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Megasecopteromorpha

Insecta

Synonymy list
YearName and author
2020Megasecopteromorpha Béthoux p. 275

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
RankNameAuthor
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
superorderPalaeodictyopterida(Bechly 1996)
—Megasecopteromorpha
—Megasecopteromorpha

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Megasecopteromorpha Béthoux 2020
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Or. †Diaphanopterodea Handlirsch 1906
+
Fm. †Alexrasnitsyniidae Prokop and Nel 2011
+
G. †Alexrasnitsynia Prokop and Nel 2011
+
†Alexrasnitsynia permiana Prokop and Nel 2011
Fm. †Asthenohymenidae Tillyard 1924
+
G. †Asthenohymen Tillyard 1924
+
†Asthenohymen affinis Tillyard 1926
†Asthenohymen apicalis Carpenter 1947
†Asthenohymen dunbari Tillyard 1924
†Asthenohymen gracilis Tillyard 1926
†Asthenohymen kansasensis Tillyard 1926
†Asthenohymen latus Carpenter 1947
†Asthenohymen micron Sinitshenkova 2004
†Asthenohymen minutum Zalessky 1939
†Asthenohymen minutus Zimmerman 1962
†Asthenohymen parvulus Carpenter 1947
†Asthenohymen pusillus Tillyard 1926
†Asthenohymen stenobasis Tillyard 1926
†Asthenohymen stigmatizans Tillyard 1926
†Asthenohymen triangularis Carpenter 1947
†Asthenohymen uralicum Zalessky 1939
†Asthenohymen zalesskyi Novokshonov and Novokshonova 1998
†Asthenohymen zonatus Sinitshenkova 2004
Superfm. †Asthenohymenidea
+
Fm. †Aenigmatidiidae Rohdendorf 1961
+
G. †Aenigmatidia Rohdendorf 1961
+
†Aenigmatidia kaltanica Rohdendorf 1961
G. †Avionbrodia Schubnel et al. 2020
+
†Avionbrodia incompleta Schubnel et al. 2020
Fm. †Biarmohymenidae Zalessky 1937
+
G. †Anomalohymen Beckemeyer and Engel 2009
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†Anomalohymen dochmus Beckemeyer and Engel 2009
G. †Biarmohymen Zalessky 1937
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†Biarmohymen bardense Zalessky 1937
G. †Diapha Kukalová-Peck 1974
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†Diapha candida Kukalová-Peck 1974
Fm. †Diaphanopteridae Handlirsch 1906
+
G. †Carrizodiaphanoptera Prokop and Kukalová-Peck 2017
+
†Carrizodiaphanoptera permiana Prokop and Kukalová-Peck 2017
G. †Diaphanoptera Brongniart 1894
+
†Diaphanoptera munieri Brongniart 1894
+
Invalid names: Diaphanoptera superba Meunier 1908 [synonym]
†Diaphanoptera vetusta Brongniart 1894
Invalid names: Diaphanoptera scudderi Brongniart 1894 [nomen dubium], Diaphanopterites Handlirsch 1919 [synonym], Pseudanthracothremma Handlirsch 1906 [synonym]
G. †Diaphpterum Pinto 1999
+
†Diaphpterum rasnitsyni Pinto 1999
G. †Philiasptilon Zalessky 1931
+
†Philiasptilon maculosum Zalessky 1931
Invalid names: Diaphanopteritidae Handlirsch 1919 [synonym]
G. †Elmodiapha Kukalová-Peck 1974
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†Elmodiapha ovata Kukalová-Peck 1974
Fm. †Elmoidae Tillyard 1937
+
G. †Elmoa Tillyard 1937
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†Elmoa trisecta Tillyard 1937
G. †Lodevohymen Prokop and Nel 2011
