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Endymenoptera

Insecta

Taxonomy
Endymenoptera was named by Engel (2022). It is extant.

It was assigned to Amphiesmenoptera by Wang et al. (2022); and to Panendymenoptera by Engel (2022).

Sister taxa
Stelloptera

Synonymy list
YearName and author
2022Endymenoptera Engel p. 6
2022Endymenoptera Wang et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
Dicondylia
Paranotalia
subclassPterygota()
RankNameAuthor
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Aparaglossata
Panorpida
superorderAmphiesmenopteraKiriakoff 1948
Metamphiesmenoptera
Euamphiesmenoptera
Panendymenoptera
Endymenoptera
Endymenoptera

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Endymenoptera Engel 2022
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Unr. Stelloptera Engel 2022
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Or. Lepidoptera Linnaeus 1758 [moth]
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Superfm. Agathiphagoidea Kristensen 1967
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Fm. Agathiphagidae Kristensen 1967 [kauri moth]
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G. †Agathiphagama Mey et al. 2021
Unr. Angiospermivora Regier et al. 2015
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Subor. Glossata Fabricius 1775
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Unr. Coelolepida Nielsen and Kristensen 1996
Superfm. Eriocranioidea Rebel 1901
G. †Protolepis Kozlov 1989
Invalid names: Dacnonypha Hinton 1946 [invalid subgroup]
Fm. †Archaeolepiidae Whalley 1985
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G. †Archaeolepis Whalley 1985
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Archaeolepis mane Whalley 1985
Fm. †Ascololepidopterigidae Zhang et al. 2013
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G. †Ascololepidopterix Zhang et al. 2013
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Ascololepidopterix multinerve Zhang et al. 2013
G. †Pegolepidopteron Zhang et al. 2013
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Pegolepidopteron latiala Zhang et al. 2013
G. †Trionolepidopteron Zhang et al. 2013
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Trionolepidopteron admarginis Zhang et al. 2013
G. †Bombycites Heer 1849
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Bombycites buchii Heer 1864
G. †Chionaemopsis Cockerell and Le Veque 1931
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Chionaemopsis quadrifasciatus Cockerell and Le Veque 1931
G. Cyphanta Walker 1865
Unr. Diurni Latreille 1802
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Invalid names: Papilionides Latreille 1802 [empty]
Subor. †Eolepidopterigina Rasnitsyn 1983
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Superfm. †Eolepidopterigoidea Rasnitsyn 1983
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Fm. †Eolepidopterigidae Rasnitsyn 1983
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Invalid names: Undopterigidae Kozlov 1988 [synonym]
G. Eucosmia Stephens 1829
G. †Gallerites Kernbach 1967
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Gallerites keleri Kernbach 1967
Subor. Heterobathmiina Kristensen and Nielsen 1983
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Superfm. Heterobathmioidea Kristensen and Nielsen 1979
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Fm. Heterobathmiidae Kristensen and Nielsen 1979
G. †Karataunia Kozlov 1989
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Karataunia lapidaria Kozlov 1989
Superfm. Micropterigoidea Herrich-Schaeffer 1855
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Fm. Micropterigidae Cotes 1889
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G. †Archmosaicus Zhang et al. 2020
G. †Auliepterix Kozlov 1989
G. †Baltimartyria Skalski 1995
G. †Bitterfeldopterix Mey 2019
G. †Electrocrania Kuznetsov 1941
G. †Klausnitzerellus Mey 2019
G. Micropterix Hübner 1825
G. †Moleropterix Engel and Kinzelbach 2008
G. †Palaeosabatinca Kozlov 1988
G. †Parasabatinca Whalley 1978
G. Sabatinca Walker 1863
G. †Saxomicropterix Mey 2019
G. †Terncladus Han et al. 2024
G. †Wichardius Mey 2019
Unr. Nocturni Latreille 1802
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Invalid names: Bombycinae Latreille 1802 [empty], Phalaenites Latreille 1802 [empty], Pterophorii Latreille 1802 [empty], Tortrices Latreille 1802 [empty]
G. †Paleolepidopterites Kozlov 2018
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Unr. †Paralepidoptera Engel 2022
