griesbachian | Greenland | Bellerophon borealis (48113 48115 112747 112756 118489 118510 118530 118600 118609 118631 118664 118677 118711 118729) Bellerophon vaceki (48114) | |
| Griesbachian | Svalbard and Jan Mayen | Bellerophon borealis (212072 212073) W. zakharovi (182611 182612) | |
| Griesbachian | Italy | Bellerophon vaceki (44451 74940 74941 74944 104956 104957) W. vaceki (127941) | |
| Smithian | Australia (Queensland) | Stachella sp. (44044 116736) | |
| Smithian | Russian Federation | W. zakharovi (90220 221139) | |
| Smithian | Pakistan | Stachella sp. (143630) W. hisakatsui (139083 139086 139141 139143 139150 140678 140683 140684) | |
| Dienerian | Russian Federation | Bellerophon borealis (214749) W. zakharovi (90218) |