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†Lodevohymen lapeyriei Prokop and Nel 2011
Fm. †Martynoviidae Tillyard 1932
+
G. †Boskovicia Dostal and Prokop 2009
+
†Boskovicia celtai Dostal and Prokop 2009
G. †Eumartynovia Carpenter 1947
+
†Eumartynovia raaschi Carpenter 1947
G. †Martynovia Tillyard 1932
+
†Martynovia halli Beckemeyer 2004
†Martynovia insignis Tillyard 1932
†Martynovia longipennis Carpenter 1947
†Martynovia protohymenoides Tillyard 1932
Invalid names: Martynoviella Tillyard 1932 [synonym]
G. †Moravoptera Dostal and Prokop 2009
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†Moravoptera hladilovae Dostal and Prokop 2009
G. †Phaneroneura Carpenter 1947
+
†Phaneroneura martynovae Carpenter 1947
†Phaneroneura minuta Béthoux et al. 2003
†Phaneroneura reducta Carpenter 1947
†Phaneroneura rineharti Sinitshenkova 2004
G. †Salagouneura Béthoux et al. 2003
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†Salagouneura chimaira Béthoux et al. 2003
Fm. †Namurodiaphidae Kukalová-Peck and Brauckmann 1990
+
G. †Camptodiapha Beckemeyer and Engel 2011
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†Camptodiapha atkinsoni Beckemeyer and Engel 2011
G. †Namurodiapha Kukalová-Peck and Brauckmann 1990
+
†Namurodiapha sippelorum Kukalová-Peck and Brauckmann 1990
Fm. †Parabrodiidae Carpenter 1934
+
G. †Parabrodia Carpenter 1934
+
†Parabrodia carbonaria Carpenter 1934
G. †Piesbergala Brauckmann and Herd 2002
+
†Piesbergala leipnerae Brauckmann and Herd 2002
G. †Paradiapha Kukalová-Peck 1974
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†Paradiapha delicatula Kukalová-Peck 1974
Fm. †Parelmoidae Rohdendorf 1962
+
G. †Parelmoa Carpenter 1947
+
†Parelmoa obtusa Carpenter 1947
†Parelmoa radialis Carpenter 1947
†Parelmoa revelata Carpenter 1947
G. †Permelmoa Prokop and Nel 2011
+
†Permelmoa magnifica Prokop and Nel 2011
G. †Permuralia Sinitshekova and Kukalová-Peck 1997
+
†Permuralia maculata Kukalová-Peck and Sinichenkova 1992
†Permuralia sharovi Kukalová-Peck and Sinichenkova 1992
Invalid names: Uralia Kukalová-Peck and Sinichenkova 1992 [replaced]
G. †Pseudelmoa Carpenter 1947
+
†Pseudelmoa ampla Carpenter 1947
G. †Sinoelmoa Yang et al. 2024
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†Sinoelmoa yangquanensis Yang et al. 2024
Fm. †Paruraliidae Kukalová-Peck and Sinichenkova 1992
+
G. †Paruralia Kukalová-Peck and Sinichenkova 1992
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†Paruralia carpenteri Kukalová-Peck and Sinichenkova 1992
†Paruralia rohdendorfi Kukalová-Peck and Sinichenkova 1992
G. †Permodiapha Kukalová-Peck 1974
+
†Permodiapha carpenteri Kukalová-Peck 1974
†Permodiapha lata Kukalová-Peck 1974
†Permodiapha specula Kukalová-Peck 1974
Fm. †Prochoropteridae Handlirsch 1911
+
G. †Euchoroptera Carpenter 1940
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†Euchoroptera longipennis Carpenter 1940
G. †Prochoroptera Handlirsch 1911
+
†Prochoroptera calopteryx Handlirsch 1911
G. †Protodiapha Kukalová-Peck 1974
+
†Protodiapha lineata Kukalová-Peck 1974
