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Alphacoleoptera

Insecta - Coleoptera

Synonymy list
YearName and author
2022Alphacoleoptera Cai et al.
2022Alphacoleoptera Engel
2022Alphacoleoptera Engel et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
Dicondylia
Paranotalia
RankNameAuthor
subclassPterygota()
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Aparaglossata
Neuropteriformia
Coleopterida
orderColeoptera
suborderAlphacoleoptera
suborderAlphacoleoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. †Alphacoleoptera Engel et al. 2022
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Superfm. †Coleopsoidea Kirejtshuk and Nel 2016
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Fm. †Coleopsidae Kirejtshuk and Nel 2016
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G. †Coleopsis Kirejtshuk et al. 2014
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Coleopsis archaica Kirejtshuk et al. 2014
Superfm. †Permocupedoidea Martynov 1933
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Fm. †Labradorocoleidae Ponomarenko 1969
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G. †Labradorocoleus Ponomarenko 1969
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Labradorocoleus carpenteri Ponomarenko 1969
Fm. †Oborocoleidae Kukalová 1968
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G. †Liberocoleus Kukalová 1968
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Liberocoleus intactus Kukalová 1968
G. †Oborocoleus Kukalová 1968
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Oborocoleus rohdendorfi Kukalová 1968
Invalid names: Tshekardocoleoidea Rodendorf 1944 [synonym]
Superfm. †Permosynoidea Tillyard 1924
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Fm. †Permosynidae Tillyard 1924
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G. †Alveolacupes Lin 1983
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Alveolacupes primus Lin 1983
G. †Alveolicupes Lin 1983
G. †Artematopodites Ponomarenko 1990
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Invalid names: Ovivagina immediata Zhang 1997 [synonym]
Artematopodites latissimus Ponomarenko 2014
Artematopodites latus Ponomarenko 1990
Artematopodites leskoviensis Yan and Zhang 2010
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Invalid names: Artematopodites longus Ponomarenko 1990 [replaced]
Artematopodites lozowskii Ponomarenko 2015
Artematopodites major Ponomarenko 1990
G. †Bistrisyne Lin 1986
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Bistrisyne tenua Lin 1986
G. †Delpuentesyne Martins-Neto and Gallego 2007
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Delpuentesyne menendezi Martins-Neto and Gallego 2007
G. †Diarcuipenna Lin 1986
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Diarcuipenna bennettitophila Ponomarenko 2011
G. †Dinoharpalus Handlirsch 1906
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Dinoharpalus coptoclavoides Ponomarenko 2015
Dinoharpalus latus Ponomarenko 2015
Dinoharpalus liasinus Giebel 1856
Dinoharpalus martynovi Ponomarenko 2018
Dinoharpalus rugosus Ponomarenko 1985
G. †Dzeregia Ponomarenko 1985
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Dzeregia ampla Ponomarenko 1990
Dzeregia byrrhoides Ponomarenko 1990
Dzeregia crassa Ponomarenko 1990
Dzeregia juxta Lin 1986
Dzeregia lata Ponomarenko 1985
Dzeregia longa Ponomarenko 1985
Dzeregia pilula Ponomarenko 1990
Dzeregia platis Ponomarenko 2014
Dzeregia striata Ponomarenko 1985
G. †Hydrobiites Heer 1864
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Hydrobiites anglicus Handlirsch 1906
Hydrobiites bellus Geinitz 1894
Hydrobiites convexus Ponomarenko 1985
Hydrobiites crassus Ponomarenko 1985
Hydrobiites dobbertinensis Handlirsch 1939
Hydrobiites giebeli Handlirsch 1906
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Invalid names: Berosus liasinus Giebel 1856 [replaced]
Hydrobiites handlirschi Ponomarenko 2011
Hydrobiites liasinus Giebel 1856
Hydrobiites longus Ponomarenko 1990
Hydrobiites minor Ponomarenko 2014
