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Carabidae (ground beetle)

Insecta - Coleoptera - Carabidae

Taxonomy
Carabici was named by Latreille (1802). It is extant. Its type is Carabus.

It was corrected as Carabiques by Dejean (1821); it was corrected as Carabidae by LeConte (1861), Cockerell (1921), Cockerell (1921), Crowson (1981), Hong (1984), Whalley (1985), Zhang (1989), Carpenter (1992), Lawrence and Newton (1995), Bouchard et al. (2011) and Cai et al. (2022).

It was assigned to Caraboidea by Crowson (1981); to Coleoptera by Latreille (1802), Dejean (1821), LeConte (1861), Cockerell (1921), Cockerell (1921), Hong (1984), Whalley (1985) and Zhang (1989); and to Adephaga by Carpenter (1992), Lawrence and Newton (1995), Bouchard et al. (2011) and Cai et al. (2022).

Subtaxa
Antemnacrassa, Apotomopterus, Ataktosites, Bembidium, Brachininae, Brachinites, Broscinae, Carabidium, Carabinae, Carabites, Carabites, Carabopteron, Cavicarabus, Celeripedes, Cicindelites, Conjunctiinae, Cratacanthus florissantensis, Elaphrinae, Elaphrotites, Fangshania, Feronites, Fonsecacarabus, Fossores, Gehringiinae, Glenopterus, Hadrocephalus, Harpalinae (syn. Licininae), Harpalomimes, Hebeicarabus, Lithocantharis, Loricera exita, Loricerinae, Meileyingia, Metagrilium, Migadopinae, Nebriinae, Nebrorabus, Neothanes, Notokistus, Omophroninae, Paussinae (syn. Ozaenidae), Platyninae, Procarabites, Protopaussinae, Protorabinae, Rhopalochlaenius, Rhysodinae, Scaritinae (syn. Promecognathinae), Siagoninae, Sinocarabus, Stereocerus, Sunocarabus, Trechinae (syn. Patrobinae), Trechoides, Trechus fractus, Tremembecarabus, Wongyekokia, Wuchangicarabus, Yunnanocarabus

Synonyms
Synonymy list
YearName and author
1802Carabici Latreille p. 80
1821Carabiques Dejean p. 1
1861Carabidae LeConte p. 4
1900Loricera exita Scudder p. 70 figs. Pl 6, fig 1
1912Trechus fractus Wickham p. 6 figs. Pl 3, fig 1
1917Cratacanthus florissantensis Wickham p. 464 figs. Pl 37 fig 2
1921Carabidae Cockerell
1981Carabidae Crowson p. 694
1984Carabidae Hong p. 3
1985Carabidae Whalley
1989Carabidae Zhang p. 420
1992Carabidae Carpenter
1995Carabidae Lawrence and Newton, Jr.
2011Carabidae Bouchard et al. p. 102
2022Carabidae Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
RankNameAuthor
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyCarabidae()
familyCarabidae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Carabidae Latreille 1802 [ground beetle]
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G. †Antemnacrassa Gomez Pallerola 1979
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†Antemnacrassa albomaculata Nikolajev 2004
†Antemnacrassa albosulcata Nikolajev 2004
†Antemnacrassa geminata Nikolajev 2004
†Antemnacrassa incerta Ponomarenko 1990
†Antemnacrassa kazakhstanica Nikolajev 2007
†Antemnacrassa latitibialis Nikolajev 1992
†Antemnacrassa maculata Nikolajev 1992
†Antemnacrassa magna Nikolajev 2004
†Antemnacrassa monreali Gomez Pallerola 1979
†Antemnacrassa nebulosa Nikolajev 2004
†Antemnacrassa nigrimontana Nikolajev 1992
†Antemnacrassa picturata Nikritin 1977
†Antemnacrassa punctata Ponomarenko 1986
†Antemnacrassa sulcata Nikritin 1977
†Antemnacrassa vitimensis Nikritin 1977
G. Apotomopterus Hope 1838
G. †Ataktosites Haupt 1956
+
†Ataktosites palustris Haupt 1956
G. Bembidium Latreille 1802
+
Subg. Bembidium (Campa) Motschulsky 1844
+
Subg. Bembidium (Testidium) Motschulsky 1864
†Bembidium absolutum Giebel 1856
†Bembidium cyaneomicans Piton 1940
†Bembidium exoletum Scudder 1876
†Bembidium expletum Scudder 1900
†Bembidium florissantensis Wickham 1913
†Bembidium fragmentum Scudder 1890
†Bembidium glaciatum Scudder 1890
†Bembidium haywardi Scudder 1900
†Bembidium infernum Heer 1856
†Bembidium laevigatum Förster 1891
†Bembidium obductum Scudder 1900
†Bembidium praeteritum Scudder 1900
†Bembidium saportanum Oustalet 1874
†Bembidium tumulorum Scudder 1900
†Bembidium vanum Scudder 1900
†Bembidium vestigium Scudder 1900
Subfm. Brachininae Bonelli 1810
+
Tr. Brachinini Bonelli 1810
+
Subtr. Brachinina Bonelli 1810
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G. Brachinus Weber 1801
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G. †Brachinites Fritsch 1882
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†Brachinites truncatus Fritsch 1882
Subfm. Broscinae Hope 1838
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Tr. Broscini Hope 1838
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Subtr. Broscina Hope 1838
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G. Miscodera Eschscholtz 1830