†Protodiapha maculifera Kukalová-Peck 1974
Fm. †Rhaphidiopsidae Handlirsch 1906
+
G. †Rhaphidiopsis Scudder 1893
+
Fm. †Sinodiaphidae Yang et al. 2020
+
G. †Sinodiapha Yang et al. 2020
+
†Sinodiapha ramosa Yang et al. 2020
G. †Stenodiapha Kukalová-Peck 1974
+
†Stenodiapha angusta Kukalová-Peck 1974
†Stenodiapha moravica Kukalová-Peck 1974
Fm. †Velisopteridae Pinto and Adami-Rodrigues 1997
+
G. †Velisoptera Pinto and Adami-Rodrigues 1997
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†Velisoptera taschi Pinto and Adami-Rodrigues 1997
Invalid names: Diaphanopteridea [empty], Eudiaphanopterodea [empty]
Or. †Megasecoptera Brongniart 1885
+
G. †Adiaphtharsia Handlirsch 1906
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†Adiaphtharsia ferrea Handlirsch 1906
Fm. †Alectoneuridae Kukalová-Peck 1975
+
G. †Alectoneura Kukalová-Peck 1975
+
†Alectoneura europaea Kukalová-Peck 1975
Fm. †Anchineuridae Carpenter 1963
+
G. †Anchineura Carpenter 1963
+
†Anchineura hispanica Carpenter 1963
Fm. †Ancopteridae Kukalová-Peck 1975
+
G. †Agaeoleptoptera Beckemeyer and Engel 2011
+
†Agaeoleptoptera uniotempla Beckemeyer and Engel 2011
G. †Ancoptera Kukalová-Peck 1975
+
†Ancoptera permiana Kukalová-Peck 1975
Fm. †Arcioneuridae Kukalová-Peck 1975
+
G. †Anconeura Kukalová-Peck 1975
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†Anconeura havlatai Kukalová-Peck 1975
G. †Arcioneura Kukalová-Peck 1975
+
†Arcioneura juveniles Kukalová-Peck 1975
Fm. †Aspidohymenidae Martynov 1930
+
G. †Aspidohymen Martynov 1930
+
†Aspidohymen chopardi Zalessky 1937
†Aspidohymen extensus Martynov 1930
Fm. †Aspidothoracidae Handlirsch 1919
+
G. †Aspidothorax Brongniart 1894
+
†Aspidothorax hispanicus Santos et al. 2025
†Aspidothorax permianus Sinitshenkova and Aristov 2020
†Aspidothorax triangularis Brongniart 1894
+
Invalid names: Aspidothorax maculatus Brongniart 1894 [synonym], Aspidothorax nanus Handlirsch 1919 [synonym]
Invalid names: Protocapnia Brongniart 1885 [nomen nudum]
Fm. †Bardohymenidae Zalessky 1937
+
G. †Actinohymen Carpenter 1962
+
†Actinohymen russelli Carpenter 1962
G. †Alexahymen Kukalová-Peck 1972
+
†Alexahymen aestatis Brauckmann 1991
†Alexahymen maruska Kukalová-Peck 1972
G. †Anthohymen Hong 1985
+
†Anthohymen hejinensis Hong 1985
G. †Bardohymen Zalessky 1937
+
†Bardohymen magnipennifer Zalessky 1937
G. †Calohymen Carpenter 1947
+
†Calohymen permianus Carpenter 1947
G. †Microhymen Yang et al. 2023
+
†Microhymen inconspicuus Yang et al. 2023
G. †Paleohymen Hong 1985
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G. †Sinohymen Hong 1985
+
G. †Sunohymen Hong 1985
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†Sunohymen xishanensis Hong 1985
G. †Sylvohymen Martynov 1940
+
†Sylvohymen ingens Carpenter 1947
†Sylvohymen marginatus Pecharová et al. 2020
†Sylvohymen peckae Brauckmann 1988
†Sylvohymen robustus Martynov 1940
†Sylvohymen sibiricus Kukalová-Peck 1972
G. †Taigahymen Sinitshenkova 2019
+
†Taigahymen borealis Sinitshenkova 2019
Fm. †Brodiidae Handlirsch 1906
+
G. †Brodia Scudder 1881
+
†Brodia jogginsensis Prokop et al. 2017
