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Nanhsiungchelyidae
Taxonomy
Nanhsiungchelyidae was named by Yeh (1966). Its type is Nanhsiungchelys.
It was assigned to Trionychoidea by Hirayama et al. (2001); to Cryptodira by Yeh (1966) and Tong et al. (2012); to Adocusia by Syromyatnikova and Danilov (2009), Danilov et al. (2013) and Syromyatnikova and Danilov (2013); to Eucryptodira by Peng et al. (2001), Hutchison and Holroyd (2003), Hutchison et al. (2013) and Mallon and Brinkman (2018); to Trionychoidae by Ke et al. (2021); and to Cryptodira by Joyce and Norell (2005), Ke et al. (2023) and Ke et al. (2024).
It was assigned to Trionychoidea by Hirayama et al. (2001); to Cryptodira by Yeh (1966) and Tong et al. (2012); to Adocusia by Syromyatnikova and Danilov (2009), Danilov et al. (2013) and Syromyatnikova and Danilov (2013); to Eucryptodira by Peng et al. (2001), Hutchison and Holroyd (2003), Hutchison et al. (2013) and Mallon and Brinkman (2018); to Trionychoidae by Ke et al. (2021); and to Cryptodira by Joyce and Norell (2005), Ke et al. (2023) and Ke et al. (2024).
Subtaxa
Anomalochelys, Basilemys, Bulganemys, Bulganemys, Charitonyx, Hanbogdemys, Hanbogdemysa, Jiangxichelys, Kharakhutulia, Nanhsiungchelys (type genus), Xianyuechelys, Yuchelys, Zangerlia
Synonyms
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Synonymy list
Year | Name and author |
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1966 | Nanhsiungchelyidae Yeh p. 191 |
1980 | Charitonyx Chkhikvadze |
2000 | Hanbogdemysa Sukhanov |
2000 | Bulganemys Sukhanov p. 342 fig. 17.24 |
2000 | Hanbogdemysa Sukhanov pp. 3340-342 fig. 17.23 |
2000 | Bulganemys Sukhanov and Narmandakh p. 342 |
2001 | Nanhsiungchelyidae Hirayama et al. p. 129 |
2001 | Nanhsiungchelyidae Peng et al. p. 26 |
2003 | Nanhsiungchelyidae Hutchison and Holroyd |
2005 | Nanhsiungchelyidae Joyce and Norell p. 4 |
2009 | Nanhsiungchelyidae Syromyatnikova and Danilov p. 75 |
2012 | Nanhsiungchelyidae Tong et al. |
2013 | Nanhsiungchelyidae Danilov et al. |
2013 | Nanhsiungchelyidae Hutchison et al. |
2013 | Nanhsiungchelyidae Syromyatnikova and Danilov |
2018 | Nanhsiungchelyidae Mallon and Brinkman |
2021 | Nanhsiungchelyidae Ke et al. |
2023 | Nanhsiungchelyidae Ke et al. |
2024 | Nanhsiungchelyidae Ke et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Nanhsiungchelyidae Yeh 1966
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G. †Basilemys Hay 1902
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†Basilemys gaffneyi Sullivan et al. 2012
†Basilemys imbricarius Cope 1876
†Basilemys morrinensis Mallon and Brinkman 2018
†Basilemys nobilis Hay 1911
†Basilemys ogmius Cope 1875
†Basilemys praeclara Hay 1911
†Basilemys sinuosa Riggs 1906
†Basilemys variolosa Cope 1876
G. †Bulganemys Sukhanov and Narmandakh 2006
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Invalid names: Bulganemys jaganchobili Sukhanov and Narmandakh 2000 [nomen nudum]
G. †Hanbogdemys Sukhanov and Narmandakh 2006
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†Hanbogdemys jaganchobili Sukhanov and Narmandakh 2006
†Hanbogdemys orientalis Sukhanov and Narmandakh 1975
G. †Jiangxichelys Tong and Mo 2010
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†Jiangxichelys ganzhouensis Tong and Mo 2010
†Jiangxichelys neimongolensis Brinkman and Peng 1996
G. †Nanhsiungchelys Yeh 1966
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†Nanhsiungchelys wuchingensis Yeh 1966
†Nanhsiungchelys yangi Ke et al. 2023
G. †Zangerlia Mlynarski 1972
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†Zangerlia dzamynchondi Sukhanov and Narmandakh 2006
†Zangerlia testudinimorpha Mlynarski 1972