Hydrobiites minutissimus Ponomarenko 1985
Hydrobiites mongolicus Ponomarenko 2014
Hydrobiites permianus Ponomarenko 2018
Hydrobiites punctatostriatus Handlirsch 1939
Hydrobiites purbeccensis Giebel 1856
Hydrobiites sulcatus Ponomarenko 1986
Hydrobiites tillyardi Ponomarenko 2011
Hydrobiites veteranus Heer 1852
Hydrobiites vladimiri Ponomarenko 2011
G. †Ischichucasyne Martins-Neto and Gallego 2005
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Ischichucasyne cladocosta Martins-Neto and Gallego 2009
Ischichucasyne santajuanaensis Martins-Neto and Gallego 2005
G. †Palademosyne Rohdendorf 1961
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Palademosyne martynovae Rohdendorf 1961
Palademosyne natalensis Ponomarenko 2005
G. †Permocrossos Martynov 1932
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Permocrossos elongatus Martynov 1932
G. †Permosyne Tillyard 1924
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Permosyne affinis Tillyard 1924
Permosyne belmontensis Tillyard 1924
Permosyne dentata Ponomarenko 2003
Permosyne elongata Ponomarenko 2005
Permosyne mitchelli Tillyard 1924
Permosyne rasnitsyni Ponomarenko 2011
G. †Platycrossos Dunstan 1923
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Platycrossos caroli Ponomarenko 2011
Platycrossos elongatus Ponomarenko 1985
Platycrossos latus Ponomarenko 2014
Platycrossos ligulatus Dunstan 1923
Platycrossos longus Ponomarenko 2014
Platycrossos loxonicus Ponomarenko 2014
Platycrossos mongolicus Ponomarenko 2014
Platycrossos ovum Ponomarenko 2014
Platycrossos petalus Ponomarenko 1985
Platycrossos punctatus Ponomarenko 1985
Platycrossos subtumidus Dunstan 1923
Platycrossos tumidus Tillyard 1916
G. †Pseudorhynchophora Handlirsch 1906
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Pseudorhynchophora convexa Ponomarenko 2008
G. †Sakmaracoleus Ponomarenko 2013
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Sakmaracoleus orenburgensis Ponomarenko 2013
G. †Tychtocoleus Rohdendorf 1961
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Tychtocoleus europeus Ponomarenko 2003
Tychtocoleus neuburgae Rohdendorf 1961
Tychtocoleus popovi Ponomarenko 2018
G. †Umkomaasia Zeuner 1960
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Umkomaasia depressa Zeuner 1960
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Asselian to the top of the Cenomanian or 298.90000 to 93.90000 Ma

Collections (76 total)


Time interval Ma Country or state Original ID and collection number
Asselian298.9 - 295.5Germany (Rhineland-Palatinate) Coleopsis archaica (156248)
Asselian298.9 - 295.5Czech Republic Oborocoleus rohdendorfi, Liberocoleus intactus (114186)
Roadian272.3 - 268.8Russian Federation (Arkangelsk) Permocrossos elongatus (104993)
Wordian266.9 - 264.28Russian Federation (Kemerovo) Palademosyne martynovae (124612)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Sakmaracoleus orenburgensis (152235)
Wuchiapingian259.9 - 254.17Russian Federation Tychtocoleus neuburgae (122565 124608)
Changhsingian254.17 - 252.17Russian Federation (Vologda) Artematopodites lozowskii, Dinoharpalus latus (105046) Permosynidae indet., Permosyne dentata, Permosyne europaea, Hydrobiites permianus, Tychtocoleus popovi, Dinoharpalus martynovi (105041)
Changhsingian254.17 - 252.17Australia (New South Wales) Permosyne affinis, Permosyne mitchelli (122424) Permosyne belmontensis (122414)
Changhsingian254.17 - 252.17Russian Federation (Vladimir) Hydrobiites tillyardi (104756) Hydrobiites tillyardi, Hydrobiites vladimiri, Permosyne rasnitsyni (113351)
Changhsingian254.17 - 252.17South Africa (Kwazulu-Natal) Palademosyne natalensis, Permosyne elongata (14356)
Olenekian251.2 - 247.2Russian Federation (Arkhangelsk) Pseudorhynchophora convexa (108318)
Longobardian242.0 - 235.0Argentina (Mendoza) Delpuentesyne menendezi (115104)
Carnian237.0 - 228.0Argentina (La Rioja) Ischichucasyne cladocosta (167475) Permosynidae indet. (123047)
Carnian237.0 - 228.0South Africa Umkomaasia depressa (151912)
Carnian237.0 - 228.0Chile (Concepción) Ischichucasyne santajuanaensis (123024)