†Brodia priscotincta Scudder 1881
+
Invalid names: Brodia fasciata Handlirsch 1919 [synonym], Brodia furcata Handlirsch 1919 [synonym], Brodia nebulosa Handlirsch 1919 [synonym], Brodia nympha Handlirsch 1919 [synonym], Brodia petiolata Handlirsch 1919 [synonym], Brodia pictipennis Handlirsch 1919 [synonym], Brodia scudderi Handlirsch 1919 [synonym]
G. †Eubrodia Carpenter 1967
+
†Eubrodia dabasinskasi Carpenter 1967
G. †Piesbergbrodia Pecharová et al. 2020
+
†Piesbergbrodia tristrata Brauckmann and Herd 2002
Fm. †Caulopteridae Kukalová-Peck 1975
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G. †Cauloptera Kukalová-Peck 1975
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†Cauloptera colorata Kukalová-Peck 1975
Fm. †Corydaloididae Handlirsch 1906
+
G. †Corydaloides Brongniart 1885
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†Corydaloides scudderi Brongniart 1885
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Invalid names: Corydaliodes gracilior Handlirsch 1919 [synonym], Corydaloides brongniarti Handlirsch 1919 [synonym], Corydaloides longipennis Handlirsch 1919 [synonym], Corydaloides simplex Brongniart 1894 [synonym]
Invalid names: Corydaliodes Handlirsch 1919 [synonym]
Fm. †Dictyoneurellidae Kukalová-Peck 1975
+
G. †Dictyoneurella Laurentiaux 1949
+
†Dictyoneurella perfecta Laurentiaux 1949
Fm. †Engisopteridae Kukalová-Peck 1975
+
G. †Engisoptera Kukalová-Peck 1975
+
†Engisoptera ivanae Rýznar 2021
†Engisoptera simplices Kukalová-Peck 1975
Subor. †Eumegasecoptera
+
Fm. †Aykhalidae Sinitshenkova 1993
+
G. †Aykhal Sinitshenkova 1993
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†Aykhal helenae Sinitshenkova 1993
G. †Namuroptera Pecharová et al. 2015
+
†Namuroptera minuta Pecharová et al. 2015
G. †Sinopalaeopteryx Pecharová et al. 2015
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†Sinopalaeopteryx olivieri Pecharová et al. 2015
†Sinopalaeopteryx splendens Pecharová et al. 2015
Fm. †Brodiopteridae Carpenter 1963
+
G. †Brodioptera Copeland 1957
+
†Brodioptera cumberlandensis Copeland 1957
†Brodioptera pintoi Brauckmann et al. 2003
†Brodioptera sinensis Pecharová et al. 2015
†Brodioptera stricklani Nelson and Tidwell 1987
Fm. †Foririidae Handlirsch 1919
+
G. †Foriria Meunier 1908
+
†Foriria maculata Meunier 1908
Fm. †Ischnoptilidae Carpenter 1951
+
G. †Ischnoptilus Brongniart 1894
+
†Ischnoptilus elegans Brongniart 1894
G. †Lameereites Handlirsch 1911
+
†Lameereites curvipennis Handlirsch 1911
Fm. †Mischopteridae Handlirsch 1906
+
G. †Mischoptera Brongniart 1894
+
†Mischoptera douglassi Carpenter and Richardson 1968
†Mischoptera nigra Brongniart 1885
+
Invalid names: Mischoptera concolor Handlirsch 1919 [synonym], Mischoptera infumata Handlirsch 1919 [synonym], Mischoptera ocellata Handlirsch 1919 [synonym], Mischoptera woodwardi Brongniart 1894 [synonym]
Invalid names: Woodwardia Brongniart 1885 [replaced]
G. †Psilothorax Brongniart 1894
+
†Psilothorax longicauda Brongniart 1894
+
Invalid names: Psilothorax pictus Handlirsch 1919 [synonym]