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Miscodera arctica Paykull 1798
Subtr. Nothobroscina Roig-Juñent 2000
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G. Mecodema Blanchard 1843
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Mecodema crenaticolle Redtenbacher 1868
G. †Carabidium Westwood 1854
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†Carabidium dejeanianum Westwood 1854
Subfm. Carabinae Latreille 1802
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G. †Atrirabus Hong and Wang 1990
+
†Atrirabus shandongensis Hong and Wang 1990
†Atrirabus tuanwangensis Hong and Wang 1990
Tr. Carabini Latreille 1802
+
G. Calosoma Weber 1801
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Subg. Calosoma (Campalita) Motschulsky 1865
+
†Campalita deplanatum Heer 1860
†Calosoma agassizi Barthelemy-Lapommeraye 1846
†Calosoma brunneum Hong 1985
†Calosoma calvini Wickham 1909
†Calosoma caraboides Heer 1860
†Calosoma catenulatum Heer 1860
†Calosoma ceresti Nel 1987
†Calosoma cockerelli Wickham 1910
†Calosoma emmonsi Scudder 1900
†Calosoma escheri Heer 1860
†Calosoma fernquisti Cockerell 1925
†Calosoma grasui Paucă and Ciobanu 1978
†Calosoma heeri Scudder 1895
Calosoma inquisitor Linnaeus 1758
†Calosoma jaccardi Heer 1860
†Calosoma kimi Lee and Nam 2021
Calosoma maderae Fabricius 1775
†Calosoma nauckianum Heer 1860
Calosoma semilaeve LeConte 1852
Calosoma sycophanta Linnaeus 1758 [forest caterpillar hunter]
G. Carabus Linnaeus 1758
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Subg. Carabus (Archicarabus) Seidlitz 1887
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Subg. Carabus (Aulonocarabus) Reitter 1896
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Subg. Carabus (Damaster) Kollar 1836
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Subg. Carabus (Hemicarabus) Gehin 1876
Subg. Carabus (Homoeocarabus) Reitter 1896
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Carabus (Homoeocarabus) maeander Fischer von Waldheim 1822
Subg. Carabus (Megadontus) Solier 1848
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Subg. Carabus (Melancarabus) Thomson 1875
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Subg. Carabus (Ohomopterus) Reitter 1896
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Subg. Carabus (Tachypus) Weber 1801
+
Subg. Carabus (Tanaocarabus) Reitter 1896
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†Carabus agassizi Barthelemy-Lapommeraye 1846
Carabus clathratus Linnaeus 1761
Carabus clatratus Linnaeus 1761
†Carabus foveolatus Piton 1935
Carabus granulatus Linnaeus 1758
Carabus hortensis Linnaeus 1758
†Carabus jeffersoni Scudder 1900
Carabus kollari Palliardi 1825
Carabus maderae Fabricius 1775
†Carabus maeandroides Lomnicki 1894
†Carabus mecothoracus Zhang 1989
†Carabus neli Deuve 1998
†Carabus novalensis Omboni 1886
†Carabus ovalus Hong and Wang 1986
†Carabus praearvensis Lomnicki 1894
Invalid names: Calosomini Bonelli 1810 [synonym]
Tr. Cychrini Perty 1830
+
G. Cychrus Fabricius 1794
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Cychrus caraboides Linnaeus 1758
G. Scaphinotus Dejean 1826
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Subg. Scaphinotus (Brennus) Motschulsky 1866
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Scaphinotus (Brennus) marginatus Fischer von Waldheim 1820
Subg. Scaphinotus (Nomaretus) Leconte 1853
+
†Nomaretus serus Scudder 1900
Scaphinotus interruptus Menetries 1843
†Scaphinotus minor Horn 1876
†Scaphinotus wheatleyi Horn 1876
G. Sphaeroderus Dejean 1831 [small snail eating beetle]
+
Sphaeroderus nitidicollis Guerin-Meneville 1829
Tr. Pamborini Hope 1838
+
G. Maoripamborus Brookes 1944
+
G. †Carabites Ponomarenko 1977
+
†Carabites caraboides Ponomarenko 1989
†Carabites nigriventris Ponomarenko 1986
G. †Carabites Heer 1852
+
†Carabites anthracinus Heer 1852
†Carabites arapahoensis Cockerell 1918
†Carabites capellinii Cecconi 1892
†Carabites creta Ponomarenko 1977
†Carabites cuneatus Cockerell 1920
†Carabites dubius Geinitz 1894
†Carabites ellipticus Haupt 1956
†Carabites eocenicus Cockerell 1921
†Carabites exanimus Scudder 1892
†Carabites exilis Heer 1862
†Carabites gardneri Cockerell 1920
†Carabites geinitzi Handlirsch 1906
†Carabites harringtoni Cockerell 1936
†Carabites islandicus Heer 1868
†Carabites kincaidi Cockerell 1908
†Carabites laesicollis Heer 1862
†Carabites parallelus Cockerell 1920
†Carabites peracutus Cockerell 1920
†Carabites rugosus Meunier 1898
†Carabites russelli Cockerell 1928
†Carabites schueli Cockerell 1926
