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Leiostegiina

Trilobita - Corynexochida

Taxonomy
Leiostegiina was named by Bradley (1925). It is not extant.

It was assigned to Corynexochida by Adrain (2011) and Wernette et al. (2020).

Synonymy list
YearName and author
1925Leiostegiina Bradley
2011Leiostegiina Adrain p. 105
2020Leiostegiina Wernette et al. p. 874

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
RankNameAuthor
phylumArthropodaLatreille 1829
subphylumArtiopoda(Hou and Bergstrom)
classTrilobitaWalch 1771
orderCorynexochida
suborderLeiostegiinaBradley 1925
suborderLeiostegiinaBradley 1925

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. †Leiostegiina Bradley 1925
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Fm. †Illaenuridae Vodges 1890
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G. †Ambonolium Raymond 1924
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†Ambonolium affine Rasetti 1944
G. †Lecanopyge Raymond 1937
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†Lecanopyge angustata Rasetti 1944
†Lecanopyge arenaria Rasetti 1944
†Lecanopyge billingsi Clark 1924
†Lecanopyge expansa Raymond 1937
†Lecanopyge intermedia Rasetti 1944
†Lecanopyge prolifica Rasetti 1944
G. †Minicephalus Lochmann and Lu 1960
G. †Olenekella Kharmova 1985
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†Olenekella alimbata Kharmova and Pegel 1985
G. †Platydiamesus Raymond 1937
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†Platydiamesus levisensis Rasetti 1944
G. †Polyariella Rozova 1963
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†Polyariella pallida Rozova 1963
G. †Rasettaspis Ivshin 1962
G. †Rasettia Lochman 1953
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†Rasettia crucensis Rusconi 1953
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Invalid names: Platycolpus excavatus Rusconi 1953 [synonym]
†Rasettia magna Bell and Ellinwood 1962
Invalid names: Platycolpus Raymond 1913 [replaced]
G. †Resseraspis Rasetti 1945
G. †Tatonaspis Kobayashi 1935
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†Tatonaspis levisensis Rasetti 1944
G. †Yurakia Rozova 1963
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†Yurakia yurakiensis Rozova 1963
Invalid names: Lecanopygidae Lochman 1953 [invalid subgroup]
Superfm. †Leiostegioidea Bradley 1925
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Fm. †Kaolishaniidae Kobayashi 1935
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Subfm. †Ceronocarinae Wernette et al. 2020
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G. †Ceronocare Shergold 1975
G. †Guangxiaspis Zhou 1977
G. †Hapsidocare Shergold 1975
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†Hapsidocare lilyensis Shergold 1980
G. †Mansuyites Shergold 1971
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†Mansuyites futiliformis Shergold 1971
Invalid names: Anhuiaspis Qian and Qiu 1983 [synonym]
G. †Taipaikia Kobayashi 1960
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†Taipaikia glabra Endo and Resser 1937
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Invalid names: Taipaikia yanshanensis Duan et al. 2005 [synonym]
G. †Eokaolishania Wittke 1984
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†Eokaolishania angustilimbata Zhu and Wittke 1989
G. †Hemikaolishania Zhang 1983
G. †Kaolishania Sun 1924
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†Kaolishania australis Shergold 1971
G. †Liaotropis Schrank 1975
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†Liaotropis tropis Schrank 1975
Subfm. †Mansuyiinae Hupe 1955
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G. †Kaolishaniella Sun 1935
G. †Mansuyia Sun 1924
G. †Paramansuyella Endo 1937
G. †Pagodia Walcott 1905
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Subg. †Pagodia (Idamea) Whitehouse 1939
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Subg. †Pagodia (Lotosoides) Shergold 1975
†Pagodia bia Walcott 1905
†Pagodia depressa Walcott 1905
†Pagodia lotos Walcott 1905
†Pagodia major Lu and Zhu 1980
†Pagodia seelyi Walcott 1912
G. †Palemansuyia Yuan and Yin 1998
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†Palemansuyia prima Yuan and Yin 1998
G. †Peichiashania Chang 1959
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†Peichiashania prima Shergold 1980
†Peichiashania quarta Shergold 1980
†Peichiashania secunda Shergold 1980
†Peichiashania tertia Shergold 1980
G. †Tangjiaella Qiu 1984
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†Tangjiaella bilira Qiu 1984
Subfm. †Tingocephalinae Hupe 1955
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G. †Palacorona Shergold 1971
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†Palacorona bacculata Shergold 1971
G. †Tingocephalus Sun 1935
Fm. †Leiostegiidae Bradley 1925
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G. †Aedotes Opik 1967
G. †Agerina Tjernvik 1956
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†Agerina acutilimbata Ghobadi Pour et al. 2011
†Agerina clymene Mergl 2006
†Agerina elongata Lu 1975
†Agerina erratica Tjernvik 1956
†Agerina ferrigena Ruzicka 1926
†Agerina parva Xiang and Zhou 1987
†Agerina quadrata Dean 1966
Invalid names: Hexianella Zhang 1983 [synonym], Plesiyuepingia Xiang and Zhou 1987 [synonym]
G. †Annamitella Mansuy 1920
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†Annamitella borealis Neuman 1964
†Annamitella brachyops Fortey and Shergold 1984
†Annamitella granulatus Weber 1948
†Annamitella insulana Dean 1973
†Annamitella kantsingensis Chang and Fan 1960
†Annamitella parva Hsu 1948