Fm. †Moravohymenidae Kukalová-Peck 1972
+
G. †Issadohymen Sinitshenkova and Aristov 2013
+
†Issadohymen ponomarenkoi Sinitshenkova and Aristov 2013
G. †Moravohymen Kukalová-Peck 1972
+
†Moravohymen vitreus Kukalová-Peck 1972
Fm. †Piesbergpteridae Rosová et al. 2022
+
G. †Piesbergptera Rosová et al. 2022
+
†Piesbergptera niesseae Rosová et al. 2022
Fm. †Protohymenidae Tillyard 1924
+
G. †Arroyohymen Prokop and Kukalová-Peck 2017
+
†Arroyohymen splendens Prokop and Kukalová-Peck 2017
G. †Carbohymen Pecharová and Prokop 2018
+
†Carbohymen testai Pecharová and Prokop 2018
G. †Ivahymen Martynov 1932
+
†Ivahymen constrictus Martynov 1932
G. †Permohymen Tillyard 1924
+
†Permohymen schucherti Tillyard 1924
G. †Protohymen Tillyard 1924
+
†Protohymen angustipennis Martynov 1932
†Protohymen anomalus Zimmerman 1962
†Protohymen bifurcatus Zimmerman 1962
†Protohymen carpenteri Novokshonov 1995
†Protohymen concinna Zalessky 1956
†Protohymen curvatus Carpenter 1947
†Protohymen elongatus Carpenter 1930
†Protohymen largus Carpenter 1947
†Protohymen latus Carpenter 1947
†Protohymen novokshonovi Pecharová and Prokop 2018
†Protohymen permianus Tillyard 1924
†Protohymen pictus Zimmerman 1962
†Protohymen readi Carpenter 1933
†Protohymen shafferi Zimmerman 1962
†Protohymen tenuis Carpenter 1939
†Protohymen venustus Carpenter 1947
Invalid names: Pseudohymen Martynov 1932 [synonym], Pseudohymenopsis Zalessky 1956 [synonym]
Fm. †Scytohymenidae Martynov 1937
+
G. †Karoohymen Riek 1976
+
†Karoohymen minutus van Dijk and Geertsema 1999
G. †Oceanoptera Shcherbakov 2009
+
†Oceanoptera elenae Shcherbakov 2009
G. †Scytohymen Martynov 1937
+
†Scytohymen extremus Martynov 1937
G. †Tshekardohymen Rohdendorf 1940
+
†Tshekardohymen martynovi Rohdendorf 1940
Fm. †Sphecopteridae Carpenter 1951
+
G. †Cyclocelis Brongniart 1894
+
†Cyclocelis chatini Brongniart 1894
+
Invalid names: Sphecoptera brongniartiana Handlirsch 1919 [synonym], Sphecoptera fenestrata Handlirsch 1919 [synonym]
†Cyclocelis devexa Sinitshenkova 2019
†Cyclocelis obscura Brongniart 1894
†Cycloscelis elegantissima Meunier 1908
G. †Sphecoptera Brongniart 1894
+
†Sphecoptera brongniarti Meunier 1908
†Sphecoptera elegantissima Meunier 1908
†Sphecoptera gracilis Brongniart 1894
†Sphecoptera minor Brongniart 1894
†Sphecoptera pulchra Brongniart 1894
Invalid names: Cyclocelis acuta Brongniart 1894 [nomen dubium]
Fm. †Sphecorydaloididae Pinto 1994
+
G. †Sphecorydaloides Pinto 1994
+
Fm. †Vorkutiidae Rohdendorf 1947
+
G. †Fragmohymen Novokshonov 1995
+
†Fragmohymen submissus Novokshonov 1995
G. †Siberiohymen Rohdendorf 1961
+
†Siberiohymen asiaticus Rohdendorf 1961
G. †Vorkutia Rohdendorf 1947
+
†Vorkutia dimina Novokshonov 1998
†Vorkutia tshernovi Rohdendorf 1947
Fm. †Xenopteraidae Pinto 1986
+
G. †Sinitshenkovae Nel et al. 2019
+
†Sinitshenkovae gallica Nel et al. 2019
†Sinitshenkovae huenickeni Pinto and de Ornellas 1978
G. †Xenoptera Pinto 1986
+
†Xenoptera riojaensis Pinto 1986