†Carabites scoticus Cockerell 1921
†Carabites vitimensis Ponomarenko 1977
G. †Carabopteron Martynov 1926
+
†Carabopteron rugosum Ponomarenko 1985
G. †Cavicarabus Hong 1985
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†Cavicarabus lucensus Hong 1985
Unr. Celeripedes Latreille 1802
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Invalid names: Barbati Latreille 1802 [empty], Crepitantes Latreille 1802 [empty], Longipalpati Latreille 1802 [empty], Melanchlaeni Latreille 1802 [empty], Metallici Latreille 1802 [empty]
G. †Cicindelites Meunier 1898
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†Cicindelites armissanti Meunier 1898
Subfm. †Conjunctiinae Ponomarenko 1977
+
G. †Conjunctia Ponomarenko 1977
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†Conjunctia longa Zhang 1997
†Conjunctia prodroma Ponomarenko 1977
G. †Mesorabus Ponomarenko 1977
+
†Mesorabus elongatus Ponomarenko 1977
Subfm. Elaphrinae Latreille 1802
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G. Blethisa Bonelli 1810
+
Blethisa catenaria Brown 1944
Blethisa inexpectata Goulet and Smetana 1983
Blethisa multipunctata Linnaeus 1758
†Blethisa pleistocenica Lomnicki 1894
Invalid names: Helobium Leach 1815 [objective synonym]
G. Diacheila Motschulsky 1844
+
Diacheila arctica Gyllenhal 1810
†Diacheila matthewsi Böcher 1995
Diacheila polita Faldermann 1835
G. Elaphrus Fabricius 1775
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Elaphrus americanus Dejean 1831
Elaphrus angusticollis Sahlberg 1844
Elaphrus californicus Mannerheim 1843
Elaphrus cupreus Duftschmid 1812
Elaphrus finitimus Casey 1920
Elaphrus lapponicus Gyllenhal 1810
Elaphrus lecontei Crotch 1876
†Elaphrus pliocenicus Piton 1939
Elaphrus purpurans Hausen 1891
Elaphrus riparius Linnaeus 1758
Elaphrus sibiricus Motschulsky 1846
Invalid names: Elaphrus irregularis Scudder 1890 [nomen dubium], Elaphrus ruscarius foveatus Pierce 1948 [synonym]
G. †Elaphrotites Haupt 1956
+
†Elaphrotites densus Haupt 1956
G. †Fangshania Hong 1981
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†Fangshania punctata Hong 1981
G. †Feronites Frič 1884
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†Feronites velenovskyi Frič 1884
G. †Fonsecacarabus Martins-Neto and Mendes 2002
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†Fonsecacarabus placidus Martins-Neto and Mendes 2002
Subfm. Gehringiinae Darlington 1933
+
Tr. Gehringiini Darlington 1933
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Subtr. Gehringiina Darlington 1933
+
G. Gehringia Darlington 1933
G. †Glenopterus Heer 1847
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G. †Hadrocephalus Handlirsch 1906
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†Hadrocephalus anglicus Handlirsch 1906
†Hadrocephalus liasinus Handlirsch 1906
†Hadrocephalus minor Handlirsch 1906
Subfm. Harpalinae Bonelli 1810
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Tr. Abacetini Chaudoir 1873
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G. Loxandrus Leconte 1852
G. †Aethocarabus Ren 1995
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Tr. Chlaeniini Brullé 1834
+
Subtr. Chlaeniina Brullé 1834
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G. Chlaenius Bonelli 1810
+
Subg. Chlaenius (Agostenus) Fischer von Waldheim 1829
Subg. Chlaenius (Chlaeniellus) Reitter 1808
†Chlaenius electrinus Giebel 1862
†Chlaenius furvus Hong 1984
+
Invalid names: Shanwangicarabus paucilus Hong and Wang 1985 [synonym]
Chlaenius gebleri Ganglbauer 1891
†Chlaenius plicatipennis Wickham 1917
†Chlaenius primaevus Piton and Théobald 1939
†Chlaenius punctulatus Horn 1876
Invalid names: Shanwangicarabus Hong 1984 [synonym]
Tr. Galeritini Kirby 1825
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Subtr. Galeritina Kirby 1825
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G. Galerita Fabricius 1801
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Galerita bicolor Drury 1770
†Galerita marshii Scudder 1900
Tr. Harpalini Bonelli 1810
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Subtr. Anisodactylina Lacordaire 1854
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G. Anisodactylus Dejean 1829
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Invalid names: Eurytrichi Le Conte 1847 [synonym]
Subtr. Harpalina Bonelli 1810
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G. Cratacanthus Dejean 1829
G. Euryderus Le Conte 1846
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Invalid names: Nothopus Leconte 1852 [objective synonym]