†Annamitella perplexa Bates 1968
†Annamitella strigifrons Fortey and Shergold 1984
†Annamitella tellecheai Rusconi 1951
†Annamitella yichangensis Xiang and Zhou 1983
Invalid names: Bathyuriscops Keller and Lisogor 1954 [synonym], Endoaspis Lochman 1956 [synonym], Monella Bates 1968 [synonym]
G. †Arcifimbria Rasetti 1963
G. †Brackebuschia Harrington and Leanza 1957
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†Brackebuschia acheila Harrington and Leanza 1957
Invalid names: Bodenbenderia Harrington and Leanza 1957 [synonym]
G. †Chiawangella Chu 1959
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†Chiawangella hunanensis Peng et al. 2004
†Chiawangella pacifica Walcott 1911
G. †Cholopilus Raymond 1924
G. †Chosenia Kobayashi 1934
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†Chosenia divergens Lu 1975
†Chosenia laticephala Kobayashi 1934
G. †Chuangiella Kobayashi 1935
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†Chuangiella elongata Kobayashi 1935
†Chuangiella shengwanensis Yang et al. 1993
G. †Chuangioides Chu 1959
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G. †Chuangiopsis Kobayashi 1955
G. †Eochuangia kobayashi 1935
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†Eochuangia hana Kobayashi 1935
G. †Geminiclavula Peng et al. 2004
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†Geminiclavula wangcunica Peng et al. 2004
G. †Iranaspis King 1937
G. †Kepisis Hu 1971
G. †Leiostegium Raymond 1913
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Subg. †Leiostegium (Euleiostegium) Zhou and Zhan 1978
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Subg. †Leiostegium (Perischodory) Raymond 1937
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†Perischodory incompertum Berg and Ross 1959
†Leiostegium manitouensis Walcott 1925
†Leiostegium proprium Boyce 1989
†Leiostegium quadratus Billings 1860
†Leiostegium spongiosum Fortey and Bruton 2013
†Leiostegium tyboensis Ross 1970
Invalid names: Alloleiostegium Zhou and Zhang 1978 [synonym]
G. †Linguchuangia Zhang 1983
G. †Lloydia Vogdes 1893
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Subg. †Lloydia (Evansaspis) Kobayashi 1955
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Subg. †Lloydia (Leiostegium) Raymond 1913
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†Lloydia (Leiostegium) douglasi Harrington 1937
†Lloydia obsoletus Phleger, Jr. 1933
G. †Meropalla Opik 1967
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†Meropalla bella Yuan and Yin 1998
†Meropalla gibbera Peng et al. 2004
Subfm. †Pagodiinae Kobayashi 1935
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G. †Lotosoides Shergold 1975
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†Lotosoides bathyora Shergold 1980
G. †Plethopeltella Kobayshi 1944
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†Plethopeltella resseri Kobayashi 1933
G. †Prochuangia Kobayashi 1935
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†Prochuangia glabella Shergold 1980
†Prochuangia leiocephala Peng et al. 1999
†Prochuangia longispina Lu and Zhu 1980
†Prochuangia turcica Dean 2006
G. †Pseudocalymene Pillet 1974
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Invalid names: Eucalymene Lu 1975 [synonym], Paraszechuanella Liu 1977 [synonym], Szechuanella Lu 1962 [synonym]
G. †Reubenella Lochmann 1966
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†Reubenella rex Lochman 1966
G. †Schantungia Lorenz 1906
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Invalid names: Chuangia Walcott 1911 [synonym]
G. †Tinaspis Robison and Pantoja-Alor 1968
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†Tinaspis totoyaca Robison and Pantoja-Alor 1968
G. †Wanshania Rong and Yang 1977
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†Wanshania stenos Yang et al. 1993
†Wanshania wanshanensis Rong and Yang 1977
Invalid names: Pagodiidae Kobayashi 1935 [invalid subgroup]
Fm. †Missisquoiidae Hupé 1953
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G. †Hardyia Walcott 1924
G. †Lunacrania Ivshin 1962
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†Lunacrania nasuta Winston and Nicholls 1967
†Lunacrania spicata Fortey 1983
†Lunacrania trisecta Kobayashi 1955
Invalid names: Paranumia [synonym]
G. †Parakoldinioidia Endo 1937
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†Parakoldinioidia akerfeldti Westrop and Adrain 2014
†Parakoldinioidia akessoni Westrop and Adrain 2014
†Parakoldinioidia inflata Winston and Nicholls 1967
†Parakoldinioidia lindgreni Westrop and Adrain 2014
†Parakoldinioidia lopezi Westrop and Adrain 2014
†Parakoldinioidia mendezi Westrop and Adrain 2014
†Parakoldinioidia stitti Fortey 1983
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Invalid names: Missisquoia typicalis Shaw 1951 [replaced]
Invalid names: Missisquoia Shaw 1951 [synonym]
G. †Pseudokoldinioidia Endo 1944
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G. †Tasmanocephalus kobayashi 1936
Fm. †Shirakiellidae Kobayashi 1935
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G. †Neoshirakiella Sun and Xiang 1979
G. †Shirakiella Kobayashi 1935
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†Shirakiella elongata Kobayashi 1935
G. †Warburtonella Sun and Jago 2009
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†Warburtonella coongiensis Sun and Jago 2009
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: actively mobileo
Life habit: nektobenthico
Diet: carnivoreo
Created: 2009-09-21 20:17:08
Modified: 2012-08-27 07:15:31
Source: o = order, c = class
References: Aberhan et al. 2004, Hendy et al. 2009

