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Helminthomorpha

Myriapoda

Synonyms
Synonymy list
YearName and author
1873Archiulidae Scudder
1887Helminthomorpha Pocock
1890Archiulidae Scudder p. 435
1930Archiulidae Pruvost p. 169
1934Archiulidae Guthörl
1969Helminthomorpha Hoffman p. 586
1975Helminthomorpha Dzik p. 17
2008Helminthomorpha Štamberg and Zajíc p. 81
2010Helminthomorpha Shear and Edgecombe p. 181
2014Helminthomorpha Riquelme et al. p. 419
2014Helminthomorpha Riquelme et al. p. 5
2018Helminthomorpha Riquelme and Hernández-Patricio p. 638
2018Helminthomorpha Ross et al. p. 101

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
RankNameAuthor
classMyriapoda()
—DignathaTiegs 1947
subclassDiplopodaGervais 1844
subclassChilognatha()
infraclassHelminthomorpha(Pocock 1887)
infraclassHelminthomorpha(Pocock 1887)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infracl. Helminthomorpha Pocock 1887
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G. †Archicambala Cook 1895
+
†Archicambala dawsoni Scudder 1868
Superor. †Archipolypoda Scudder 1882
+
G. †Albadesmus Wilson and Anderson 2004
+
†Albadesmus almondi Wilson and Anderson 2004
G. †Anaxeodesmus Wilson 2005
+
†Anaxeodesmus diambonotus Wilson 2005
G. †Anthracodesmus Peach 1899
+
Or. †Archidesmida Wilson and Anderson 2004
+
Fm. †Archidesmidae Scudder 1885
+
G. †Archidesmus Peach 1882
+
†Archidesmus macnicoli Peach 1882
Fm. †Zanclodesmidae Wilson et al. 2005
+
G. †Orsadesmus Wilson et al. 2005
+
†Orsadesmus rubecollus Wilson et al. 2005
G. †Zanclodesmus Wilson et al. 2005
+
†Zanclodesmus willetti Wilson et al. 2005
Or. †Cowiedesmida Wilson and Anderson 2004
+
Fm. †Cowiedesmidae Wilson and Anderson 2004
+
G. †Cowiedesmus Wilson and Anderson 2004
+
†Cowiedesmus eroticopodus Wilson and Anderson 2004
Or. †Euphoberiida Hoffman 1969
+
Fm. †Euphoberiidae Scudder 1882
+
G. †Acantherpestes Meek and Worthen 1868
+
†Acantherpestes foveolatus Fritsch 1899
†Acantherpestes gigas Fritsch 1895
†Acantherpestes inequalis Scudder 1890
†Acantherpestes major Meek and Worthen 1868
†Acantherpestes ornatus Fritsch 1899
†Acantherpestes vicinus Fritsch 1899
G. †Chonionotus Jordan 1854
G. †Euphoberia Meek and Worthen 1868
+
†Euphoberia absens Fritsch 1899
†Euphoberia anguilla Scudder 1882
†Euphoberia armigera Meek and Worthen 1868
†Euphoberia brownii Woodward 1871
†Euphoberia carri Scudder 1882
†Euphoberia cuspidata Scudder 1890
†Euphoberia ferox Salter 1863
+
Invalid names: Acantherpestes brodiei Scudder 1882 [synonym]
†Euphoberia flabellata Scudder 1882
†Euphoberia granosa Scudder 1882
†Euphoberia histrix Fritsch 1899
†Euphoberia horrida Scudder 1882
†Euphoberia hystricosa Scudder 1890
†Euphoberia lithanthracis Jordan 1854
†Euphoberia simplex Scudder 1890
†Euphoberia spinulosa Scudder 1890
†Euphoberia tracta Scudder 1890
†Euphoberia varians Fritsch 1899
G. †Myriacantherpestes Burke 1979
G. †Kampecaris Page 1856
+
†Kampecaris dinmorensis Clarke 1951
†Kampecaris forfarensis Peach 1882
G. †Lauravolsella Haug et al. 2023
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†Lauravolsella willemeni Haug et al. 2023
G. †Palaeodesmus Wilson and Anderson 2004
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†Palaeodesmus tuberculata Brade-Birks 1923
Or. †Palaeosomatida Hannibal and Krzemiński 2005
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Fm. †Palaeosomatidae Hannibal and Krzemiński 2005
+
G. †Palaeosoma Jackson et al. 1919
+
†Palaeosoma giganteum Baldwin 1911
†Palaeosoma robustum Baldwin 1911
+