G. Harpalus Latreille 1802
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†Harpalus abolitus von Heyden and von Heyden 1866
†Harpalus bruckmanni Heer 1862
†Harpalus conditus Scudder 1900
†Harpalus constrictus Heer 1862
†Harpalus deletus Oustalet 1874
†Harpalus dilatatus Theobald 1937
†Harpalus excavatus Förster 1891
†Harpalus lheritieri Piton 1940
†Harpalus maceratus Wickham 1911
†Harpalus nero Oustalet 1874
†Harpalus nitens Heer 1870
†Harpalus nuperus Scudder 1900
†Harpalus obtusus Förster 1891
†Harpalus oustaleti Meunier 1915
†Harpalus pleistocenicus Lomnicki 1894
Harpalus pusillus Motschulsky 1850
†Harpalus redivivus Wickham 1917
†Harpalus sinis Heer 1847
†Harpalus stierlini Heer 1862
†Harpalus stygius Heer 1862
†Harpalus tabidus Heer 1847
†Harpalus tardigradus Heer 1862
†Harpalus ulomaeformis Wickham 1917
†Harpalus veterum Cockerell 1921
Harpalus vittatus Gelber 1833
†Harpalus whitfieldii Scudder 1900
G. Ophonus Dejean 1821
+
Subg. Ophonus (Metophonus) Bedel 1897
G. Selenophorus Dejean 1829
Invalid names: Dapti Le Conte 1847 [synonym]
Subtr. Stenolophina Kirby 1837
+
G. Bradycellus Erichson 1837
+
Bradycellus ruficollis Stephens 1828
G. Dicheirotrichus Jacquelin du Val 1857
+
Subg. Dicheirotrichus (Trichocellus) Ganglbauer 1891
G. Stenolophus Dejean 1821
+
†Stenolophus religatus Scudder 1900
Tr. Helluonini Hope 1838
+
Subtr. Omphrina Jedlicka 1941
+
G. Creagris Nietner 1857
G. Helluomorpha De Laporte 1834
+
†Helluomorpha protogaea Giebel 1862
Tr. Lebiini Bonelli 1810
+
G. †Agatoides Motschulsky 1856
+
†Agatoides carinulatus Motschulsky 1856
Subtr. Calleidina Chaudoir 1873
+
G. Plochionus Latreille and Dejean 1821
+
†Plochionus lesquereuxi Scudder 1900
Subtr. Cymindidina Laporte 1834
+
G. Cymindis Latreille 1806
+
Subg. Cymindis (Tarulus) Bedel 1906
Cymindis angularis Gyllenhal 1810
Cymindis arctica Kryzhanovskij and Emets 1979
†Cymindis aurora Horn 1876
†Cymindis extorpescens Scudder 1898
†Cymindis pulchella Heer 1847
G. †Cymindoides Motschulsky 1856
+
†Cymindoides sculptipennis Motschulsky 1856
Subtr. Dromiusina Bonelli 1810
+
G. †Antephilorhizus Kirichenko-Babko et al. 2021
+
†Antephilorhizus pripiatiensis Kirichenko-Babko et al. 2021
†Antephilorhizus zerovae Kirichenko-Babko and Perkovsky 2022
G. Dromius Bonelli 1810
+
Dromius angusticollis Sahlberg 1880
†Dromius bakeri Abdullah 1969
†Dromius resinatus Germar 1813
Invalid names: Lebina Germar 1813 [synonym]
G. Microlestes Schmidt-Göbel 1846
+
G. Paradromius Fowler 1887
+
Paradromius linearis Olivier 1795
G. Syntomus Hope 1838
+
Syntomus americanus Dejean 1831
Syntomus truncatellus Linnaeus 1761
Subtr. Lebiina Bonelli 1810
+
G. Lebia Latreille 1802 [colorful foliage ground beetle]
+
†Lebia amissa von Heyden and von Heyden 1865
†Lebia harrelli Cockerell 1936
†Lebia minuscula Piton 1936
†Lebia newberryi Scudder 1900
†Lebia protospiloptera Cockerell 1921
G. †Miolebidia Zhang 1989
+
†Miolebidia gaura Zhang 1989
Subtr. Pericalina Hope 1838
+
G. Coptodera Dejean 1825
+
†Coptodera elektra Gamboa and Ortuño 2015
G. †Protoscalidion Schaufuss 1889
+
†Protoscalidion rugiae Schaufuss 1889
Invalid names: Brachinus repressus Scudder 1900 [nomen dubium]
Tr. Licinini Bonelli 1810
+
Subtr. Dicaelina Laporte 1834
+
G. Dicaelus Bonelli 1813
+
†Dicaelus alutaceus Horn 1876
G. Diplocheila Brullé 1834
+
†Diplocheila henshawi Scudder 1890
Invalid names: Rembus Latreille 1817 [replaced]
Subtr. Licinina Bonelli 1810
+
G. Badister Clairville 1806
+
†Badister antecursor Scudder 1900
†Badister debilis Heer 1847
†Badister fragilis Heer 1862
†Badister grandis Heer 1862
†Badister prodromus Heer 1847
Tr. Odacanthini Laporte 1834
+
Invalid names: Casnoniae LeConte 1861 [synonym]
Tr. Panagaeini Bonelli 1810
+
Subtr. Panagaeina Bonelli 1810
+
G. Panagaeus Latreille 1802
+
†Panagaeus dryadum Oustalet 1874
Tr. Pseudomorphini Hope 1838
+
G. †Burmapseudomorphus Beutel et al. 2020
+
†Burmapseudomorphus planus Beutel et al. 2020
Tr. Pterostichini Bonelli 1810
+
G. Cyclotrachelus Chaudoir 1838
+
Subg. Cyclotrachelus (Evarthrus) Leconte 1853
†Cyclotrachelus tenebricus Scudder 1900
Subtr. Pterostichina Bonelli 1810
+
G. Abax Bonelli 1810
+
†Abax durhamianus Lesne 1926
Abax parallelus Duftschmid 1812
G. Myas Dejean 1828
+
†Myas rigefactus Scudder 1900
†Myas umbrarum Scudder 1900
G. Platysma Bonelli 1810
G. Plocamostethus Britton 1940
+
G. Poecilus Bonelli 1810
+
Poecilus kugelanni Panzer 1797
Poecilus nearcticus Lindroth 1966
†Poecilus virgatus Pongrácz 1935
Invalid names: Feronia Latreille 1816 [synonym]
G. Pterostichus Bonelli 1810
+
Subg. Pterostichus (Argutor) Dejean 1821