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Fm. †Mesokristenseniidae Huang et al. 2010
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G. †Kladolepidopteron Zhang et al. 2013
G. †Mesokristensenia Huang et al. 2010
G. †Petisca Martins-Neto 1998
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Petisca dryellina Martins-Neto 1998
G. †Phylledestes Cockerell 1907
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Phylledestes vorax Cockerell 1907
G. †Pyralites Heer 1856
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Pyralites obscurus Heer 1856
Pyralites preecei Jarzembowski 1980
G. Schaefferia Houlbert 1918
G. †Sphingidites Kernbach 1967
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Sphingidites weidneri Kernbach 1967
Invalid names: Aglossata Speidel 1977 [invalid subgroup], Monotrysia Borner 1939 [empty], Papilionida Laicharting 1781 [synonym], Phalaena Linnaeus 1758 [invalid subgroup], Zalepidoptera Engel 2022 [empty], Zeugloptera Chapman 1917 [empty]
Or. Trichoptera Kirby 1815 [caddisfly]
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G. †Amechanites Haupt 1956
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Amechanites costatus Haupt 1956
G. †Archotaulius Handlirsch 1906
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Archotaulius bavaricus Handlirsch 1906
G. †Axiomaldia Melnitsky and Ivanov 2020
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Axiomaldia flinti Melnitsky and Ivanov 2020
G. †Calamodontus Botosaneanu and Wichard 1983
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Calamodontus grandaevus Botosaneanu and Wichard 1983
Fm. †Cladochoristidae Riek 1953
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G. †Cladochorista Tillyard 1926
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Cladochorista belmontensis Tillyard 1926
Cladochorista curta Sukatsheva and Sinitshenkova 2023
Cladochorista issadica Sukatsheva and Aristov 2013
Cladochorista multivenosa Sukatsheva 1973
G. †Cladochoristella Riek 1955
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Cladochoristella ryzhkovae Sukatsheva and Aristov 2013
Cladochoristella sola Sukatsheva and Sinitshenkova 2023
G. †Meloclada Melnitsky and Ivanov 2020
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Meloclada diuturna Melnitsky and Ivanov 2020
Meloclada frequentatoria Melnitsky and Ivanov 2020
G. †Conchindusia Vyalov and Sakacheva 1976
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Conchindusia longaeva Sukacheva 2000
Conchindusia rasnitsyni Jarzembowski 1995
G. †Coprindusia Ivanov 2006
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Coprindusia pallida Ivanov 2006
Unr. Eutrichoptera Engel 2022
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Subor. Annulipalpia Martynov 1924
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G. †Cretolype Ivanov 2006
G. †Macropteryx Hong 1976
Superfm. Philopotamoidea Stephens 1829
Superfm. Psychomyioidea Curtis 1835
G. †Qinquania Hong 1982
Invalid names: Rhyacophiloidea Stephens 1836 [empty]
Subor. Integripalpia Martynov 1924
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G. †Cathayamodus Gao et al. 2016
Fm. Glossosomatidae Wallgren 1891
Fm. Hydrobiosidae Ulmer 1905
Superfm. Hydroptiloidea Stephens 1836
Infraor. Phryganides Latreille 1805
Fm. Rhyacophilidae Stephens 1836
G. †Folindusia Berry 1927
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Subg. †Folindusia (Acrindusia) Vialov 1973
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Subg. †Folindusia (Echinindusia) Vialov and Sukatsheva 1976
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Subg. †Folindusia (Folindusia) Berry 1927
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Folindusia (Folindusia) derosa Sukatsheva 1993
Folindusia (Folindusia) laeta Sukatsheva 1993
Folindusia (Folindusia) panda Sukatsheva 1993
Subg. †Folindusia (Spirindusia) Vialov and Sukacheva 1976
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Folindusia (Spirindusia) zeuneri Vialov and Sukacheva 1976
Folindusia adumbrata Sukacheva 2000
Folindusia digna Sukacheva 2000
Folindusia manlaica Sukatsheva 1980
Folindusia megasporae Sukacheva 2000
Folindusia miocenica Berry 1928
Folindusia pilarius Sukacheva 2000
Folindusia wilcoxiana Berry 1927
Fm. †Kalophryganidae Haupt 1956
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G. †Kalophryganea Haupt 1956
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G. †Litholimnephilops Robinson et al. 2018
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Litholimnephilops yinani Robinson et al. 2018