Invalid names: Archaeoscolex Matthew 1889 [nomen dubium], Archaeoscolex corneus Matthew 1889 [nomen dubium], Protohymenoptera Tillyard 1924 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Kinderscoutian to the top of the Changhsingian or 321.80000 to 251.90200 Ma

Collections (105 total)


Time interval Ma Country or state Original ID and collection number
Bashkirian323.4 - 315.2Argentina (La Rioja) Xenoptera riojaensis (108150)
Kinderscoutian321.8 - 321.3USA (Utah) Brodioptera stricklani (122484)
Marsdenian321.3 - 320.0Germany (Ruhr) Namurodiapha sippelorum, Sylvohymen peckae (110939) Sylvohymen pintoi (152121)
Westphalian A318.7 - 316.9Canada (New Brunswick) Archaeoscolex corneus (114468)
Westphalian A318.7 - 316.9United Kingdom (England) Brodia priscotincta (167683)
Westphalian A318.7 - 316.9Canada (Nova Scotia) Brodia jogginsensis (187589) Brodioptera cumberlandensis (122485)
Westphalian318.7 - 307.4USA (Alabama) Camptodiapha atkinsoni, Agaeoleptoptera uniotempla (81525)
Westphalian318.7 - 307.4USA (Rhode Island) Rhaphidiopsis diversipenna (112701)
Westphalian B316.9 - 314.6United Kingdom (Staffordshire) Brodia furcata, Brodia nebulosa, Brodia pictipennis, Brodia nympha (34906) Brodia priscotincta (167682) Brodia priscotincta, Brodia scudderi (152262)
Westphalian B316.9 - 314.6France (Nord-Pas-de-Calais) Dictyoneurella perfecta (152792)
Westphalian B316.9 - 314.6United Kingdom (England) Brodia fasciata, Brodia nympha (125238)
Westphalian B - Westphalian C316.9 - 309.8China (Ningxia) Brodioptera sinensis, Sinopalaeopteryx splendens, Sinopalaeopteryx olivieri, Namuroptera minuta, Sinodiapha ramosa, Microhymen inconspicuus (139576)
Westphalian C314.6 - 309.8United Kingdom (England) Brodia priscotincta (127525)
Westphalian C314.6 - 309.8United Kingdom (Staffordshire) Brodia priscotincta (167681)
Westphalian C - Westphalian D314.6 - 307.4Germany (Lower Saxony) Aspidothorax tristrata (122514)
Westphalian C - Westphalian D314.6 - 307.4France Megasecoptera indet., Avionbrodia incompleta (122504)
Westphalian D309.8 - 307.4Germany (Lower Saxony) Aspidothorax aestatis (172341) Aspidothorax tristrata (122515) Piesbergala leipnerae (122518) Piesbergptera niesseae (224159)
Westphalian D309.8 - 307.4USA (Illinois) Adiaphtharsia ferrea (146857) Carbohymen testai (127721) Eubrodia dabasinskasi (151848) Mischoptera douglassi (122538 161914) Mischoptera sp. (124839) Prochoroptera calopteryx (122493 124837 124840 124841) Prochoroptera calopteryx, Eubrodia dabasinskasi (112637) Prochoroptera calopteryx, Eubrodia dabasinskasi, Mischoptera douglassi (151852) Prochoroptera calopteryx, Lameereites curvipennis (112630)
Kasimovian307.0 - 303.7USA (Kansas) Megasecoptera indet., Euchoroptera longipennis (122507) Parabrodia carbonaria (122510)
Kasimovian - Gzhelian307.0 - 298.9Argentina (San Luis) Megasecoptera indet., Philiasptilon huenickeni, Velisoptera taschi, Diaphpterum rasnitsyni, Sphecorydaloides lucchesei (35057)
Gzhelian303.7 - 298.9Spain (Leon) Anchineura hispanica (105374)
Gzhelian303.7 - 298.9Russian Federation (Sakha) Aykhal helenae (126060)
Gzhelian303.7 - 298.9Spain (León) Aspidothorax hispanicus (239637)
