PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Chasmosaurinae

Reptilia - Ceratopsidae

Taxonomy

Chasmosaurinae was named by Lambe (1915). It is not extant. Its type is Chasmosaurus.

It was assigned to Ceratopsia by Lambe (1915); and to Ceratopsidae by Lehman (1989), Dodson and Currie (1990), Lehman (1990), Godfrey and Holmes (1995), Lehman (1996), Dodson (1997), Dodson (1997), Sereno (1997), Eberth et al. (2001), Dodson et al. (2004), Farke (2004), Ryan and Evans (2005), Ryan and Russell (2005), Wu et al. (2007), Ryan (2007), Longrich (2010), Xu et al. (2010), Maidment and Barrett (2011), Farke et al. (2011), Longrich (2013), Longrich (2014), Mallon et al. (2014), Brown and Henderson (2015), Morschhauser et al. (2019) and Fowler and Freedman Fowler (2020).

Subtaxa

Agujaceratops, Anchiceratops, Arrhinoceratops, Bisticeratops, Bravoceratops, Ceratops (syn. Proceratops), Chasmosaurus (syn. Mojoceratops, Eoceratops, Protorosaurus) (type genus), Coahuilaceratops, Judiceratops, Kosmoceratops, Navajoceratops, Pentaceratops, Sierraceratops, Spiclypeus, Terminocavus, Triceratopsini, Utahceratops, Vagaceratops (syn. Mercuriceratops)

Synonyms

Ceratops montanus was named by Marsh (1888). Its type specimen is USNM 2411, a skull, and it is a 3D body fossil. Its type locality is Ceratops type site, Cow Creek, which is in a Campanian/Campanian terrestrial horizon in the Judith River Formation of Montana. It is the type species of Ceratops, Proceratops.

It was recombined as Proceratops montanus by Lull (1906); it was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Dodson (1990) and Penkalski and Dodson (1999); it was considered a nomen dubium by Dodson et al. (2004), Ryan (2007), Ryan et al. (2010) and Mallon et al. (2016).
Ceratops was named by Marsh (1888). Its type is Ceratops montanus. It is the type genus of Ceratopsidae, Ceratopsoidea, Ceratopsinae.

It was synonymized subjectively with Monoclonius by Baur (1891); it was synonymized subjectively with Agathaumas by Lydekker (1893); it was replaced with Proceratops by Lull (1906); it was considered a nomen dubium by Dodson and Currie (1990), Dodson (1990), Lehman (1990), Dodson et al. (2004), Ryan (2007) and Mallon et al. (2016).

It was assigned to Ceratopsia by Zittel (1890); to Stegosauria by Winkler (1893); to Ceratopsia by Gadow (1898); to Ceratopidae by Mansel-Pleydell (1902); to Ceratopsia by Gadow (1897) and Lull (1924); to Ceratopsinae by Nopcsa (1928), Huene (1950) and Tatarinov (1964); and to Ceratopsidae by Marsh (1888), Marsh (1888), Marsh (1895), Depéret (1896), Marsh (1896), Nopcsa (1901), Hay (1902), Hatcher et al. (1907), Huene (1909), Lambe (1911), von Zittel (1911), Brown (1914), Matthew (1915), Gilmore (1919), Gregory and Mook (1925), Huene (1927), Hay (1930), Lull (1933), Romer (1956), Kuhn (1964), Romer (1966), Charig (1967), Swinton (1970), Powell (1973), Laub (1979), Lehman (1981) and Ostrom and Wellnhofer (1986).
Proceratops was named by Lull (1906). Its type is Ceratops montanus.

It was synonymized subjectively with Ceratops by Hay (1930), Romer (1956) and Romer (1966).

It was assigned to Agathaumidae by Lull (1906).