G. †Ostracindusia Vyalov 1973
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Ostracindusia martynovae Sukacheva 2000
Ostracindusia modesta Sukacheva 1982
Ostracindusia vyalovi Jarzembowski 1995
G. †Paleodicella Sukatsheva 2014
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Paleodicella anomala Sukatsheva 2014
G. †Pelindusia Vyalov and Sakacheva 1976
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Pelindusia percealleni Jarzembowski 1995
G. †Petropteron Cockerell 1912
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Petropteron mirandum Cockerell 1912
G. †Phryganeidium Westwood 1854
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Phryganeidium pytho Westwood 1854
G. †Piscindusia Jarzembowski 1995
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Piscindusia complexa Heads 2006
Piscindusia sukachevae Jarzembowski 1995
Fm. †Prorhyacophilidae Riek 1955
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G. †Prorhyacophila Riek 1955
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Prorhyacophila batkenica Sukatsheva and Sinitshenkova 2023
Prorhyacophila furcata Sukatsheva 1973
Prorhyacophila rara Sukatsheva and Sinitshenkova 2023
Prorhyacophila rasnitsyni Sukatsheva and Aristov 2013
Subor. †Protomeropina Tillyard 1926
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G. †Adoptysma Novokshonov 1993
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Adoptysma anomalanala Novokshonov 1993
Fm. †Terminoptysmatidae Melnitsky and Ivanov 2020
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G. †Terminoptysma Melnitsky and Ivanov 2020
G. †Sanctipaulus Pinto 1956
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Sanctipaulus mendesi Pinto 1956
G. †Secrindusia Vialov and Sukacheva 1976
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Secrindusia expressa Sukacheva 2000
G. †Terrindusia Vialov 1973
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Terrindusia antiqua Heer 1849
Terrindusia buriatica Sukatsheva and Vasilenko 2019
Terrindusia fluvialis Sukatsheva 1980
Terrindusia khasurtica Sukatsheva and Vasilenko 2019
Terrindusia laxa Sukatsheva 1980
Terrindusia micacea Frič 1869
G. †Trichopterella Cockerell 1924
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Trichopterella torta Cockerell 1924
G. †Tuanwangica Zhang 1985
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Tuanwangica aethoneura Zhang 1985
G. †Zygobasis Hong 1982
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Zygobasis mesozoica Hong 1982
Invalid names: Aphelocheira Stephens 1836 [invalid subgroup], Aspatherium geinitzii Kolenati 1851 [nomen nudum], Bittacus validus Hagen 1856 [nomen dubium], Cyrnus subatomarius Kolenati 1851 [nomen nudum], Hallesus retusus Hagen 1856 [nomen dubium], Hydropsyche submaculata Kolenati 1851 [nomen nudum], Hydropsyche subvariabilis Kolenati 1851 [nomen nudum], Kalasiroptera Engel 2022 [empty], Nebritus willistoni Melander 1949 [nomen dubium], Phryganea arenacea Fritsch 1899 [nomen dubium], Phryganea kolbi Frič 1889 [nomen dubium], Phryganea solitaria Fritsch 1899 [nomen dubium], Phryganeina Sukatsheva 1968 [empty], Phryganoidea Handlirsch 1906 [synonym], Rhyacophila subumbrosa Kolenati 1851 [nomen nudum], Rhyacophila succinica Kolenati 1851 [nomen nudum], Rhyacophila succinica major Kolenati 1851 [nomen nudum], Rhyacophila succinica media Kolenati 1851 [nomen nudum], Rhyacophila succinica minor Kolenati 1851 [nomen nudum], Spicipalpia [invalid subgroup]
Or. †Tarachoptera Mey et al. 2017
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Fm. †Tarachocelidae Mey et al. 2017
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G. †Kinitocelis Mey et al. 2017
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Kinitocelis brevicostata Mey et al. 2017
Kinitocelis dashengi Wang et al. 2022
Kinitocelis divisonotata Mey et al. 2017
Kinitocelis hennigi Mey et al. 2017
Kinitocelis macroptera Mey and Wichard 2020
Kinitocelis patrickmuelleri Mey and Wichard 2023
Kinitocelis sparsella Mey et al. 2018
G. †Retortocelis Mey et al. 2018
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Retortocelis longella Mey et al. 2018
Retortocelis minimella Mey et al. 2018
Retortocelis spicipalpia Mey and Wichard 2020
Retortocelis tyloptera Mey et al. 2018
G. †Tarachocelis Mey et al. 2017
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Tarachocelis emmarossae Mey and Wichard 2023
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Westphalian C to the top of the Holocene or 314.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 309.8 Ma