Gzhelian303.7 - 298.9France Aspidothorax triangularis, Aspidothorax maculatus, Aspidothorax nanus, Woodwardia nigra, Psilothorax longicauda, Sphecoptera gracilis, Cyclocelis minor, Cyclocelis chatini, Sphecoptera brongniartiana, Cyclocelis obscura, Ischnoptilus elegans, Corydaloides scudderi, Corydaloides brongniarti, Corydaloides longipennis, Corydaloides simplex, Corydaliodes gracilior, Diaphanoptera munieri, Diaphanoptera vetusta, Anthracothremma scudderi, Mischoptera woodwardi, Cyclocelis acuta, Sphecoptera pulchra, Mischoptera concolor, Mischoptera ocellata, Mischoptera infumata, Psilothorax pictus, Sphecoptera fenestrata (103635) Mischoptera nigra, Sphecoptera brongniarti, Foriria maculata, Diaphanoptera superba, Cycloscelis elegantissima (121979)
Gzhelian303.7 - 298.9Russian Federation Philiasptilon maculosum (124591) Siberiohymen asiaticus (124584)
Asselian298.9 - 293.52Czechia (Moravia) Engisoptera ivanae (217298) Martynovia insignis, Boskovicia celtai, Moravoptera hladilovae (117144)
Asselian298.9 - 293.52Czechia Alexahymen maruska, Moravohymen vitreus, Elmodiapha ovata, Protodiapha maculifera, Protodiapha lineata, Diapha candida, Permodiapha sp., Permodiapha carpenteri, Permodiapha lata, Permodiapha specula, Stenodiapha angusta, Stenodiapha moravica, Paradiapha delicatula, Cauloptera colorata, Arcioneura juveniles, Anconeura havlatai, Ancoptera sp., Ancoptera permiana, Engisoptera simplices, Alectoneura europaea (114186)
Asselian298.9 - 293.52USA (New Mexico) Arroyohymen splendens (189778) Asthenohymenidae indet., Phaneroneura rineharti, Asthenohymen micron, Asthenohymen zonatus (124386) Martynovia insignis, Carrizodiaphanoptera permiana (122481)
Asselian298.9 - 293.52China (Shanxi) Sinoelmoa yangquanensis (235985)
Sakmarian293.52 - 290.1China (Shanxi) Paleohymen shanxiensis, Paleohymen renmazhuangensis, Sinohymen gancaoshanensis (140933)
Artinskian290.1 - 283.3USA (Oklahoma) Anomalohymen dochmus (104828) Asthenohymen apicalis (104820) Elmoa trisecta, Parelmoa radialis, Phaneroneura martynovae, Protohymen curvatus, Protohymen latus, Asthenohymen apicalis, Asthenohymen latus (104822) Martynovia halli (104818) Martynovia longipennis (104890) Parelmoa revelata, Parelmoa radialis, Parelmoa obtusa, Pseudelmoa ampla, Calohymen permianus, Eumartynovia raaschi, Phaneroneura martynovae, Phaneroneura reducta, Protohymen curvatus, Protohymen venustus, Protohymen largus, Asthenohymen triangularis, Asthenohymen apicalis, Asthenohymen parvulus, Asthenohymen latus (104824) Protohymen bifurcatus, Protohymen readi, Asthenohymen minutus (104826) Protohymen curvatus, Protohymen largus (104823) Protohymen shafferi, Protohymen anomalus, Protohymen pictus (104827) Sylvohymen ingens (104889)
Artinskian290.1 - 283.3USA (Texas) Actinohymen russelli (111540)