Invalid names: Euphoberia woodwardi Baldwin 1911 [synonym]
Or. †Pleurojulida Wilson and Hannibal 2005
+
Fm. †Pleurojulidae Schneider and Werneburg 1998
+
G. †Isojulus Fritsch 1899
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†Isojulus constans Frič 1875
+
Invalid names: Isojulus marginatus Fritsch 1899 [synonym], Isojulus setipes Fritsch 1899 [synonym], Pleurojulus falcifer Fritsch 1899 [synonym], Pleurojulus longipes Fritsch 1899 [synonym]
G. †Pleurojulus Fritsch 1899
+
†Pleurojulus biornatus Fritsch 1899
†Pleurojulus levis Fritsch 1899
+
Invalid names: Pleurojulus aculeatus Fritsch 1899 [synonym], Pleurojulus pinguis Fritsch 1899 [synonym]
G. †Pneumodesmus Wilson and Anderson 2004
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†Pneumodesmus newmani Wilson and Anderson 2004
Fm. †Woodesmidae Ross et al. 2018
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G. †Woodesmus Ross et al. 2018
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†Woodesmus sheari Ross et al. 2018
Invalid names: Euphoberidae Scudder 1882 [empty]
G. †Archiulus Scudder 1868
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†Archiulus brassi Dohrn 1868
†Archiulus euphoberioides Scudder 1895
†Archiulus glomeratus Scudder 1890
†Archiulus lyelli Scudder 1895
†Archiulus xylobioides Scudder 1868
Unr. Colobognatha Brandt 1834
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Or. Platydesmida de Saussure 1860
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Fm. Andrognathidae Cope 1869
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G. Andrognathus Cope 1869
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†Andrognathus burmiticus Moritz and Wesener 2019
Or. Polyzoniida Cook 1895
+
Fm. Polyzoniidae Gervais 1844
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G. Polyzonium Brandt 1837
Fm. Siphonotidae Cook 1895
+
G. Siphonotus Brandt 1837
Or. Siphonophorida Newport 1844
+
Fm. Siphonophoridae Newport 1844
+
G. Siphonocybe Pocock 1903
G. Siphonophora Brandt 1837
+
†Siphonophora hoffmani Santiago-Blay and Poinar 1992
†Siphonophora hui Jiang et al. 2019
†Siphonophora velezi Santiago-Blay and Poinar 1992
Fm. Siphonorhinidae Cook 1895
+
G. Siphonorhinus Pocock 1894
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†Siphonorhinus globosus Su et al. 2024
†Siphonorhinus peculiaris Su et al. 2024
G. †Dolesea Hannibal and May 2020
+
†Dolesea subtila Hannibal and May 2020
Unr. Eugnatha Attems 1898
+
Superor. Juliformia Attems 1926
+
Or. Julida Leach 1814
+
Superfm. Juloidea Leach 1814
+
Fm. Julidae Leach 1814
Fm. Parajulidae Bollman 1893
Fm. Nemasomatidae Bollman 1893
+
G. Blaniulus Gervais 1836
Fm. †Proglomeridae Fritsch 1899
+
G. †Archiscudderia Fritsch 1899
+
†Archiscudderia coronata Fritsch 1899
†Archiscudderia paupera Fritsch 1899
†Archiscudderia regularis Fritsch 1899
†Archiscudderia tapeta Fritsch 1899
Or. Spirobolida Cook 1895
+
Fm. †Anthracojulidae Hoffman 1969
+
G. †Anthracojulus Fritsch 1899
G. †Gobiulus Dzik 1975
+
†Gobiulus sabulosus Dzik 1975
Fm. Rhinocricidae Brölemann 1913
+
G. Rhinocricus Karsch 1881
Fm. Spirobolidae Bollman 1893
+
G. Hiltonius Chamberlin 1918
G. Spirobolus Brandt 1833
Invalid names: Atopetholidae Chamberlin 1918 [empty]
Or. Spirostreptida Cook 1895
+
Subor. Cambalidea Bollman 1893
+
Fm. Cambalidae Bollman 1893
Fm. Pseudonannolenidae Silvestri 1895
+
G. Epinannolene Broleman 1903
Superfm. †Xyloiuloidea Cook 1895
+
Fm. †Gaspestriidae Wilson 2006
+
G. †Gaspestria Wilson 2006
G. †Sigmastria Wilson 2006
G. †Karstiulus Hannibal and May 2020