†Zangerlia ukhaachelys Joyce and Norell 2005
Invalid names: Zangerlia dzamynchondi Sukhanov and Narmandakh 2000 [nomen nudum]
Invalid names: Bulganemys Sukhanov and Narmandakh 2000 [nomen nudum], Charitonyx Chkhikvadze 1980 [nomen dubium], Hanbogdemysa Sukhanov 2000 [nomen nudum]
Diagnosis
Reference | Diagnosis | |
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H.-k. Yeh 1966 | With the diagnosis of the type genus Nanhsiungchelys. | |
R. Hirayama et al. 2001 | Large trionychoids with shell length well exceeding over 50 cm, and up to 90 cm; shell surface deeply sculptured by the linearly arranged ridges and vallies; marginal region of carapace and plastron thickened; neural series complete to suprapygals except for Basilemys nobilis; anterior portion of epiplastron projected and enormously thickened; humero-pectoral sulcus deeply intersecting the entoplastron; incisura columellae auris closed posterior; coracoid fan-shaped; thelial process of ilium absent; two or fewer short and stout phalanges in manus and pes; limbs covered by epidermal bones with pitted sculpture. |
Measurements
No measurements are available
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Source: subo = suborder, c = class, subp = subphylum | |||||
References: Carroll 1988, Hendy et al. 2009, Uetz 2005 |
Age range: base of the Late/Upper Aptian to the top of the Lancian or 122.46000 to 66.00000 Ma
Collections (159 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Aptian | Japan (Fukui) | Basilemys sp. (38020) | |
Aptian - Albian | Mongolia (Omnogov) | Nanhsiungchelyidae indet. (154401) | |
Aptian - Albian | Mongolia (Dornogov) | Nanhsiungchelyidae indet. (168802) | |
Late/Upper Aptian - Early/Lower Albian | Mongolia (Dornogov) | Nanhsiungchelyidae indet. (53159) | |
Late/Upper Albian - Cenomanian | Mongolia (Dalanzagad) | Nanhsiungchelyidae indet. (60457) | |
Cenomanian | Japan (Hokkaido) | Anomalochelys angulata (129339) | |
Cenomanian - Turonian | China (Henan) | Yuchelys nanyangensis (196440) | |
Cenomanian - Santonian | Mongolia (Omnogov) | Basilemys orientalis (67974) Nanhsiungchelyidae indet. (13411 55254 108340 127666) Nanhsiungchelyidae indet., Hanbogdemys sp. (79724) | |
Cenomanian - Santonian | Kazakhstan (Ongtüstik Qazaqstan) | Nanhsiungchelyidae indet. (58720) | |
Cenomanian - Santonian | Mongolia (Dornogov) | Nanhsiungchelyidae indet. (108342 189006) Nanhsiungchelyidae indet., Kharakhutulia kalandadzei, Hanbogdemys sp., Charitonyx tajanikolaevae (118167) | |
Cenomanian - Middle Campanian | China (Guangdong) | Nanhsiungchelys yangi (230296) | |
Early/Lower Cenomanian | Uzbekistan (Navoi) | Basilemys sp. (58208 64685) | |
Early/Lower Cenomanian | Uzbekistan (Karakalpakstan) | Nanhsiungchelyidae indet. (58466 64689 189009) | |
Late/Upper Cenomanian - Early/Lower Turonian | Japan (Kumamoto) | Nanhsiungchelyidae indet. (51963) | |
Turonian | Japan (Iwate) | Nanhsiungchelyidae indet. (165694) | |
Early/Lower Turonian - Middle Campanian | China (Henan) | Nanhsiungchelyidae indet. (91210) | |
Coniacian - Santonian | Japan | Basilemys sp. (189008) | |
Late/Upper Santonian | Canada (Alberta) | Basilemys sp. (48834 71485) | |
Campanian | China (Nei Mongol) | Zangerlia neimongolensis (131829) | |
Campanian | Mongolia (Ömnögovi Aimag) | Nanhsiungchelyidae indet. (130798) | |
Campanian | Mongolia (Omnogov (=Southern Gobi)) | Zangerlia dzamynchondi (137600) | |
Campanian | Canada (Alberta) | Basilemys variolosa (177143) | |
Campanian | Mongolia (Omnogov) | Adocus sp. (48983) Bulganemys jaganchobili (127739) | |
Early/Lower Campanian | USA (Texas) | Basilemys sp. (200396) | |
Early/Lower Campanian | USA (Utah) | Basilemys sp. (167530) | |
