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Coptoclavidae

Insecta - Coleoptera - Coptoclavidae

Synonymy list
YearName and author
1961Coptoclavidae Ponomarenko p. 68
1992Coptoclavidae Carpenter
2007Coptoclavidae Soriano et al. p. 527
2011Coptoclavidae Bouchard et al.
2022Coptoclavidae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCoptoclavidaePonomarenko 1961
familyCoptoclavidaePonomarenko 1961

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Coptoclavidae Ponomarenko 1961
show all | hide all
G. †Agrascapha Lin 1992
+
†Agrascapha curta Lin 1992
G. †Amblycephalonius Bode 1953
+
+
Invalid names: Barycephalus nudatus Bode 1953 [synonym], Tricyrtus tenuistriatus Bode 1953 [synonym]
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
+
G. †Charonoscapha Ponomarenko 1977
+
†Charonoscapha grossa Ponomarenko 1977
†Charonoscapha ovata Ponomarenko 1977
G. †Charonoscaphidia Ponomarenko 1977
+
†Charonoscaphidia elongata Ponomarenko 1977
G. †Coptoclavia Ping 1928
+
†Coptoclavia africana Teixeira 1975
Subfm. †Coptoclavinae Ponomarenko 1961
+
G. †Bolbonectes Ponomarenko 1987
+
†Bolbonectes intermedius Ponomarenko 1987
†Bolbonectes occidentalis Ponomarenko 1993
†Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs 2000
G. †Coptoclava Ping 1928
+
†Coptoclava longipoda Ping 1928
G. †Hoyaclava Soriano et al. 2007
+
†Hoyaclava buscalionae Soriano et al. 2007
G. †Megacoptoclava Ponomarenko and Martínez-Delclòs 2000
+
†Megacoptoclava longiurogomphia Ponomarenko and Martínez-Delclòs 2000
G. †Tigrivia Bao et al. 2023
+
†Tigrivia baii Bao et al. 2023
Subfm. †Coptoclaviscinae Soriano et al. 2007
+
G. †Coptoclavella Ponomarenko 1980
+
†Coptoclavella elegans Ponomarenko 1980
†Coptoclavella inexpecta Soriano et al. 2007
†Coptoclavella jurassica Ponomarenko 2014
†Coptoclavella minor Ponomarenko 1980
†Coptoclavella purbeckensis Ponomarenko et al. 2005
†Coptoclavella striata Ponomarenko 1986
†Coptoclavella vittata Ponomarenko 1986
G. †Coptoclavisca Ponomarenko 1987
+
†Coptoclavisca nigricollinus Ponomarenko 1987
G. †Holcoptera Handlirsch 1906
+
†Holcoptera alisonae Kelly et al. 2017
†Holcoptera giebeli Handlirsch 1906
†Holcoptera pigmentatus Kelly et al. 2017
†Holcoptera rasnitsyni Ponomarenko and Fedorenko 2022
†Holcoptera schlotheimi Giebel 1856
+
Invalid names: Holcoptera confluens Cockerell 1915 [synonym]
†Holcoptera solitensis Kelly et al. 2017
Invalid names: Holcoelytrum Handlirsch 1906 [synonym], Stargelytron Ponomarenko 2015 [synonym]
G. †Euroscapha Lin 1992
+
Subfm. †Hispanoclavinae Soriano et al. 2007
+
G. †Hispanoclavina Soriano et al. 2007
+
†Hispanoclavina diazromerali Soriano et al. 2007
†Hispanoclavina gratshevi Soriano et al. 2007
Subfm. †Timarchopsinae Wang et al. 2010
+
G. †Daohugounectes Wang et al. 2009
+
†Daohugounectes primitivus Wang et al. 2009
G. †Protonectes Prokin and Ponomarenko 2013
+
†Protonectes germanicus Prokin and Ponomarenko 2013
G. †Timarchopsis Brauer et al. 1889
+
†Timarchopsis aquaticus Ponomarenko 1977
†Timarchopsis cyrenaicus Ponomarenko 1977
†Timarchopsis czekanowskii Brauer et al. 1889
†Timarchopsis gigas Ponomarenko 1977
†Timarchopsis gobiensis Ponomarenko 1987
†Timarchopsis latus Ponomarenko 1977
†Timarchopsis longus Ponomarenko 2014
†Timarchopsis mongolicus Ponomarenko 1985
†Timarchopsis sainshandensis Ponomarenko 1987
Invalid names: Necronectes Ponomarenko 1977 [synonym]
Invalid names: Necronectinae Ponomarenko 1977 [replaced]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Bithynian to the top of the Albian or 244.99000 to 100.50000 Ma