Age range: base of the Taoyuanian to the top of the Caradoc or 498.50000 to 449.50000 Ma

Collections (577 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Cambrian538.8 - 485.4China (Jilin Provence) Leiostegium floodi (156930)
Marjumian504.5 - 497.0USA (Wisconsin) Pagodia sp. (230)
Wangcunian504.5 - 497.0China (Hunan) Geminiclavula wangcunica (49274)
Mayan - Merioneth504.5 - 485.4Iran (Semnan) Chuangia sp., Kaolishania sp., Prochuangia sp. (22823)
Guzhangian500.5 - 497.0China (Nei Mongol) Chuangioides subaiyingouensis (129775 130254)
Guzhangian500.5 - 497.0China (Liaoning) Chiawangella pacifica (129879)
Guzhangian500.5 - 497.0China (Shanxi) Chuangioides punctatus (129927) Shirakiellidae indet. (129812)
Guzhangian500.5 - 497.0China (Henan) Mansuyia sp. (127078) Wanshania stenos (127077)
Guzhangian500.5 - 497.0China (Hunan) Chiawangella hunanensis (107050 126439) Chiawangella hunanensis, Wanshania wanshanensis (49182) Meropalla bella (126348 126440) Meropalla bella, Wanshania wanshanensis (49181) Meropalla gibbera (126325) Wanshania constricta (128930) Wanshania wanshanensis (49174 49176 49177 49178 49179 49180 49183 49303 126312 126314 126322 126328)
Guzhangian500.5 - 497.0China (Guizhou) Meropalla bella, Palemansuyia prima (125072)

Youngest occurrences
Time interval Ma Country or state PBDB collection number
Llanvirn468.0 - 458.4China (Gansu) Bathyuriscops sp. (23594)
Llanvirn468.0 - 458.4China (Xinjiang) Pseudocalymene quadrata (170841 170842)
Llanvirn468.0 - 458.4Canada (Newfoundland and Labrador) Lloydia sp. (78097)
Llanvirn468.0 - 458.4Argentina (La Rioja) Proetiella tellecheai (116146)
Darriwilian467.3 - 458.4Argentina (San Juan) Proetiella tellecheai (116105 116106 116108 116111 116114 116115)
Darriwilian467.3 - 458.4China (Sichuan) Pseudocalymene sp. (64235)
Darriwilian467.3 - 458.4Australia (Tasmania) Tasmanocephalus sp. (2674)
Llandeilo461.1 - 458.4China (Guizhou) Annamitella sp. (188421 188422 188425)
Llandeilo - Caradoc461.1 - 449.5Canada (Northwest Territories) Agerina norrisi (316)
Katian453.0 - 445.2Kazakhstan Agerina acutilimbata (162197 162198)