Early/Lower Campanian - Middle Campanian | USA (Utah) | Basilemys sp. (167916) | |
Judithian | USA (Wyoming) | Basilemys sp. (14458) | |
Late/Upper Campanian | Mongolia (Omnogov Aimak) | Basilemys sp. (37184) | |
Late/Upper Campanian | USA (New Mexico) | Aspideretes sp. (64094) Basilemys gaffneyi (47193 62317 76107 96814 138830 144519 154382 154386 154419 154594 154620 154621 154622 154623 154624 154625 154626 154627) Basilemys nobilis (100677) Basilemys sp. (36449 55882 55883) | |
Late/Upper Campanian | Mongolia (Omnogov) | Basilemys sp. (45090 127668) Basilemys sp., Nanhsiungchelyidae indet. (127678) Testudines indet. (46152) | |
Late/Upper Campanian | Canada (Alberta) | Basilemys sp. (36264 221937) Basilemys variolosa (22697 34320 34322) | |
Late/Upper Campanian | Mexico (Baja California) | Basilemys sp. (197022) | |
Late/Upper Campanian | USA (Utah) | Basilemys nobilis (187686 187690 187698 187702 187711) | |
Late/Upper Campanian | USA (Montana) | Basilemys sp. (22689 59373) | |
Late/Upper Campanian | Mexico (Chihuahua) | Basilemys sp. (35707 210630) | |
Middle Campanian | Canada (Alberta) | Basilemys sp. (45566 181024) | |
Middle Campanian | Mongolia (Omngov) | Testudinata indet. (127677) | |
Middle Campanian | USA (Utah) | Basilemys sp. (167521) | |
Middle Campanian - Late/Upper Campanian | USA (Montana) | Basilemys sp. (50907 51682 222122) Ceratopsia indet. (14462) | |
Middle Campanian - Late/Upper Campanian | USA (Texas) | Basilemys sp. (25073 68012 68014 68015 212185) | |
Middle Campanian - Late/Upper Campanian | Mongolia (Omnogov) | Nanhsiungchelyidae indet. (154408 189005) Testudinata indet. (181536) Zangerlia testudinimorpha (189007) | |
Middle Campanian - Late/Upper Campanian | Canada (Alberta) | Adocus variolosus (150748 150749) Basilemys indet. (167986) | |
Middle Campanian - Late/Upper Campanian | Canada (Saskatchewan) | Basilemys sp. (14494) | |
Late/Upper Campanian - Maastrichtian | Canada (Alberta) | Basilemys sp. (45757) | |
Maastrichtian | USA (California) | Basilemys sp. (76729) | |
Maastrichtian | China (Guangdong) | Nanhsiungchelys wuchingensis (71863) | |
Maastrichtian | China (Jiangxi) | Jiangxichelys ganzhouensis (127742) Nanhsiungchelys sp., Jiangxichelys ganzhouensis (147070) Xianyuechelys yingliangi (235257) | |
Maastrichtian | USA (Montana) | Basilemys sp. (85498) | |
Maastrichtian | USA (North Dakota) | Basilemys sinuosa (45313) Basilemys sp. (231466) | |
Early/Lower Maastrichtian | Canada (Alberta) | Basilemys morrinensis (193696) | |
Lancian | USA (North Dakota) | Basilemys sp. (137706 137715 137724 137728 137732) | |
Lancian | USA (Colorado) | Basilemys sp. (210050) | |
Lancian | USA (Montana) | Basilemys sp. (14508 14513 14536 14654) | |
Lancian | USA (Wyoming) | Aspideretoides sp. (14544 70343) Basilemys sinuosa (75290) Basilemys sp. (14514 14519 14542 14565 14579 75280 75297 75443 137795) Basilemys sp., Basilemys sinuosa (14505) | |
Lancian | USA (South Dakota) | Basilemys sinuosa (48235) | |
Late/Upper Maastrichtian | USA (Montana) | Basilemys sinuosus (99441) Basilemys sp. (189003) | |
Late/Upper Maastrichtian | Canada (Saskatchewan) | Basilemys sp. (45675 54105) | |
Late/Upper Maastrichtian | USA (Colorado) | Basilemys sinuosa (49538 49539 176740 176741) | |
Late/Upper Maastrichtian | USA (South Dakota) | Basilemys sinuosa (45390) Basilemys sp. (50082 150891 150893 224484) | |
Late/Upper Maastrichtian | Canada (Alberta) | Compsemys ogmius (58157) | |
Late/Upper Maastrichtian | USA (Wyoming) | Basilemys sp. (14585 137798) | |
Late/Upper Maastrichtian | USA (North Dakota) | Basilemys sinuosa (45099 45118 45144 45145 45381) Basilemys sp. (137720) |