Synonymy list

Year	Name and author
1888	Ceratops Marsh
1888	Ceratops montanus Marsh p. 477
1888	Ceratops Marsh p. 478
1890	Ceratops Zittel p. 751
1890	Ceratops montanus Zittel p. 751
1893	Ceratops Winkler p. 105
1893	Ceratops montanus Winkler p. 105
1895	Ceratops Marsh
1896	Ceratops Depéret p. 185
1896	Ceratops montanus Hatcher p. 113
1896	Ceratops Marsh p. 243
1897	Ceratops Gadow p. 205
1898	Ceratops Gadow p. 23
1900	Ceratops montanus Walcott p. 23
1901	Ceratops Nopcsa p. 216
1901	Ceratops montanus Nopcsa p. 216
1902	Ceratops Hay p. 500
1902	Ceratops montanus Hay p. 500
1902	Ceratops Mansel-Pleydell p. lxxxiv
1902	Ceratops montanus Mansel-Pleydell p. lxxxv
1906	Proceratops Lull p. 144
1906	Proceratops montanus Lull p. 144
1907	Ceratops Hatcher et al. p. 11
1907	Ceratops montanus Hatcher et al. pp. 100, 172
1909	Ceratops Huene p. 18
1910	Ceratops monatus Lull p. 772
1911	Ceratops Lambe p. 9
1911	Ceratops von Zittel p. 296
1914	Ceratops Brown p. 377
1915	Ceratops montanus Lambe p. 10
1915	Chasmosaurinae Lambe p. 20
1915	Ceratops Matthew p. 30
1919	Ceratops Gilmore p. 64
1924	Ceratops Lull p. 245
1925	Ceratops Gregory and Mook p. 5
1927	Ceratops Huene p. 259
1927	Ceratops montanus Huene p. 261
1928	Ceratops Nopcsa p. 185
1930	Ceratops monatus Arldt p. 702
1930	Ceratops Hay p. 223
1930	Ceratops montanus Hay p. 223
1930	Ceratops montanus Russell p. 142
1931	Ceratops montanus Johnson p. 368
1933	Ceratops Lull p. 96
1933	Ceratops montanus Lull p. 96
1950	Ceratops Huene p. 351
1956	Ceratops Romer p. 639
1964	Ceratops Kuhn p. 55
1964	Ceratops montanus Kuhn p. 55
1964	Ceratops montanus Russell p. 10
1964	Ceratops Tatarinov p. 585
1964	Ceratops montanus Tatarinov p. 585
1966	Ceratops Romer p. 371
1967	Ceratops Charig p. 717
1970	Ceratops Swinton p. 254
1973	Ceratops Powell p. 118
1979	Ceratops Laub p. 4
1979	Ceratops montanus Laub p. 4
1981	Ceratops Lehman p. 207
1986	Ceratops Ostrom and Wellnhofer p. 116
1989	Chasmosaurinae Lehman p. 138
1990	Chasmosaurinae Dodson and Currie p. 612
1990	Chasmosaurinae Lehman p. 212
1995	Chasmosaurinae Godfrey and Holmes p. 727
1996	Chasmosaurinae Lehman p. 505 fig. 10
1997	Chasmosaurinae Dodson p. 12
1997	Chasmosaurinae Sereno p. 443
2001	Chasmosaurinae Eberth et al. p. 59
2001	Ceratops montanus Horner et al. p. 10
2004	Chasmosaurinae Dodson et al. p. 495
2004	Chasmosaurinae Farke p. 5
2005	Chasmosaurinae Ryan and Evans pp. 323-324
2005	Chasmosaurinae Ryan and Russell p. 1381 fig. 10
2007	Chasmosaurinae Ryan p. 391 fig. 12
2007	Chasmosaurinae Wu et al. p. 1261 fig. 14
2010	Chasmosaurinae Longrich p. 692 fig. 10
2010	Chasmosaurinae Xu et al. p. 1634 fig. 3
2011	Chasmosaurinae Farke et al. p. 699
2011	Chasmosaurinae Maidment and Barrett p. 4
2013	Chasmosaurinae Longrich
2014	Chasmosaurinae Longrich p. 303 fig. 8
2014	Chasmosaurinae Mallon et al.
2015	Chasmosaurinae Brown and Henderson
2019	Chasmosaurinae Morschhauser et al. p. 129 fig. 10
2020	Chasmosaurinae Fowler and Freedman Fowler

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()

Rank	Name	Author
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Cerapoda
—	Marginocephalia
—	Ceratopsia	()
family	Ceratopsidae
subfamily	Chasmosaurinae
subfamily	Chasmosaurinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Chasmosaurinae Lambe 1915

show all | hide all

G. †Agujaceratops Lucas et al. 2006 [Aguja horned face]

†Agujaceratops mariscalensis Lehman 1989

†Agujaceratops mavericus Lehman et al. 2017

G. †Anchiceratops Brown 1914

†Anchiceratops ornatus Brown 1914

Invalid names: Anchiceratops longirostris Sternberg 1926 [synonym]

G. †Arrhinoceratops Parks 1925

†Arrhinoceratops brachyops Parks 1925

G. †Bisticeratops Dalman et al. 2022

†Bisticeratops froeseorum Dalman et al. 2022

G. †Bravoceratops Wick and Lehman 2013

†Bravoceratops polyphemus Wick and Lehman 2013

G. †Chasmosaurus Lambe 1914

†Chasmosaurus belli Lambe 1902

†Chasmosaurus russelli Sternberg 1940

Invalid names: Mojoceratops perifania Longrich 2010 [synonym]