Artinskian290.1 - 283.3USA (Kansas) Elmoa trisecta, Asthenohymen sp., Protohymen permianus, Permohymen schucherti, Asthenohymen dunbari, Asthenohymen affinis, Asthenohymen gracilis, Asthenohymen stigmatizans, Asthenohymen kansasensis, Asthenohymen stenobasis, Asthenohymen pusillus, Martynovia insignis, Martynoviella protohymenoides (111219) Elmoa trisecta, Protohymen permianus, Asthenohymen dunbari, Asthenohymen affinis, Martynovia protohymenoides (111320) Elmoa trisecta, Protohymen readi, Protohymen permianus, Asthenohymen dunbari, Asthenohymen affinis, Asthenohymen pusillus, Protohymen elongatus, Martynovia insignis (111319) Protohymen permianus, Permohymen schucherti, Asthenohymen dunbari, Asthenohymen pusillus, Protohymen tenuis (114152) Protohymen permianus, Permohymen schucherti, Protohymen elongatus, Doter affinis, Doter pusillus, Doter minor (111217) Protohymen readi, Protohymen permianus, Permohymen schucherti, Asthenohymen dunbari, Asthenohymen pusillus, Protohymen elongatus (114151)
Artinskian290.1 - 283.3France (Languedoc) Martynovia halli, Permelmoa magnifica (105044) Martynoviidae indet. (122923) Martynoviidae indet., Phaneroneura minuta, Salagouneura chimaira (122883)
Artinskian290.1 - 283.3China (Shanxi) Sunohymen xishanensis (140458)
Kungurian283.3 - 274.4Russian Federation (Suksun) Protohymen novokshonovi (122753) Pseudohymen sp., Pseudohymen sylvaensis, Sylvohymen robustus (116641) Sylvohymen marginatus (113380) Sylvohymen sibiricus, Pseudohymen carpenteri, Uralia maculata, Uralia sharovi, Paruralia rohdendorfi, Paruralia carpenteri (108206) Tshekardohymen martynovi (124241)
Kungurian283.3 - 274.4France Alexrasnitsynia permiana (117604) Martynoviidae indet., Martynovia insignis, Phaneroneura minuta, Salagouneura chimaira (103637)
Kungurian283.3 - 274.4Russian Federation (Komi republic) Cyclocelis devexa (201683) Vorkutia tshernovi (160744)
Kungurian283.3 - 274.4China (Shanxi) Anthohymen hejinensis (140456)
Kungurian283.3 - 274.4Russian Federation (Kishertskij) Pseudohymen angustipennis (111188)
Kungurian283.3 - 274.4Russian Federation Asthenohymen uralicum, Asthenohymen minutum (155817) Biarmohymen bardense (155819) Pseudohymenopsis concinna, Bardohymen magnipennifer (161939) Pseudohymenopsis minor (102504) Scytohymen extremus, Fragmohymen submissus, Asthenohymen uralicum (114207) Scytohymenidae indet., Oceanoptera elenae (115214) Vorkutia dimina, Asthenohymen uralicum (107787)
Kungurian283.3 - 274.4France (Languedoc) Lodevohymen lapeyriei, Sinitshenkovae gallica (105045) Martynoviidae indet. (122884)
Kungurian283.3 - 274.4Russian Federation (Komi Republic) Taigahymen borealis (201684)
Kungurian - Early/Lower Kazanian283.3 - 266.9Russian Federation Scytohymen extremus (116612)
Roadian274.4 - 266.9Russian Federation (Perm) Aspidothorax permianus (210714)
Roadian274.4 - 266.9Russian Federation (Kazan) Aspidohymen extensus (116803)
Roadian274.4 - 266.9Russian Federation (Kemerovo) Aenigmatidia kaltanica (122569)
Roadian274.4 - 266.9Russian Federation Aspidohymen chopardi (122871)
Roadian274.4 - 266.9Russian Federation (Arkangelsk) Ivahymen constrictus (111187)
Wuchiapingian259.51 - 254.14Russian Federation (Vologda) Issadohymen ponomarenkoi (104754)
Changhsingian254.14 - 251.902South Africa (Natal) Karoohymen delicatulus, Karoohymen minutus (124428)