+
†Karstiulus fortsillensis Hannibal and May 2020
Fm. †Nyraniidae Hoffman 1969
+
G. †Nyranius Hoffman 1963
Fm. †Plagiascetidae Hoffman 1969
+
G. †Plagiascetus Hoffman 1963
Fm. †Xyloiulidae Cook 1895
+
G. †Blanziulus Langiaux and Sotty 1976
G. †Xyloiulus Cook 1895
+
Invalid names: Xylobius Dawson 1860 [replaced]
Invalid names: Projulidae Fritsch 1899 [synonym]
Superor. Merochaeta Cook 1895
+
Or. Polydesmida Pocock 1887
+
Subor. Leptodesmidea Brolemann 1916
+
Superfm. Chelodesmoidea Cook 1895
Superfm. Platyrhacoidea Pocock 1895
Superfm. Xystodesmoidea Cook 1895
Fm. Polydesmidae Leach 1815
+
G. Polydesmus Latreille 1802
G. Propolydesmus Verhoeff 1895
Subor. Polydesmidea Pocock 1887
+
G. Dasyodontus Loomis 1936
G. Inodesmus Cook 1896
Superfm. Pyrgodesmoidea Silvestri 1896
Superfm. Trichopolydesmoidea Verhoeff 1910
Fm. Sphaeriodesmidae Humbert and de Saussure 1869
+
G. Cyclodesmus Humbert and de Saussure 1869
Invalid names: Chelodesmidea Cook 1895 [empty], Chytodesmidae Cook 1896 [empty]
Superor. Nematophora Verhoeff 1913
+
Or. Callipodida Pocock 1894
+
Subor. †Burmanopetalidea Stoev et al. 2019
+
Fm. †Burmanopetalidae Stoev et al. 2019
G. †Hannibaliulus Shear et al. 2009
+
†Hannibaliulus wilsonae Shear et al. 2009
Invalid names: Dorypetalidae Verh 1909 [empty], Lysiopetaloidea Verh 1900 [synonym]
Or. Stemmiulida Cook 1895
+
Fm. Stemmiulidae Pocock 1894
+
G. †Parastemmiulus Riquelme et al. 2014
Fm. †Hexecontasomatidae Hannibal 2000
+
G. †Hexecontasoma Hannibal 2000
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†Hexecontasoma carinatum Hannibal 2000
Fm. †Oklahomasomatidae Hannibal and May 2020
+
G. †Oklahomasoma Hannibal and May 2020
+
†Oklahomasoma richardsspurense Hannibal and May 2020
G. †Sinosoma Huang and Hannibal 2018
+
†Sinosoma luopingense Huang and Hannibal 2018
Or. Siphoniulida Pocock 1894
+
Fm. Siphoniulidae Pocock 1894
+
G. Siphoniulus Pocock 1894
+
†Siphoniulus muelleri Liu et al. 2017
†Siphoniulus preciosus Liu et al. 2017
Invalid names: Archiulidae Scudder 1873 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Locomotion: actively mobilep
Created: 2004-02-29 09:38:49
Modified: 2004-02-29 11:38:49
Source: p = phylum
Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Homerian to the top of the Rancholabrean or 430.50000 to 0.01200 Ma
Minimum age of oldest fossil (stem group age): 427.4 Ma

Collections (110 total)


Time interval Ma Country or state Original ID and collection number
Wenlock433.4 - 427.4United Kingdom (Scotland) Cowiedesmus eroticopodus, Albadesmus almondi, Pneumodesmus newmani (26907)
Homerian430.5 - 427.4United Kingdom (Scotland) Albadesmus almondi (155916) Cowiedesmus eroticopodus, Albadesmus almondi (155913) Diplopoda indet. (155915) Pneumodesmus newmani (84333)
Pridoli423.0 - 419.2United Kingdom (England) Kampecaris dimnorensis (11128)
Lochkovian419.2 - 410.8United Kingdom (Scotland) Archidesmus macnicoli (155912 155918) Archidesmus macnicoli, Sigmastria dilata (155933) Kampecaris forfarensis (195121 195122 195123 195127 195128)
Lochkovian - Pragian419.2 - 407.6United Kingdom (England) Kampecaris dinmorensis (233872)
Emsian407.6 - 393.3United Kingdom (Scotland) Kampecaris tuberculata (120607)
Late/Upper Emsian402.5 - 391.9Canada (New Brunswick) Gaspestria genselorum (155932)
Late/Upper Emsian402.5 - 391.9Canada (Quebec) Gaspestria genselorum (155931)
Frasnian382.7 - 372.2Canada (Quebec) Zanclodesmus willetti (155925)
Late/Upper Famennian364.7 - 360.7USA (Pennsylvania) Orsadesmus rubecollus (35044)