Invalid names: Chasmosaurus brevirostris Lull 1933 [nomen dubium], Chasmosaurus kaiseni Brown 1933 [nomen dubium], Eoceratops Lambe 1915 [synonym], Mojoceratops Longrich 2010 [synonym], Monoclonius canadensis Lambe 1902 [nomen dubium], Protorosaurus Lambe 1914 [synonym]

G. †Coahuilaceratops Loewen et al. 2010

†Coahuilaceratops magnacuerna Loewen et al. 2010

G. †Judiceratops Longrich 2013

†Judiceratops tigris Longrich 2013

G. †Kosmoceratops Sampson et al. 2010

†Kosmoceratops richardsoni Sampson et al. 2010

G. †Navajoceratops Fowler and Freedman Fowler 2020

†Navajoceratops sullivani Fowler and Freedman Fowler 2020

G. †Pentaceratops Osborn 1923

†Pentaceratops sternbergii Osborn 1923

Invalid names: Pentaceratops fenestratus Wiman 1930 [synonym]

G. †Sierraceratops Dalman et al. 2022

†Sierraceratops turneri Dalman et al. 2022

G. †Spiclypeus Mallon et al. 2016

†Spiclypeus shipporum Mallon et al. 2016

G. †Terminocavus Fowler and Freedman Fowler 2020

†Terminocavus sealeyi Fowler and Freedman Fowler 2020

Unr. †Triceratopsini Longrich 2011

G. †Eotriceratops Wu et al. 2007

†Eotriceratops xerinsularis Wu et al. 2007

G. †Regaliceratops Brown and Henderson 2015

†Regaliceratops peterhewsi Brown and Henderson 2015

G. †Titanoceratops Longrich 2011

†Titanoceratops ouranos Longrich 2011

G. †Torosaurus Marsh 1891

†Torosaurus latus Marsh 1891

Invalid names: Torosaurus gladius Marsh 1891 [objective synonym]

†Torosaurus utahensis Gilmore 1946

G. †Triceratops Marsh 1889

†Triceratops horridus Marsh 1889

Invalid names: Nedoceratops hatcheri Lull 1905 [synonym], Triceratops albertensis Sternberg 1949 [synonym], Triceratops brevicornus Hatcher and Lull 1905 [synonym], Triceratops calicornis Marsh 1898 [synonym], Triceratops elatus Marsh 1891 [synonym], Triceratops eurycephalus Schlaikjer 1935 [synonym], Triceratops flabellatus Marsh 1889 [synonym], Triceratops obtusus Marsh 1898 [synonym], Triceratops serratus Marsh 1890 [synonym]

†Triceratops prorsus Marsh 1890

Invalid names: Nedoceratops Ukrainsky 2007 [synonym], Ojoceratops Sullivan and Lucas 2010 [synonym], Sterrholophus Marsh 1891 [synonym], Tatankaceratops Ott and Larson 2010 [synonym], Triceratops sulcatus Marsh 1890 [nomen dubium]

G. †Utahceratops Sampson et al. 2010

†Utahceratops gettyi Sampson et al. 2010

G. †Vagaceratops Sampson 2010

†Vagaceratops irvinensis Holmes et al. 2001

Invalid names: Mercuriceratops gemini Ryan et al. 2014 [synonym], Pentaceratops aquilonius Longrich 2014 [synonym]

Invalid names: Mercuriceratops Ryan et al. 2014 [synonym]

Invalid names: Ceratops Marsh 1888 [nomen dubium]

Diagnosis

Reference	Diagnosis
S. C. R. Maidment and P. M. Barrett 2011	Modified from Lehman (1989): large ceratopsian dinosaurs with long, low facial region (preorbital length/height ratio = 1.4 to 3.0); inter-premaxillary fossae present; premaxilla and predentary with horizontal or poorly developed lateral cutting flange; nasal horn core, if present, small and formed in part by a separate ossification (epinasal); postfrontal foramen present; frontoparietal fontanelle walled mostly by the postorbital bones; frontal bones reduced; prefrontal bones small; cranial frill long (0.94 to 1.70 basal length of skull, except in Triceratops); triangular squamosal bones (length/height ratio = 2.0 to 3.5); prominent longitudinal channel in ventral surface of sacrum; triangular acromial process of the scapula arises about half way along the scapula blade; deep triceps fossa on the caudolateral deltopectoral crest of the humerus; foramen on the caudomedial humerus located dorsal to the apex of the deltopectoral crest; strongly curved ischium.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Maximum body mass:

10000 kgg

Minimum body mass:

1000 kgg

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialsubo

Locomotion:

actively mobileo

Life habit:

ground dwellingsubo

Diet:

herbivoresubo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2005-09-15 11:08:09

Modified:

2005-09-15 13:08:09

Source: g = genus, subo = suborder, o = order

References: Marsh 1875, Peczkis 1995, Norman and Weishampel 1991

Age range: base of the Judithian to the top of the Lancian or 83.50000 to 66.00000 Ma

Collections (301 total)

Time interval	Ma	Country or state	Original ID and collection number
Campanian	83.6 - 72.1	Mexico (Chihuahua)	Chasmosaurinae indet. (226324)
Campanian	83.6 - 72.1	USA (Wyoming)	Ceratopsia indet. (45861)
Campanian	83.6 - 72.1	Canada (Alberta)	Chasmosaurus russelli (64401)
Judithian	83.5 - 70.6	USA (Texas)	Chasmosaurus mariscalensis (48941 52462 52463 52464)
Late/Upper Campanian	83.5 - 70.6	USA (Utah)	Ceratopsia indet. (97805 97810) Chasmosaurinae indet. (69799 97800 97801 97802 97806 97808 97809 97811 131868) Kosmoceratops richardsoni (97804)
Late/Upper Campanian	83.5 - 70.6	Mexico (Coahuila)	Ceratopsidae indet. (24258)
Late/Upper Campanian	83.5 - 70.6	Mexico (Chihuahua)	Chasmosaurus mariscalensis (227230)
Late/Upper Campanian	83.5 - 70.6	USA (Wyoming)	Ceratopsia indet. (45862)
Late/Upper Campanian	83.5 - 70.6	USA (Montana)	Mercuriceratops gemini (156538) Spiclypeus shipporum (55274)
Late/Upper Campanian	83.5 - 70.6	USA (New Mexico)	Ceratops sp. (62549 196540) Ceratopsidae indet. (53982) Chasmosaurinae indet. (138832 210631) Navajoceratops sullivani (138830) Pentaceratops fenestratus (39231) Pentaceratops sp. (46991 64094 64096) Pentaceratops sternbergi (52058) Pentaceratops sternbergii (34826 34827 34828 39229 196511 199764 222481) Torosaurus sp., Chasmosaurinae indet. (227186)
Late/Upper Campanian	83.5 - 70.6	Canada (Alberta)	Anchiceratops sp. (51317) Arrhinoceratops sp. (152403) Chasmosaurus belli (45730 45731 58909 58954 59047 191258) Chasmosaurus brevirostris (58912) Chasmosaurus irvinensis (86515) Chasmosaurus kaiseni (35379) Chasmosaurus russelli (64400) Chasmosaurus sp. (22656 22659 24891 24893 59006 59014 59136 191255) Eoceratops canadensis (59026) Mercuriceratops gemini (156536) Monoclonius belli (60873) Triceratops horridus (162141)
Late/Upper Campanian	83.5 - 70.6	USA (Texas)	Agujaceratops mariscalensis (236770 236774) Agujaceratops sp. (94373 236771) Pentaceratops sp. (191054)
Middle Campanian	83.5 - 70.6	USA (Montana)	Judiceratops tigris (26779)
Middle Campanian	83.5 - 70.6	USA (Texas)	Agujaceratops mariscalensis (236772 236773 236775)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Canada (Saskatchewan)	Chasmosaurinae indet. (75030)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Canada (Alberta)	Anchiceratops sp. (51318) Chasmosaurus russelli (64402) Monoclonius canadensis (60874)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	USA (Montana)	Ceratops montanus (46123) Chasmosaurus sp. (50907)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	USA (Texas)	Agujaceratops mariscalensis (236769) Chasmosaurus mariscalensis (22709 25073)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mexico (Chihuahua)	Chasmosaurus mariscalensis (35353)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	Canada (Alberta)	Anchiceratops longirostris (28548) Anchiceratops ornatus (191402) Anchiceratops sp. (52072) Arrhinoceratops brachyops (67612) Ceratopsia indet. (64629)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	Mexico (Coahuila)	Chasmosaurinae indet. (226295) Chasmosaurus sp. (91053)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	USA (Colorado)	Chasmosaurinae indet. (76715)
Late/Upper Campanian - Maastrichtian	83.5 - 66.0	USA (New Mexico)	Torosaurus sp. (178745) Triceratops sp. (55286)
Maastrichtian	72.1 - 66.0	USA (South Dakota)	Torosaurus latus (53128) Triceratops horridus (225317 225924) Triceratops sp. (74175)
