PPP# 361 - Escudo de Veraguas [Escudo de Veraguas Fm] (Pliocene of Panama)

Also known as NMB Loc. 17834

Where: Bocas, Panama (9.1° N, 81.6° W: paleocoordinates 9.0° N, 81.1° W)

• coordinate stated in text

• outcrop-level geographic resolution

When: Escudo de Veraguas Formation (Bocas del Toro Group), Piacenzian (3.6 - 2.6 Ma)

• STRATIGRAPHIC RELATIONS: The upper and lower contacts of the formation are not exposed. BIOSTRATIGRAPHY: from Collins et al. (1999), not specified to Early or Late Pliocene; direct date from Cheetham et al. (1999).

• bed-level stratigraphic resolution

Environment/lithology: deep-water; poorly lithified, burrowed, bioturbated, fine-grained, argillaceous sandstone and poorly lithified, burrowed, bioturbated, argillaceous siltstone

• ENVIRONMENT: from Collins et al. 1999 (p. 94), mixed middle neritic and outer nertic/upper bathyal, estimated depth of 125 m.
• GENERAL LITHOLOGY: from Collins et al. (1999), lower part of formation at the stratotype is moderately indurated, fine, silty sandstone and clayey siltstone, pervasively bioturbated and containing frequent, cemented, irregular burrow-concentrations and horizons with dense thalassinoid burrow systems.

Size class: macrofossils

Reposited in the BMNH, NMB, USNM

Collection methods: bulk, sieve,

• COVERAGE: Exhaustive for gastropoda, bivalvia and scaphopoda. COLLECTORS: unknown (PPP staff), 6/4/88. REPOSITORIES:collections presumably held at USNM, BMNH and Naturhistorisches Museum Basel (NMB),

Primary reference: L. S. Collins and A. G. Coates. 1999. A paleobiotic survey of Caribbean faunas from the Neogene of the Isthmus of Panama. Bulletins of American Paleontology (357) [A. Miller/A. Hendy/A. Hendy]more details

Purpose of describing collection: paleoecologic analysis

PaleoDB collection 79989: authorized by Austin Hendy, entered by Kate Bulinski on 31.03.2008, edited by Austin Hendy

Creative Commons license: CC BY (attribution)