Tournaisian358.9 - 346.7United Kingdom (Scotland) Anthracodesmus macconochiei (127550) Archidesmida indet. 1 (195789) Archipolypoda indet. 2 (195401) Archipolypoda indet. 3 (194781) Juliformia indet. 4, Woodesmus sheari (183472) Woodesmus sheari (195790)
Brigantian336.0 - 330.9Germany Euphoberia sp. (153594)
Serpukhovian330.9 - 323.2Poland Palaeosoma robustum (195184)
Westphalian A318.7 - 316.9United Kingdom (England) Acantherpestes giganteus, Euphoberia robusta, Euphoberia armigera (127287) Euphoberia woodwardi (96873) Eurypterus ferox (155939) Xylobius platti (34905) Xyloiulus sp. (124367)
Westphalian A318.7 - 316.9Canada (Nova Scotia) Xylobius sigillariae, Archiulus xylobioides (155665) Xylobius sigillariae, Archiulus xylobioides, Archiulus lyelli, Xylobius similis, Xylobius fractus, Xylobius dawsoni, Archiulus euphoberioides (28069)
Westphalian A318.7 - 316.9Belgium Acantherpestes sp., Xylobius renieri (155630)
Westphalian A318.7 - 316.9Netherlands (Limburg) Lauravolsella willemeni (110966)
Westphalian B316.9 - 314.6United Kingdom (Staffordshire) Anaxeodesmus diambonotus (153546)
Westphalian B316.9 - 314.6United Kingdom (England) Euphoberia ferox, Myriacantherpestes sp. (125238) Lepidoptera caterpillar indet. (155938)
Westphalian B316.9 - 314.6Belgium Euphoberia ferox (155631)
Westphalian B316.9 - 314.6United Kingdom Euphoberia brownii (195182) Euphoberia ferox (195183)
Moscovian315.2 - 307.0Czech Republic (Pilsen) Euphoberia histrix, Euphoberia varians, Euphoberia absens, Acantherpestes gigas, Acantherpestes vicinus, Acantherpestes ornatus, Acantherpestes foveolatus, Julus constans, Isojulus setipes, Isojulus marginatus, Pleurojulus biornatus, Pleurojulus levis, Pleurojulus aculeatus, Pleurojulus longipes, Pleurojulus falcifer, Pleurojulus pinguis, Xylobius pstrossi, Julus costulatus, Xylobius sellatus, Xylobius tabulatus, Archiscudderia paupera, Archiscudderia tapeta, Archiscudderia coronata, Archiscudderia regularis, Archiscudderia problematica (128574)
Westphalian C314.6 - 309.8Germany (Saarland) Chonionotus lithanthracis (122623)
Westphalian C - Westphalian D314.6 - 307.5Canada (Nova Scotia) Xylobius sp. (80111)
Westphalian D309.8 - 307.5USA (Illinois) Euphoberia armigera (155944) Euphoberia armigera, Acantherpestes major, Euphoberia carri (122528) Euphoberia armigera, Acantherpestes major, Euphoberia flabellata (154202) Euphoberia armigera, Euphoberia anguilla (155943) Euphoberia armigera, Euphoberia horrida, Euphoberia granosa, Euphoberia carri (155941) Euphoberia armigera, Euphoberia major, Euphoberia granosa (152106) Euphoberia armigera, Xylobius frustulentus, Acantherpestes major, Euphoberia granosa, Euphoberia spinulosa, Euphoberia simplex, Euphoberia tracta (122521) Euphoberia armigera, Xylobius frustulentus, Acantherpestes major, Euphoberia horrida, Euphoberia granosa, Euphoberia carri, Euphoberia anguilla, Archiulus glomeratus (122527) Euphoberia sp., Pleurojulus biornatus, Euphoberia armigera, Xylobius mazonus, Xylobius frustulentus, Acantherpestes major, Euphoberia granosa, Euphoberia anguilla, Acantherpestes inequalis, Euphoberia hystricosa, Euphoberia cuspidata, Euphoberia spinulosa, Euphoberia simplex, Euphoberia tracta, Archiulus glomeratus (21981) Hexecontasoma carinatum (122493 132455) Pleurojulus biornatus (112630 112638 133734)
Westphalian D309.8 - 307.5USA (Ohio) Plagiascetus lateralis, Xylobius sp. (85292)
Westphalian D309.8 - 307.5United Kingdom (Wales) Euphoberia sp., Pleurojulus levis (127543)