Maastrichtian	72.1 - 66.0	USA (North Dakota)	Torosaurus latus (45103)
Maastrichtian	72.1 - 66.0	USA (Montana)	Triceratops elatus (49016) Triceratops prorsus (173841) Triceratops sp. (38544 55353 55355 64428 64429 100852 109660 109661 142376 189572 228491)
Maastrichtian	72.1 - 66.0	USA (Wyoming)	Triceratops sp. (225318 225319)
Maastrichtian	72.1 - 66.0	USA (Texas)	Bravoceratops polyphemus (145482) Torosaurus sp. (52056) Torosaurus utahensis (227185)
Early/Lower Maastrichtian	70.6 - 66.0	Canada (Alberta)	Anchiceratops longirostris (97797) Anchiceratops ornatus (64630 76062 191401) Arrhinoceratops brachyops (156799) Regaliceratops peterhewsi (191308)
Early/Lower Maastrichtian	70.6 - 66.0	Mexico (Coahuila)	Coahuilaceratops magnacuerna (97757)
Lancian	70.6 - 66.0	USA (Montana)	Triceratops sp. (14516 14550 14615 14616 14649 81041 81042 144502 179991)
Lancian	70.6 - 66.0	USA (Colorado)	Triceratops sp. (14640)
Lancian	70.6 - 66.0	USA (South Dakota)	Triceratops sp. (14537 192251)
Lancian	70.6 - 66.0	Canada (Saskatchewan)	Triceratops sp. (14558 128461)
Lancian	70.6 - 66.0	USA (Wyoming)	Ceratopsia indet. (165877) Chasmosaurinae indet. (14540 14541 14542 14543 14544 70341 70343 75280 75281 75284 75287 75289 75290 75292 75294 75297 75298 75429 75431 75432 75439 75441 75442 75443 75447) Triceratops sp. (14505 14527 14538 14557 14579 14601 14609 165878)
Lancian	70.6 - 66.0	USA (North Dakota)	Triceratops sp. (137710 137728 224667)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Texas)	Pentaceratops sp. (85188) Torosaurus utahensis (69211)
Late/Upper Maastrichtian	70.6 - 66.0	Canada (Alberta)	Eotriceratops xerinsularis (75475) Torosaurus sp. (47130) Triceratops albertensis (47127 47128)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Montana)	Triceratops horridus (217183) Triceratops prorsus (14674 173842) Triceratops sp. (13142 55354 67129 72143 88990 99441 109659 142326 157187 162855 179968 183109 183110 189003 221004 221005 221007 227182 227283)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Utah)	Ceratopsia indet. (45719) Triceratops sp. (48549)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Wyoming)	Ceratops horridus (46119) Ceratopsia indet. (52089) Diceratops hatcheri (46133) Torosaurus gladius (46135) Torosaurus latus (46134) Triceratops brevicornus (46132 52683) Triceratops calicornis (50931) Triceratops elatus (46130 52682 52691 52692) Triceratops eurycephalus (52107) Triceratops flabellatus (46136 165876) Triceratops obtusus (46131) Triceratops prorsus (46127 46129 52690 52694 189513) Triceratops serratus (46126) Triceratops serratus, Ceratopsia indet. (52686) Triceratops sp. (14585 26760 45878 50086 50087 52088 52681 52684 52687 52688 52693 52695 54177 54179 57161 57162 57166 57168 142325 164991 164992 164993 191271 226054 226055) Triceratops sp., Triceratops sulcatus (52685) Triceratops sulcatus (46138 52689)
Late/Upper Maastrichtian	70.6 - 66.0	USA (North Dakota)	Triceratops horridus (191374) Triceratops prorsus (24851) Triceratops sp. (45123 57919)
Late/Upper Maastrichtian	70.6 - 66.0	USA (New Mexico)	Chasmosaurinae indet. (68042) Ojoceratops fowleri (47819 97795 97796 138833 171027)
Late/Upper Maastrichtian	70.6 - 66.0	Canada (Saskatchewan)	Ceratopsidae indet. (152402) Chasmosaurinae indet. (52837) Triceratops prorsus (152400 152401 228825 228826 228827) Triceratops sp. (14555 52836 54105 152399 225845) Triceratops sp., Triceratops prorsus (45675)
Late/Upper Maastrichtian	70.6 - 66.0	USA (South Dakota)	Tatankaceratops sacrisonorum (97798) Torosaurus sp., Triceratops sp. (224484) Triceratops horridus (179969 224486) Triceratops sp. (50082 124934 150890 150892 150893 179970 216603 224488 224489 224490 224492)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Colorado)	Ceratopsia indet. (49527) Triceratops sp. (49529 49536 73927 182584)