Taxonomic list

• NOMENCLATURE: modern nomenclature, though only identified to subgenus resolution. IDENTIFIER: presumably identified by J. Jackson, J. Todd, H. Fortunato, and P. Jung.
Show authors, comments, and common names
Scaphopoda
 Gadilida - Gadilidae
Gadilopsis sp.
Gadilopsis sp. Woodring 1925 tusk shell
 Gadilida - Entalinidae
Bathoxiphus sp.
Bathoxiphus sp. Pilsbry 1897 tusk shell
Bivalvia
 Lucinida - Lucinidae
Gonimyrtea sp., Parvilucina sp.
Gonimyrtea sp. Marwick 1929 clam
Parvilucina sp. Dall 1901 clam
sensu lato
 Pholadida - Corbulidae
"Varicorbula sp." = Corbula (Varicorbula), Caryocorbula sp.
"Varicorbula sp." = Corbula (Varicorbula) Grant and Gale 1931 clam
Caryocorbula sp. Gardner 1926 clam
 Cardiida - Cardiidae
Laevicardium sp., Trigoniocardia sp., Microcardium sp.
Laevicardium sp. Swainson 1840 cockle
Trigoniocardia sp. Dall 1900 cockle
sensu lato
Microcardium sp. Thiele 1934 cockle
 Cardiida - Semelidae
Abra sp.
Abra sp. Lamarck 1818 abra
 Cardiida - Tellinidae
"Eurytellina sp." = Tellina (Eurytellina), Angulus sp.
"Eurytellina sp." = Tellina (Eurytellina) Fischer 1887 tellin clam
Angulus sp. Megerle 1811 tellin clam
 Cardiida - Veneridae
"Ventricolaria sp." = Globivenus, Gouldia sp.
"Ventricolaria sp." = Globivenus Coen 1934 venus clam
Gouldia sp. Adams 1847 venus clam
 Cardiida - Chamidae
Chama sp.
Chama sp. Linnaeus 1758 jewel box
 Cardiida - Kelliellidae
Kelliella sp.
Kelliella sp. Sars 1865 clam
 Carditida - Crassatellidae
Crassinella sp.
Crassinella sp. Guppy 1874 clam
 Mytilida - Crenellidae
Crenella sp.
Crenella sp. Brown 1827 clam
 Pectinida - Pectinoidae
? Spathochlamys sp., Spathochlamys sp., Argopecten sp.
? Spathochlamys sp. Waller 1993 scallop
Spathochlamys sp. Waller 1993 scallop
Argopecten sp. Monterosato 1889 calico scallop
 Pectinida - Propeamussiidae
Cyclopecten sp.
Cyclopecten sp. Verrill 1897 mud scallop
 Ostreida - Gryphaeidae
? Hyotissa sp.
? Hyotissa sp. Stenzel 1971 oyster
 Arcida - Limopsidae
Limopsis sp.
Limopsis sp. Sassi 1827 clam
 Arcida - Arcidae
Anadara (Rasia) sp.
Anadara (Rasia) sp. Gray 1857 ark
 Arcida - Glycymerididae
Tucetona sp.
Tucetona sp. Iredale 1939 clam
 Nuculida - Nuculidae
Nucula (Lamellinucula) sp.
Nucula (Lamellinucula) sp. Schenck 1944 nut clam
 Nuculanida - Nuculanidae
Saccella sp.
Saccella sp. Woodring 1925 pointed nut clam
Gastropoda
 Neotaenioglossa - Calyptraeidae
Crucibulum (Crucibulum) sp.
Crucibulum (Crucibulum) sp. Schumacher 1817 slipper shell
 Pseudomelanoidea - Pseudomelaniidae
? Miraclathurella sp., Compsodrillia sp., Carinodrillia (? Buchema) sp.
? Miraclathurella sp. Woodring 1928 snail
Compsodrillia sp. Woodring 1928 snail
 Sorbeoconcha - Caecidae
"Brochina sp." = Caecum (Brochina)
"Brochina sp." = Caecum (Brochina) Gray 1837 snail
 Sorbeoconcha - Vitrinellidae
Cyclostremiscus (Ponocyclus) sp.
 Sorbeoconcha - Vermetidae
Serpulorbis sp.
Serpulorbis sp. Sasso 1827 worm shell
 Sorbeoconcha - Xenophoridae
Xenophora sp.
Xenophora sp. Fischer von Waldheim 1807 carrier shell
 Sorbeoconcha - Strombidae
Strombus sp.
Strombus sp. Linnaeus 1758 conch
 Neogastropoda - Cystiscidae
Granulina sp., Persicula sp.
Granulina sp. Jousseaume 1888 snail
Persicula sp. Schumacher 1817 snail
 Neogastropoda - Marginellidae
Dentimargo sp., Volvarina sp.
Dentimargo sp. Cossmann 1899 margin shell
Volvarina sp. Hinds 1844 margin shell
 Neogastropoda - Muricidae
Haustellum sp., Typhis sp.
Haustellum sp. Schumacher 1817 murex snail
Typhis sp. Montfort 1810 murex snail
 Neogastropoda - Volutidae
Voluta sp.
Voluta sp. Linnaeaus 1758 volute
 Neogastropoda - Buccinidae
Antillophos (Antillophos) sp.
Antillophos (Antillophos) sp. Woodring 1928 true whelk
 Neogastropoda - Columbellidae
Sincola (? Sinaxila) sp., Mitrella sp., Nassarina (Nassarina) sp.
Sincola (? Sinaxila) sp. Olsson and Harbison 1953 snail
Mitrella sp. Risso 1826 snail
Nassarina (Nassarina) sp. Dall 1889 snail
 Neogastropoda - Nassariidae
Nassarius sp.
Nassarius sp. Duméril 1806 snail
sensu lato
 Neogastropoda - Ancillariidae
"Ancilla (Eburna) sp." = Eburna
"Ancilla (Eburna) sp." = Eburna Lamarck 1801 snail