Stephanian B301.5 - 301.0France Blanziulus parriati (122490 161212)
Stephanian B301.5 - 301.0Czech Republic (Rakovník) Julus pictus (154179)
Stephanian B301.5 - 301.0Czech Republic Anthracojulus pictus (155305)
Asselian298.9 - 295.5USA (New Mexico) Spirobolida indet. (124386)
Asselian - Sakmarian298.9 - 290.1Germany (Rhineland-Palatinate) Julus brassii (122606)
Artinskian290.1 - 279.3USA (Texas) Xylobius sp. (11149)
Artinskian290.1 - 279.3USA (Oklahoma) Helminthomorpha indet., Karstiulus fortsillensis, Oklahomasoma richardsspurense, Dolesea subtila (67862)
Kungurian279.3 - 272.3USA (Texas) Xylobius sp. (195185)
Aegean247.2 - 242.0France (Moselle) Hannibaliulus wilsonae (161211)
Pelsonian247.2 - 242.0China (Yunnan) Sinosoma luopingense (127488)
Early/Lower Cenomanian99.6 - 93.5Myanmar Polydesmida indet., Siphonophoridae indet., Polyzoniida indet. (196547) Polydesmida indet., Siphonophoridae indet., Polyzoniida indet., Andrognathidae indet. (144466) Siphonophora hui (199763) Siphonophora hui, Monstrodesmus grimaldii, Propolydesmus cretaceus, Siphonorhinus globosus, Siphonorhinus peculiaris (161726) Stemmiulida indet., Cambalidea indet., Cambalidae indet., Polydesmida indet., Siphonotidae indet., Siphoniulus muelleri, Siphoniulus preciosus, Andrognathus burmiticus, Callipodida indet. (184444) Stemmiulida indet., Cambalidea indet., Polydesmida indet., Siphonophorida indet., Siphonophoridae indet., Polyzoniida indet., Siphonotidae indet., Siphonorhinidae indet., Andrognathidae indet. (184692)
Middle Campanian - Late/Upper Campanian83.5 - 70.6Mongolia (Omnogov) Diplopoda indet. (13776 28243 37022)
Bridgerian50.3 - 46.2USA (Wyoming) Iulus telluster (121306)
Priabonian38.0 - 33.9Poland Julus laevigatus (109625) Julus politus, Julus badius, Julus rubens (120811)
Chadronian37.2 - 33.9USA (Colorado) Parajulus cockerelli (123119)
MP 3028.4 - 23.03Germany (Nordrhein-Westfalen) Julus antiquus (127628)
Chattian28.1 - 23.03France Protosilvestria sculpta (160861 160862)
Aquitanian - Burdigalian23.03 - 15.97Australia (Queensland) Julida indet. (106497)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Anbarrhacus adamantis (217222) Maatidesmus paachtun (217221) Myrmecodesmus sp. (217216) Parastemmiulus elektron (157206) Polydesmida indet. (198592 198593 217215) Siphonophora sp. (217217) Spirobolida indet., Juliformia indet., Stemmiulidae indet., Polydesmida indet., Xystodesmidae indet. (172774) Stemmiulidae indet., Polydesmida indet., Platyrhacidae indet., Siphonophora sp., Xystodesmidae indet. (217220)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Docodesmus sp., Siphonophora sp., Lophodesmus sp., Inodesmus sp., Siphonophora hoffmani, Siphonophora velezi (159268) Siphonocybe sp., Docodesmus brodzinskyi (122473) Stemmiulidae indet., Chelodesmidae indet., Docodesmus sp., Epinannolene sp., Siphonophora sp., Siphonotus sp., Iomus sp., Psochodesmus sp., Dasyodontus sp., Inodesmus sp., Siphonophora hoffmani (122472)
Sarmatian12.7 - 11.608Germany (Wurttemberg) Julidae indet., Julus antiquus (113752)
Pliocene5.333 - 2.588USA (Arizona) Parajulus onychis (152091)
Early/Lower Pleistocene2.588 - 0.781Hungary (Barany) Julus sp. (35459)
Rancholabrean0.3 - 0.012USA (California) Spirobolus australis (150336)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Julus occidentalis (74235)
Late/Upper Pleistocene (glacial)0.129 - 0.0117USA (California) Julus cavicola (155393)
Late/Upper Pleistocene0.129 - 0.0117Jamaica Rhinocricus spp., Chondrotropis sp., Cyclodesmus porcellanus, Caraibodesmus verrucosus (224573)