Carnian237.0 - 228.0Austria Platycrossos caroli, Hydrobiites handlirschi, Diarcuipenna bennettitophila (102471)
Tuvalian232.0 - 221.5Germany Dinoharpalus coptoclavoides (153474)
Norian228.0 - 208.5China (Hunan) Diarcuipenna heterosa, Ademosynoides juxta (155362)
Norian228.0 - 208.5Australia (Queensland) Ademosyne tumida (111777) Platycrossos ligulatus, Platycrossos subtumidus (111558)
Rhaetian208.5 - 201.3China (Hunan) Alveolacupes primus, Alveolacupes secundes, Bistrisyne tenua (155364)
Rhaetian208.5 - 201.3United Kingdom (England) Coleoptera indet. 4, Coleoptera indet. 3 (114677) Harpalus liasinus, Berosus sp. (115246)
Hettangian201.3 - 199.3Poland (Swietokrzyskie) Hydrobiites sp., Artematopodites spp. (123446)
Angulata201.3 - 196.5Switzerland Carabites bellus, Hydrobius veteranus (131766)
Hettangian - Sinemurian201.3 - 190.8China (Xinjiang) Ovivagina prolixa, Ovivagina propinqua (103477)
Early/Lower Toarcian183.0 - 182.0Germany Hydrobiites dobbertinensis, Hydrobiites punctatostriatus (122986)
Toarcian182.7 - 174.1Australia (Western Australia) Platycrossos sp. (114239)
Toarcian182.7 - 174.1Russian Federation (Kemerovo) Hydrobiites bellus (125473)
Toarcian182.7 - 174.1Russian Federation (Irkutsk) Hydrobiites crassus (122950 125476)
Toarcian182.7 - 174.1Russian Federation (Krasnoyarsk) Dzeregia lata (110819)
Toarcian182.7 - 174.1China (Xinjiang) Ovivagina insculpta (103478)
Toarcian182.7 - 174.1Mongolia Dzeregia striata (124855)
Toarcian - Aalenian182.7 - 170.3Russian Federation (Kemerovo) Dinoharpalus rugosus, Dzeregia striata, Dzeregia longa (125475) Hydrobiites bellus (117989) Hydrobiites convexus, Dinoharpalus rugosus (125471) Platycrossos punctatus, Dzeregia lata (125479)
Aalenian174.1 - 170.3Russian Federation (Altai) Dzeregia lata (125480 125481)
Aalenian - Bajocian174.1 - 168.3China (Xinjiang) Ovivagina immediata (103479)
Bajocian170.3 - 168.3Russian Federation (Kemerovo) Hydrobiites convexus, Dzeregia longa (125477)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Hydrobiites bellus, Hydrobiites minutissimus, Platycrossos petalus, Platycrossos elongatus, Platycrossos punctatus, Dzeregia lata (118010)
Bathonian168.3 - 166.1Mongolia Dzeregia striata, Dzeregia lata, Dzeregia longa (119621)
Bathonian168.3 - 166.1Russian Federation (Krasnoyarsk) Platycrossos punctatus (125474)
Callovian - Oxfordian166.1 - 157.3China (Hebei) Sinonitidulina longa (154788)
Callovian - Oxfordian166.1 - 157.3Russian Federation (Chita) Hydrobiites crassus, Dzeregia longa (86023)
Tithonian152.1 - 145.0Russian Federation (Chita) Artematopodites latus (152168) Dzeregia byrrhoides (125407)
Tithonian152.1 - 145.0Mongolia Artematopodites crispulus, Artematopodites lepidus (121393) Artematopodites maximus (125977 165036) Artematopodites maximus, Artematopodites crispulus, Artematopodites lepidus, Hydrobiites mongolicus, Platycrossos mongolicus (104748) Hydrobiites mongolicus (125192) Hydrobiites mongolicus, Hydrobiites minor, Dzeregia platis, Artematopodites latissimus, Platycrossos latus, Platycrossos loxonicus, Platycrossos longus, Platycrossos ovum (121394)
Tithonian152.1 - 145.0Russian Federation Artematopodites latus (125408) Hydrobiites crassus (119398)
Middle Berriasian145.0 - 140.2United Kingdom (England) Hydrophilidae indet. (104008)
Berriasian145.0 - 139.8Russian Federation (Chita) Dzeregia ampla (152170)
Valanginian139.8 - 132.9China (Shaanxi) Artematopodites shaanbeiensis (140460 140461 140462)
Barremian129.4 - 125.0Russian Federation (Chita) Artematopodites longus (151642) Hydrobiites longus (152180)
Aptian125.0 - 113.0Mongolia (Kobdo aimak) Hydrobiites sulcatus (104054)
Aptian125.0 - 113.0Russian Federation (Buryatia) Artematopodites sp. (113349 113350)
Aptian125.0 - 113.0Mongolia (Gobi-Altai aimak) Hydrobiites sulcatus (104055)
Early/Lower Aptian121.4 - 122.46Russian Federation (Chita) Artematopodites major, Dzeregia pilula (114316) Dzeregia crassa (119499)
Cenomanian100.5 - 93.9Canada (Newfoundland and Labrador) Labradorocoleus carpenteri (119438)