Subg. Pterostichus (Bothriopterus) Chaudoir 1835
Subg. Pterostichus (Cryobius) Chaudoir 1838
Subg. Pterostichus (Eosteropus) Tschitschérine 1902
Subg. Pterostichus (Hypherpes) Chaudoir 1838
Subg. Pterostichus (Lamenius) Bousquet 1999
Subg. Pterostichus (Lenapterus) Berlov 1996
Subg. Pterostichus (Melanius) Bonelli 1810
+
Invalid names: Pterostichus (Omaseus) Dejean 1821 [synonym]
Subg. Pterostichus (Metallophilus) Chaudoir 1838
+
Invalid names: Pterostichus (Lyperopherus) Motschulsky 1844 [synonym]
Subg. Pterostichus (Phonias) des Gozis 1886
Subg. Pterostichus (Pseudomaseus) Chaudoir 1838
Subg. Pterostichus (Steropus) Dejean 1821
+
Invalid names: Pterostichus (Steroderus) Motschulsky 1850 [synonym]
Pterostichus abnormis Sahlberg 1880
†Pterostichus abrogatus Scudder 1890
Pterostichus cancellatus Motschulsky 1860
Pterostichus costatus Ménétriès 1851
†Pterostichus depletus Scudder 1900
†Pterostichus destitutus Scudder 1890
†Pterostichus destructus Scudder 1890
†Pterostichus distinctus Piton and Théobald 1939
†Pterostichus dormitans Scudder 1890
†Pterostichus fernquisti Wickham 1931
†Pterostichus fractus Scudder 1890
†Pterostichus gelidus Scudder 1877
†Pterostichus gurnetensis Cockerell 1921
†Pterostichus laevigatus Scudder 1890
Pterostichus minor Gyllenhal 1827
†Pterostichus molestus Theobald 1937
Pterostichus niger Schaller 1783
Pterostichus nigrita Paykull 1790
Pterostichus parens Tschitscherine 1897
†Pterostichus pumpellyi Scudder 1900
Pterostichus sublaevis Sahlberg 1880
Pterostichus tundrae Tschitscherine 1894
†Pterostichus vetustus Heer 1862
†Pterostichus walcotti Scudder 1900
G. Stomis Clairville 1806
+
†Stomis elegans Oustalet 1874
Invalid names: Feroniens Dejean 1825 [synonym]
G. †Sinis Heer 1862
+
†Sinis brevicollis Heer 1862
Tr. Sphodrini Laporte 1834
+
Subtr. Calathina Laporte 1834
+
G. Calathus Bonelli 1810
+
Subg. Calathus (Neocalathus) Ball and Negre 1972
Calathus fuscipes Goeze 1777
Calathus piceus Marsham 1802
G. †Quasicalathus Schmidt and Will 2022
+
†Quasicalathus agonicollis Schmidt and Will 2022
†Quasicalathus conservans Schmidt and Will 2022
†Quasicalathus elpis Ortuño and Arillo 2009
Subtr. Sphodrina Laporte 1834
+
G. Pristonychus Dejean 1828
G. Sphodrus Clairville 1806
Subtr. Synuchina Lindroth 1956
+
G. Synuchus Gyllenhal 1810
Tr. Zabrini Bonelli 1810
+
Subtr. Amarina Zimmerman 1832
+
G. Amara Bonelli 1810
+
Subg. Amara (Amara) Bonelli 1810
Subg. Amara (Bradytus) Stephens 1827
Subg. Amara (Celia) Zimmerman 1832
Subg. Amara (Curtonotus) Stephens 1827
Subg. Amara (Paracelia) Bedel 1899
Amara aenea De Geer 1774
Amara arcticola Poppius 1906
Amara aulica Panzer 1796
+
Invalid names: Carabus ruficornis De Geer 1774 [replaced]
Amara bifrons Gyllenhal 1810
Amara brunnea Gyllenhal 1810
†Amara cockerelli Wickham 1912
Amara conflata Le Conte 1855
†Amara danae Scudder 1900
Amara erratica Duftschmid 1812
†Amara foersteri Handlirsch 1907
+
Invalid names: Amara sinuata Förster 1891 [replaced]
Amara insignis Dejean 1831
Amara municipalis Duftschmid 1812
†Amara occlusa Scudder 1900
†Amara paleomelas Scudder 1879
†Amara pinguicula Heer 1862
†Amara powellii Scudder 1900
†Amara primigenia Heer 1862
†Amara princeps Heer 1862
†Amara procera Förster 1891
Amara pseudobrunnea Lindroth 1968
†Amara revocata Scudder 1900
†Amara sterilis Scudder 1900
†Amara veterata Scudder 1900
Subtr. Zabrina Bonelli 1810
+
G. Zabrus Clairville 1806
+
Tr. Zuphiini Bonelli 1810
+
Subtr. Zuphiina Bonelli 1810
+
G. Polistichus Bonelli 1810
+
†Polystichus hopei Oustalet 1874
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]
G. †Harpalomimes Handlirsch 1906
+
†Harpalomimes burmeisteri Giebel 1856
G. †Hebeicarabus Hong 1984
+
G. †Lithocantharis Lin 1997
+
Subfm. Loricerinae Bonelli 1810
+
Tr. Loricerini Bonelli 1810
+
G. †Cretoloricera Liu et al. 2023
+
†Cretoloricera electra Liu et al. 2023
G. Loricera Latreille 1802
+
Loricera decempunctata Eschscholtz 1833
†Loricera electrica Klausnitzer 2003
†Loricera groehni Cai et al. 2017
Loricera pilicornis Fabricius 1775
Invalid names: Loriceritae Bonelli 1810 [empty]
G. †Meileyingia Hong 1987
+
†Meileyingia spinosa Hong 1987
G. †Metagrilium Handlirsch 1906
+
†Metagrilium westwoodi Handlirsch 1906
Subfm. Migadopinae Chaudoir 1861
+
G. †Cretomigadops Liu et al. 2022
+
†Cretomigadops bidentatus Liu et al. 2022
Tr. Migadopini Chaudoir 1861
+
Subtr. Migadopina Chaudoir 1861
+
G. Migadops Waterhouse 1842
+
Migadops latus Guerin-Meneville 1841
Subfm. Nebriinae Laporte 1834
+
G. †Ledouxnebria Deuve 1998
+
†Ledouxnebria brisaci Deuve 1998