Collections (137 total)


Time interval Ma Country or state Original ID and collection number
Bithynian244.99 - 244.24Germany Stargelytron altus (175694)
Bithynian244.99 - 244.24Germany (Bavaria) Stargelytron altus (115969)
Carnian237.0 - 227.3China (Xinjiang) Agrascapha curta, Euroscapha hoplopoda, Euroscapha rectabdomena (156325)
Late/Upper Triassic237.0 - 201.4Angola Coptoclavia africana (210174)
Tuvalian233.6 - 227.3Germany Coptoclavidae indet. (163985) Protonectes germanicus, Coptoclavidae indet. (153474) Stargelytron larissae (175695)
Norian227.3 - 205.7USA (North Carolina) Holcoptera solitensis (189406 189407)
Norian227.3 - 205.7Ukraine (Kharkiv) Holcoptera rasnitsyni (105057)
Sevatian211.18 - 205.7Italy (Lombardia) Holcoptera schlotheimi (94207)
Rhaetian205.7 - 201.4United Kingdom (England) Harpalinae indet. 1, Harpalinae indet. (114643) Holcoptera pigmentatus (113957) Holcoptera schlotheimi (114644 115223 115246 187761) Holcoptera schlotheimi, Holcoptera giebeli (113961)
Rhaetian - Early/Lower Hettangian205.7 - 199.5United Kingdom (England) Holcoptera confluens (124688)
Early/Lower Hettangian201.4 - 199.5United Kingdom (England) Holcoptera schlotheimi (113956 113958 123986)
Planorbis201.4 - 199.5United Kingdom (England) Holcoptera schlotheimi (115460 123982)
Hettangian - Sinemurian201.4 - 192.9USA (Connecticut) Holcoptera giebeli (153636) Holcoptera schlotheimi (187762)
Hettangian - Sinemurian201.4 - 192.9USA (Massachusetts) Holcoptera giebeli (153637)
Obtusum199.5 - 192.9United Kingdom (Dorset) Holcoptera alisonae (189408) Holcoptera giebeli, Holcoptera alisonae (189405)
Obtusum199.5 - 192.9United Kingdom (England) Holcoelytrum giebeli, Holcoptera schlotheimi (114638 114639) Holcoptera alisonae (189404) Holcoptera giebeli (114637 189411)
Turneri199.5 - 192.9United Kingdom (Dorset) Holcoptera giebeli, Holcoptera alisonae (189409 189410) Holcoptera schlotheimi, Holcoptera giebeli (187763)
Early/Lower Toarcian184.2 - 182.9Germany (Niedersachsen) Amblycephalonius tenuistriatus, Tricyrtus tenuistriatus (107555) Barycephalus nudatus (105002)
Toarcian184.2 - 174.7Kyrgyzstan (Batken) Necronectes latus (105003)
Toarcian184.2 - 174.7Russian Federation Necronectes aquaticus (123146)
Toarcian184.2 - 174.7Russian Federation (Irkutsk) Stygeonectes jurassicus (104946 113471) Timarchopsis czekanowskii (114696)
Aalenian174.7 - 170.9Mongolia Timarchopsis sainshandensis (123958)
Bathonian168.2 - 165.3China (Xinjiang) Tuhanectes xinjiangensis (165545)
Bathonian168.2 - 165.3Mongolia Timarchopsis mongolicus (119621)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Daohugounectes primitinus (125080 125081 125082) Daohugounectes primitivus (113371)
Callovian - Oxfordian165.3 - 154.8Russian Federation (Chita) Coptoclavella minor (86023)
Callovian - Oxfordian165.3 - 154.8Russian Federation (Buryat) Stygeonectes jurassicus (114660)
Callovian - Kimmeridgian165.3 - 149.2Russian Federation (Chita) Coptoclavella minor (125485)
Oxfordian161.5 - 154.8Russian Federation (Buryatia) Necronectes aquaticus, Stygeonectes jurassicus (123147) Stygeonectes jurassicus (113483 123194 123195)
Oxfordian161.5 - 154.8Russian Federation (Buryat) Stygeonectes jurassicus, Bolbonectes occidentalis (105017)
Oxfordian - Kimmeridgian161.5 - 149.2Russian Federation (Buryat) Stygeonectes jurassicus (104950)
Late/Upper Jurassic161.5 - 143.1Russian Federation Stygeonectes jurassicus (114317)
Kimmeridgian154.8 - 149.2Kazakhstan Charonoscapha grossa, Charonoscapha ovata (105426) Charonoscapha ovata (113439 115717) Charonoscapha ovata, Charonoscaphidia elongata (105971) Charonoscaphidia elongata (106102 114238) Exedia plana (106101) Necronectes gigas, Exedia plana, Charonoscapha grossa (105970)