Collections (470 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Moscovian315.2 - 307.0Czech Republic Phryganea kolbi, Phryganea arenacea (125662)
Westphalian C314.6 - 309.8Czech Republic Phryganea solitaria (155300)
Kungurian279.3 - 272.3Russian Federation (Komi) Adoptysma anomalanala (97621)
Wuchiapingian259.9 - 254.17Russian Federation (Vologda) Cladochorista issadica (104754)
Changhsingian254.17 - 252.17South Africa (Natal) Cladochoristidae indet. (124428)
Changhsingian254.17 - 252.17Russian Federation (Vladimir) Cladochoristella ryzhkovae, Prorhyacophila rasnitsyni (113351)
Changhsingian254.17 - 252.17Australia (New South Wales) Cladochorista belmontensis (122423 122433)
Ladinian242.0 - 237.0Brazil (Rio Grande do Sul) Sanctipaulus mendesi (116119)
Ladinian242.0 - 237.0Kyrgyzstan (Osh) Cladochorista curta, Cladochoristella sola, Prorhyacophila rara, Prorhyacophila batkenica (163987) Cladochorista multivenosa, Prophilopotamus asiaticus (106088) Prorhyacophila furcata (110765)
Longobardian242.0 - 235.0China (Shaanxi) Folindusia sp. (183209)
Carnian237.0 - 228.0China (Xinjiang) Terrindusia sp. (196151)
Norian228.0 - 208.5Australia (Queensland) Cladochoristella bryani, Prorhyacophila colliveri (106484)
Rhaetian208.5 - 201.3Germany Lepidoptera indet., Glossata indet. (191173)
Planorbis201.3 - 196.5Germany (Niedersachsen) Lepidoptera indet., Glossata indet. (191174)
Turneri196.5 - 189.6United Kingdom (England) Archaeolepis mane (114645)
Early/Lower Toarcian183.0 - 182.0Germany Liadotaulius maior (114991) Paratrichopteridium breve, Paratrichopteridium angustum, Liadotaulius acutipennis, Paratrichopteridium areatum, Orthophlebia furcata (123987)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Liadotaulius maior, Eolepidopterigidae indet., Lepidoptera indet., Micropterygidae indet. (114993)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Dolophilodes shurabica (105088)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Baga bakharica (121709)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Archiphilopotamus kubekovensis, Archiphilopotamus luxus, Archiphilopotamus absurdus, Archiphilopotamus mancus, Oncovena borealis, Juraphilopotamus funeralis, Juraphilopotamus similis (118010)