Tr. Nebriini Laporte 1834
+
G. †Archaeonebria Kavanaugh and Schmidt 2019
+
†Archaeonebria inexspectata Schmidt and Kavanaugh 2019
G. Leistus Frölich 1799
G. Nebria Latreille 1802
+
Nebria brevicollis Fabricius 1792
†Nebria fossilis Piton 1935
Nebria frigida Sahlberg 1844
Nebria metallica Fischer von Waldheim 1822
Nebria nivalis Paykull 1790
†Nebria pluto Heer 1862
Nebria rufescens Strøm 1768
+
Invalid names: Nebria gyllenhali Schönherr 1806 [synonym]
Nebria sahlbergi Fischer von Waldheim 1828
†Nebria tisiphone Oustalet 1874
Tr. Notiophilini Motschulsky 1850
+
G. Notiophilus Duméril 1805
+
Notiophilus aeneus Herbst 1806
Notiophilus aquaticus Linnaeus 1758
Notiophilus biguttatus Fabricius 1779
Notiophilus palustris Duftschmid 1812
†Notiophilus vancouveri Pierce 1950
Tr. Opisthiini Dupuis 1912
+
G. Opisthius Kirby 1837
+
Tr. Pelophilini Kavanaugh 1996
+
G. Pelophila Dejean 1821
+
Pelophila borealis Paykull 1790
G. †Nebrorabus Ponomarenko 1989
+
†Nebrorabus baculum Ponomarenko 1989
†Nebrorabus capitatus Ponomarenko 1989
†Nebrorabus nebrioides Ponomarenko 1989
G. †Neothanes Scudder 1890
+
†Neothanes europaeus Ponomarenko 2019
†Neothanes testeus Scudder 1878
G. †Notokistus Handlirsch 1906
+
†Notokistus brodiei Handlirsch 1906
Subfm. Omophroninae Bonelli 1810
+
G. †Cretomophron Rosová et al. 2023
+
†Cretomophron mutilum Rosová et al. 2023
Subfm. Paussinae Latreille 1806 [ant nest beetle]
+
G. †Cerapterites Wasmann 1926
+
†Cerapterites primaevus Wasmann 1926
Tr. Paussini Latreille 1806
+
Subtr. †Arthropteritina Luna de Carvalho 1961
+
G. †Arthropterites Wasmann 1926
+
†Arthropterites klebsi Wasmann 1926
Subtr. Carabidomemnina Wasmann 1928
+
G. Eohomopterus Wasmann 1919
+
†Eohomopterus poinari Nagel 1997
†Eohomopterus simojovelensis Solórzano Kraemer 2006
Subtr. Cerapterina Billberg 1820
+
G. Arthropterus M'Leay 1838
+
†Arthropterus hermenaui Wasmann 1926
G. †Protocerapterus Wasmann 1926
+
†Protocerapterus incola Wasmann 1929
Subtr. †Eopaussina Luna de Carvalho 1951
+
G. †Eopaussus Wasmann 1926
+
†Eopaussus balticus Wasmann 1926
Subtr. Paussina Latreille 1806
+
G. Homopterus Westwood 1841
+
G. Paussus Linnaeus 1775
G. Pleuarthropterus Wasmann 1929
+
G. †Paussoides Motschulsky 1856
+
†Paussoides mengei Motschulsky 1856
G. †Paussopsis Cockerell 1911
+
†Paussopsis nearctica Cockerell 1911
G. †Succinarthropterus Kolbe 1926
+
†Succinarthropterus helmi Schaufuss 1896
†Succinarthropterus kolbei Wasmann 1926
Invalid names: Arthropterillus Wasmann 1926 [synonym]
Invalid names: Ozaenidae Hope 1838 [synonym], Ozaenini [empty]
Subfm. Platyninae Bonelli 1810
+
Tr. Platynini Bonelli 1810
+
G. Agonum Bonelli 1810
+
Subg. Agonum (Olisares) Motschulsky 1865
+
†Agonum abstractum Scudder 1900
†Agonum antiquum Piton 1940
Agonum micans Nicolai 1822
Agonum muelleri Herbst 1784
Agonum quinquepuncatum Motschulsky 1844
Agonum sahlbergi Chaudoir 1850
Agonum sexpunctatum Linnaeus 1758
Agonum versutum Sturm 1824
G. Anchomenus Bonelli 1810
+
†Anchomenus bipunctatus Förster 1891
†Anchomenus orphanus Heer 1847
G. Blackburnia Sharp 1878
+
Subg. Blackburnia (Blackburnia) Sharp 1878
+
†Blackburnia (Blackburnia) burneyi Liebherr and Porch 2015
†Blackburnia (Blackburnia) cryptipes Liebherr and Porch 2015
†Blackburnia (Blackburnia) godzilla Liebherr and Porch 2015
†Blackburnia (Blackburnia) menehune Liebherr and Porch 2015
†Blackburnia (Blackburnia) mothra Liebherr and Porch 2015
†Blackburnia (Blackburnia) ovata Liebherr and Porch 2015
†Blackburnia (Blackburnia) rugosa Liebherr and Porch 2015
Invalid names: Disenochus Blackburn 1878 [synonym]
G. Oxypselaphus Chaudoir 1843
+
G. Paranchus Lindroth 1974
+
Paranchus albipes Fabricius 1794
Subtr. Platynina Bonelli 1810
+
G. Limodromus Motschulsky 1850
+
†Limodromus emetikos Gamboa and Ortuño 2017
†Limodromus hoffeinsorum Schmidt 2015
G. Platynus Bonelli 1810
+
Subg. Platynus (Batenus) Motschulsky 1865
+
†Platynus caesus Scudder 1890
†Platynus calvini Wickham 1917
†Platynus casus Scudder 1890
†Platynus desuetus Scudder 1890
†Platynus dilapidatus Scudder 1895
†Platynus dissipatus Scudder 1890
†Platynus exterminatus Scudder 1900
†Platynus florissantensis Wickham 1913
Platynus funebris LeConte 1854
†Platynus halli Scudder 1890
†Platynus harttii Scudder 1890
†Platynus hindei Scudder 1890
†Platynus insculptipennis Wickham 1917
†Platynus interglacialis Scudder 1900
†Platynus interitus Scudder 1900
†Platynus longaevus Scudder 1900
†Platynus pleistocenicus Wickham 1917
†Platynus senex Scudder 1878
†Platynus subgelidus Wickham 1917
†Platynus tartareus Scudder 1900