Kimmeridgian - Valanginian154.8 - 132.6Russian Federation (Chita) Coptoclavella minor (152182)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Actea sphinx, Prodytiscus eichstaettensis, Silphites cetoniformis, Ditomoptera dubia, Cerambycites dubius, Prodytiscus longispinosus (123985) Actea sphinx, Pseudohydrophilus avitus, Ditomoptera dubia (123144) Blabera avita (152935) Oryctes pluto, Cetonia defossa, Buprestis lapidelythris, Palaeobelostoma hartingi (123906) Pseudohydrophilus longispinosus, Cerambycites minor, Sphenoptera sphinx, Cerambycites dubius (133578)
Early/Lower Tithonian149.2 - 145.06Germany (Bavaria) Actea sphinx, Pseudohydrophilus avitus, Ditomorptera dubia, Ditomorptera minor (122586 139876)
Tithonian149.2 - 143.1Mongolia Coptoclavidae indet. (121393) Coptoclavidae indet., Coptoclavella jurassica, Timarchopsis longus (104748) Timarchopsis longus (121394 152133)
Tithonian149.2 - 143.1Mongolia (Dund-Gov’ Aymag) Timarchopsis gobiensis (117695)
Tithonian149.2 - 143.1Russian Federation (Chita) Stygeonectes jurassicus (152174 152178 152181)
Tithonian149.2 - 143.1Russian Federation Bolbonectes intermedius (123957 125408)
Late/Upper Berriasian139.1 - 137.05United Kingdom (England) Coptoclavella purbeckensis (103833)
Valanginian - Barremian137.05 - 121.4Mongolia (Dornogov) Coptoclavisca nigricollinus (113407)
Hauterivian132.6 - 125.77Algeria Necronectes cyrenaicus (123148)
Hauterivian132.6 - 125.77Russian Federation (Chita) Coptoclava longipoda (152177) Coptoclavella minor (125409)
Late/Upper Hauterivian132.6 - 125.77China (Hebei) Coptoclava longipoda (111077)
Barremian125.77 - 121.4Russian Federation (Chita) Coptoclava longipoda (152180) Stygeonectes jurassicus (152176) Stygeonectes jurassicus, Bolbonectes intermedius (151642)
Early/Lower Barremian125.77 - 121.4China Coptoclava longipoda (235409)
Early/Lower Barremian125.77 - 121.4Spain Bolbonectus lithographicus (123437) Coptoclavella inexpecta (104052)
Late/Upper Barremian125.77 - 121.4Spain (Castilla-La Mancha) Hispanoclavina diazromerali, Hispanoclavina gratshevi, Hoyaclava buscalionae, Ovonectes pilosum, Megacoptoclava longiurogomphia (28006)
Late/Upper Barremian125.77 - 121.4China (Jilin) Coptoclava longipoda (155010)
Early/Lower Aptian121.4 - 119.57China (Zhejiang) Coptoclava longipoda (155611)
Early/Lower Aptian121.4 - 119.57Russian Federation (Chita) Coptoclava longipoda (152179) Stygeonectes jurassicus, Coptoclavella minor (119499)
Aptian121.4 - 113.2China (Liaoning) Coptoclava longipoda (150814 151536)
Aptian121.4 - 113.2China (Jilin) Coptoclava longipoda (151876 172391)
Aptian121.4 - 113.2China (Nei Mongol) Coptoclava longipoda (156008)
Aptian121.4 - 113.2Russian Federation (Buryatia) Coptoclava longipoda (105969 113350)
Aptian121.4 - 113.2China (Gansu) Coptoclava longipoda (154779)
Aptian121.4 - 113.2China (Shandong) Coptoclava longipoda (106313 154887 154888 154890 154894 154897 156554 156555 156601 161254) Coptoclavisca grandioculus (125196)
Aptian121.4 - 113.2Mongolia Coptoclava longipoda (123933) Coptoclavella elegans (151594 151596) Coptoclavella elegans, Coptoclavella minor (151591)
Aptian121.4 - 113.2Russian Federation (Chita) Coptoclava longipoda (126001 138148) Stygeonectes jurassicus (138072)
Aptian121.4 - 113.2Mongolia (Govi-Altai) Coptoclavella striata, Coptoclavella vittata (122332) Coptoclavella vittata (136020)
Aptian121.4 - 113.2Russian Federation Coptoclava longipoda (124811)
Aptian121.4 - 113.2China (Hebei) Tigrivia baii (241724)
Albian113.2 - 100.5South Korea Coptoclava sp. (184781 184782 184783)