 Neogastropoda - Olividae
Olivella (Minioliva) sp., Olivella (Macgintiella) sp.
Olivella (Minioliva) sp. Olsson 1956 olive snail
Olivella (Macgintiella) sp. Olsson 1956 olive snail
 Neogastropoda - Borsoniidae
Microdrillia sp.
Microdrillia sp. Casey 1903 snail
 Neogastropoda - Conidae
Conus sp.
Conus sp. Linnaeus 1758 cone shell
 Neogastropoda - Drilliidae
Syntomodrillia sp., Leptadrillia sp., Clathrodrillia sp.
Syntomodrillia sp. Woodring 1928 snail
Leptadrillia sp. Woodring 1928 snail
Clathrodrillia sp. Dall 1918 snail
 Neogastropoda - Terebridae
Strioterebrum sp.
Strioterebrum sp. Sacco 1891 auger snail
 Neogastropoda - Clathurellidae
Nannodiella sp., Glyphostoma (Glyphostoma) sp.
Nannodiella sp. Dall 1919 snail
 Neogastropoda - Mangeliidae
Ithycythara sp., Saccharoturris sp., Kurtziella (Kurtziella) sp., Tenaturris sp.
Ithycythara sp. Woodring 1928 snail
Saccharoturris sp. Woodring 1928 snail
Tenaturris sp. Woodring 1928 snail
 Neogastropoda - Raphitomidae
Pleurotomella sp.
Pleurotomella sp. Verrill 1872 snail
 Neogastropoda - Turridae
Polystira sp.
Polystira sp. Woodring 1928 turrid
 Neogastropoda - Costellariidae
Fusiturricula sp.
Fusiturricula sp. Woodring 1928 ribbed mitre
 Neogastropoda - Mitridae
Subcancilla sp.
Subcancilla sp. Olsson and Harbison 1953 mitre shell
 Sorbeoconcha - Naticidae
Natica (? Natica) sp., Polinices sp.
Natica (? Natica) sp. Scopoli 1777 moon snail
Polinices sp. Monfort 1810 moon snail
 Sorbeoconcha - Atlantidae
Atlanta sp.
Atlanta sp. Lesueur 1817 snail
 Sorbeoconcha - Cerithiopsidae
Cerithiopsis sp.
Cerithiopsis sp. Hanley 1849 snail
 Sorbeoconcha - Rissoidae
Rissoidae indet., Alvania sp.
Rissoidae indet. Gray 1847 snail
Alvania sp. Risso 1826 snail
 Cerithioidea - Cerithiidae
Alabina sp.
Alabina sp. Dall 1902 cerith snail
 Cerithioidea - Turritellidae
Vermicularia sp., Turritella sp.
Vermicularia sp. Lamarck 1799 turret shell
Turritella sp. Lamarck 1799 turret shell
 Vanikoroidea - Eulimidae
Eulimidae indet., Niso sp.
Eulimidae indet. Philippi 1853 snail
Niso sp. Risso 1826 snail
 Epitonioidea - Epitoniidae
Epitonium (Asperiscala) sp.
Epitonium (Asperiscala) sp. de Boury 1909 wentletrap
 Trochoidea - Trochidae
Trochidae indet., Microgaza sp., Calliostoma sp.
Trochidae indet. Rafinesque 1815 top snail
Microgaza sp. Dall 1881 top snail
Calliostoma sp. Swainson 1840 top snail
 Opisthobranchia - Haminoeidae
"Atys (Aliculastrum) sp." = Aliculastrum
"Atys (Aliculastrum) sp." = Aliculastrum Pilsbry 1896 snail
 Opisthobranchia - Retusidae
Volvulella (Volvulella) sp., Sulcoretusa sp.
Volvulella (Volvulella) sp. Newton 1891 snail
Sulcoretusa sp. Burch 1945 snail
 Opisthobranchia - Cylichnidae
Acteocina sp., Scaphander sp.
Acteocina sp. Gray 1847 barrel-bubble
Scaphander sp. Monfort 1810 snail
 Thecosomata - Cavoliniidae
Diacria sp., Cavolinia sp., Styliola sp., Clio sp.
Diacria sp. Gray 1847 snail
Cavolinia sp. Avilgaard 1791 snail
Styliola sp. Gray 1850 snail
Clio sp. Linnaeus 1767 snail
 Heterostropha - Architectonicidae
Pseudotorinia sp., Architectonica sp.
Pseudotorinia sp. Sacco 1892 snail
Architectonica sp. Röding 1798 sundial
 Heterostropha - Ringiculidae
"Ringincula (Ringiculella) sp." = Ringicula (Ringiculella)
"Ringincula (Ringiculella) sp." = Ringicula (Ringiculella) Sacco 1892 snail
 Heterostropha - Murchisonellidae
Anisocycla (Saccoina) sp.
Anisocycla (Saccoina) sp. Dall and Bartsch 1904 snail
 Heterostropha - Pyramidellidae
Pyramidellidae indet., Eulimella (Ebalina) sp., Ondina sp., Besla sp.
Pyramidellidae indet. Gray 1840 snail
Eulimella (Ebalina) sp. Thiele 1929 snail
Ondina sp. de Folin 1870 snail
Besla sp. Dall and Bartsch 1904 snail
Gymnolaemata
 Cheilostomata - Gemelliporellidae
Gemelliporella glabra
 Cheilostomata - Mamilloporidae
Mamillopora "n. sp. 1"
 Cheilostomata - Phidoloporidae
Schedocleidochasma cleidostoma
 Cheilostomata - Petraliidae
Petraliella bisinuata
 Cheilostomata - Smittinidae
Parasmittina sp.
 Cheilostomata - Lepraliellidae
Celleporaria albirostris
 Cheilostomata - Membraniporidae
Cupuladria aff. canariensis, Cupuladria aff. biporosa
Cupuladria aff. biporosa Canu and Bassler 1923