G. †Praeanchodemus Schmidt et al. 2017
+
†Praeanchodemus punctaticeps Schmidt et al. 2017
G. †Procarabites Handlirsch 1906
Subfm. Protopaussinae Gestro 1892
+
G. Protopaussus Gestro 1892
+
†Protopaussus pristinus Nagel 1997
Subfm. †Protorabinae Ponomarenko 1977
+
G. †Cordorabus Ponomarenko 1977
+
†Cordorabus antennatus Ponomarenko 1977
†Cordorabus minimus Ponomarenko 1977
†Cordorabus notatus Ponomarenko 1977
†Cordorabus striatus Ponomarenko 1986
†Cordorabus vittatus Ponomarenko 1980
†Cordorabus wuqiensis Hong 1995
G. †Cretorabus Ponomarenko 1977
+
†Cretarabus medius Hong and Wang 1990
†Cretorabus capitatus Ponomarenko 1977
†Cretorabus latus Ponomarenko 1977
†Cretorabus orientalis Ponomarenko 1989
†Cretorabus ovalis Ponomarenko 1989
†Cretorabus rasnitsyni Wang and Zhang 2011
†Cretorabus sulcatus Ponomarenko et al. 2005
G. †Lirabus Hong 1992
+
†Lirabus granulatus Hong 1992
G. †Lithorabus Ponomarenko 1977
+
†Lithorabus incertus Ponomarenko 1977
G. †Magnirabus Hong and Wang 1990
+
†Magnirabus furvus Hong and Wang 1990
G. †Ovrabites Ponomarenko 1977
+
†Ovrabites incertus Ponomarenko 1993
†Ovrabites jurassicus Ponomarenko 1977
†Ovrabites ovalis Ponomarenko 1977
G. †Penecupes Ren 1995
+
†Penecupes rapax Ren 1995
G. †Protorabus Ponomarenko 1977
+
†Protorabus crassus Ponomarenko 1989
†Protorabus elongatus Ponomarenko 1986
†Protorabus kobdoensis Ponomarenko 1986
†Protorabus magnus Ponomarenko 1977
†Protorabus minisculus Zhang 1997
†Protorabus nigrimonticola Ponomarenko 1977
†Protorabus planus Ponomarenko 1977
†Protorabus tsagensis Ponomarenko 1989
G. †Rhopalochlaenius Zhang et al. 1994
+
†Rhopalochlaenius phalanthus Zhang et al. 1994
Subfm. Scaritinae Bonelli 1810
+
Tr. Clivinini Rafinesque 1815
+
G. Camptodontus Dejean 1826
Subtr. Clivinina Rafinesque 1815
+
G. Clivina Latreille 1802
Tr. Dyschiriini Kolbe 1880
+
G. Dyschiriodes Jeannel 1941
+
Dyschiriodes aeneus Dejean 1825
Dyschiriodes luticola Chaudoir 1850
Dyschiriodes nigricornis Motschulsky 1844
+
Invalid names: Dyschirius septentrionum Munster 1923 [synonym]
G. Dyschirius Bonelli 1810
+
Dyschirius montanus Le Conte 1879
+
Invalid names: Dyschirius arenosus Stephens 1827 [synonym]
Invalid names: Helophorus rigescens Scudder 1890 [nomen dubium]
G. †Dyschiriomimus Yablokov-Khnzorian 1960
+
†Dyschiriomimus stackelbergi Yablokov-Khnzorian 1960
Tr. †Palaeoaxinidiini McKay 1991
+
G. †Palaeoaxinidium McKay 1991
+
Tr. Scaritini Bonelli 1810
+
Subtr. Scaritina Bonelli 1810
+
G. Scarites Fabricius 1775
+
†Scarites mancus Zhang 1989
†Scarites robustiventris Theobald 1937
Invalid names: Scarites haidingeri Heer 1861 [nomen nudum]
Invalid names: Promecognathinae Le Conte 1853 [synonym]
G. †Sinocarabus Hong 1982
+
G. Stereocerus Kirby 1837
+
Stereocerus rubripes Motschulsky 1860
G. †Sunocarabus Hong 1987
+
†Sunocarabus brunneus Hong 1987
Subfm. Trechinae Bonelli 1810
+
Tr. Patrobini Kirby 1837
+
Subtr. Patrobina Kirby 1837
+
G. Patrobus Dejean 1821
+
Patrobus assimilis Chaudoir 1844
Patrobus fossifrons Eschscholtz 1823
Patrobus foveocollis Eschscholtz 1823
†Patrobus gasiorowskii Lomnicki 1894
†Patrobus henshawi Wickham 1917
†Patrobus kudiensis Cockerell 1925
Patrobus stygicus Chaudoir 1872
+
Invalid names: Patrobus decessus Scudder 1900 [synonym], Patrobus frigidus Scudder 1900 [synonym], Patrobus gelatus Scudder 1890 [synonym]
G. †Trechinites Heer 1862
+
†Trechinites oblongus Heer 1862
Unr. Trechitae Bonelli 1810
+
G. †Balticeler Schmidt and Maddison 2021
+
†Balticeler kerneggeri Schmidt and Maddison 2021
Tr. Bembidiini Stephens 1827
+
G. Asaphidion Des Gozis 1886
+
Asaphidion alaskanum Wickham 1919
Asaphidion yukonense Wickham 1919
G. Bembidion Latreille 1802
+
Subg. Bembidion (Asioperyphus) Vysoky 1986
Subg. Bembidion (Bembidion) Latreille 1802
Subg. Bembidion (Bracteon) Bedel 1879
+
Invalid names: Bembidion (Chrysobracteon) Netolitzky 1914 [synonym]
Subg. Bembidion (Diplocampa) Bedel 1896
Subg. †Bembidion (Eodontium) Schmidt and Michalik 2017
Subg. Bembidion (Eupetedromus) Netolitzky 1911
Subg. Bembidion (Furcacampa) Netolitzky 1931
Subg. Bembidion (Hirmoplataphus) Lindroth 1963
Subg. Bembidion (Hydrium) LeConte 1847
+
Invalid names: Bembidion (Eurytrachelus) Motschulsky 1850 [synonym]
Subg. Bembidion (Melomalus) Casey 1918
Subg. Bembidion (Metallina) Motschulsky 1850
Subg. Bembidion (Neobembidion) Bousquet 2006
Subg. Bembidion (Nesocidium) Sharp 1903
Subg. Bembidion (Notaphus) Dejean 1821
Subg. Bembidion (Ochthedromus) LeConte 1847
Subg. Bembidion (Peryphanes) Jeannel 1941
Subg. Bembidion (Peryphus) Dejean 1821
Subg. Bembidion (Philochthemphanes) Netolitzky 1943
+
Invalid names: Bembidion (Archaeophilochthus) Ortuño and Arillo 2010 [synonym]
Subg. Bembidion (Philochthus) Stephens 1828
Subg. Bembidion (Phyla) Motschulsky 1844
Subg. Bembidion (Plataphus) Motschulsky 1864
+
Invalid names: Bembidion (Plataphodes) Ganglbauer 1891 [synonym]
Subg. Bembidion (Princidium) Motschulsky 1864
Subg. Bembidion (Semicampa) Netolitzky 1910
Subg. Bembidion (Terminophanes) Müller-Motzfeld 1998
Subg. Bembidion (Testedium) Motschulsky 1864
Subg. Bembidion (Trepanedoris) Netolitzky 1918
Subg. Bembidion (Trepanes) Motschulsky 1864
Bembidion aeneum Germar 1824
†Bembidion damnosum Scudder 1900
†Bembidion everestae Pierce 1944
†Bembidion festivum Zhang et al. 1994
Bembidion gilvipes Sturm 1825
Bembidion lunatum Duftschmid 1812
Bembidion minimum Fabricius 1792
Bembidion obliquum Sturm 1825
†Bembidion strenum Zhang 1989
Bembidion virens Gyllenhal 1827
G. Ocys Stephens 1828
+
Ocys harpaloides Audinet-Serville 1821
G. Sinechostictus Motschulsky 1864
+
Subg. Sinechostictus (Sinechostictus) Motschulsky 1864
Invalid names: Bembidiina Stephens 1827 [invalid subgroup], Tachyina Motschulsky 1862 [empty]
Tr. Pogonini Laporte 1834
+
G. Pogonus Dejean 1821
+
Subg. Pogonus (Pogonoidius) Carret 1903
Tr. Tachyini Motschulsky 1862
+
Subtr. Tachyina Motschulsky 1862
+
G. Elaphropus Motschulsky 1839
G. Polyderis Motschulsky 1862
G. Tachys Dejean 1821
Tr. Trechini Bonelli 1810
+
G. †Antarctotrechus Ashworth and Erwin 2016
+
†Antarctotrechus balli Ashworth and Erwin 2016
G. Trechiama Jeannel 1927
Subtr. Trechina Bonelli 1810
+
G. Trechoblemus Ganglbauer 1891
G. Trechus Clairville 1806
Invalid names: Bembidion succini Giebel 1856 [nomen dubium], Patrobinae Kirby 1837 [synonym]
G. †Trechoides Motschulsky 1856
+
†Trechoides fasciatus Motschulsky 1856
G. †Tremembecarabus Martins-Neto 1998
+
†Tremembecarabus rotundus Martins-Neto 1998
G. †Wongyekokia Lin 1997
+
G. †Wuchangicarabus Hong 1985
+
†Wuchangicarabus latus Hong 1985
G. †Yunnanocarabus Lin 1977
+
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Hettangian to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma

Collections (412 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Rhaetian - Early/Lower Hettangian208.5 - 196.5United Kingdom (England) Coleoptera indet. 3, Coleoptera indet. 6, Coleoptera indet. 5, Coleoptera indet. 4 (124688)
Angulata201.3 - 196.5Switzerland Carabites anthracinus (131766)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Lithorabus incertus (105087)
Obtusum196.5 - 189.6United Kingdom (England) Carabidae indet. (114637)
Early/Lower Toarcian183.0 - 182.0Germany Bellingeria laticollis, Elaterium triopas (123987)
Toarcian182.7 - 174.1China (Hubei) Wuchangicarabus latus (156563) Wuchangicarabus latus, Cavicarabus lucensus (156562)
Toarcian - Aalenian182.7 - 170.3Russian Federation (Kemerovo) Carabopteron rugosum (125471)
Bajocian170.3 - 168.3China (Yunnan) Yunnanocarabus litus (165273)
Bajocian - Bathonian170.3 - 166.1China (Anhui) Carabidae indet. (106695)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Carabopteron punctatolineatum (115792) Cordorabus minimus (106101) Cordorabus notatus, Cordorabus antennatus, Protorabus planus, Protorabus nigrimonticola (105971) Cordorabus notatus, Ovrabites ovalis, Protorabus planus, Mesorabus elongatus (113439) Cordorabus notatus, Protorabus nigrimonticola, Antemnacrassa kazakhstanica, Antemnacrassa nigrimontana, Antemnacrassa albosulcata (105426) Ovrabites jurassicus (115717) Ovrabites ovalis, Antemnacrassa albosulcata (106102) Ovrabites ovalis, Ovrabites jurassicus, Protorabus planus, Holcorobeus nigrimontanus (106279) Ovrabites ovalis, Protorabus magnus (105970)
Tithonian152.1 - 145.0Mongolia Protorabinae indet. (152133)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Carabidium dejeanianum, Carabidae indet. (105966) Cretorabus sulcatus (123443)
Berriasian145.0 - 139.8United Kingdom (England) Coleoptera indet. (127481)
Berriasian - Valanginian145.0 - 132.9Hong Kong Lithocantharis lunglokshuiensis (173064) Wongyekokia angustris (173068 173072)
Berriasian - Valanginian145.0 - 132.9Russian Federation (Chita) Nebrorabus capitatus (125410)
Valanginian139.8 - 132.9China (Shaanxi) Cordorabus wuqiensis (140459)
Valanginian - Barremian139.8 - 125.0Mongolia (Dornogov) Cretorabus orientalis (113407)
Late/Upper Hauterivian - Aptian136.4 - 113.0Japan Harpalinae indet. (153093)
Hauterivian132.9 - 129.4Russian Federation (Chita) Protorabus crassus (125409)
Early/Lower Barremian130.0 - 125.45Spain Antemnacrassa monreali (152685)