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Pelomedusoides
Taxonomy
Pelomedusoides was named by Broin (1988) [incorrectly credited by Leckey, based on Hutchison et al., to E. D. Cope 1865]. It was considered monophyletic by Pérez-García (2017).
It was reranked as the unranked clade Pelomedusoides by Gaffney et al. (2006), Cadena et al. (2012), Weems and Knight (2013), Cadena (2020), Antunes and Pérez-García (2024).
It was assigned to Eupleurodira by de la Fuente and Iturralde-Vinent (2001), Gaffney et al. (2006); to Panpelomedusoides by Cadena et al. (2012); to Pleurodira by Pérez-García et al. (2013), Pérez-García and Lapparent de Broin (2013); to Testudines by Menegazzo et al. (2015); to Pleurodira by Cadena et al. (2012), Pérez-García et al. (2013), Pérez-Garcia (2020), Georgalis et al. (2020), Cadena (2020), Maniel et al. (2021), Antunes and Pérez-García (2024); and to Pleurodira by Joyce et al. (2004), Hutchison et al. (2004), Oliveira and Kellner (2007), Weems and Knight (2013), Ferreira et al. (2015), de Araujo Carvalho et al. (2016), Pérez-García and Chapman (2017), Pérez-García (2017), Pérez-García and Ortega (2018), Ferreira et al. (2018), Pérez-García (2020), de Lapparent de Broin et al. (2020), López-Conde et al. (2021), Cadena et al. (2025).
It was reranked as the unranked clade Pelomedusoides by Gaffney et al. (2006), Cadena et al. (2012), Weems and Knight (2013), Cadena (2020), Antunes and Pérez-García (2024).
It was assigned to Eupleurodira by de la Fuente and Iturralde-Vinent (2001), Gaffney et al. (2006); to Panpelomedusoides by Cadena et al. (2012); to Pleurodira by Pérez-García et al. (2013), Pérez-García and Lapparent de Broin (2013); to Testudines by Menegazzo et al. (2015); to Pleurodira by Cadena et al. (2012), Pérez-García et al. (2013), Pérez-Garcia (2020), Georgalis et al. (2020), Cadena (2020), Maniel et al. (2021), Antunes and Pérez-García (2024); and to Pleurodira by Joyce et al. (2004), Hutchison et al. (2004), Oliveira and Kellner (2007), Weems and Knight (2013), Ferreira et al. (2015), de Araujo Carvalho et al. (2016), Pérez-García and Chapman (2017), Pérez-García (2017), Pérez-García and Ortega (2018), Ferreira et al. (2018), Pérez-García (2020), de Lapparent de Broin et al. (2020), López-Conde et al. (2021), Cadena et al. (2025).
Subtaxa
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1897 | Chrysemys montolivensis Roman |
| 1908 | Naiadochelys Hay figs. 125-126 |
| 1908 | Naiadochelys ingravata Hay fig. 133 |
| 1908 | Taphrosphys dares Hay p. 120 figs. Figs. 121-124 |
| 1908 | Naiadochelys Hay p. 125 |
| 1908 | Naiadochelys ingravata Hay p. 125 figs. fig. 133 |
| 1930 | Taphrosphys dares Hay p. 74 |
| 1930 | Naiadochelys Hay p. 75 |
| 1930 | Naiadochelys ingravata Hay p. 75 |
| 1948 | Naiadochelys Zangerl p. 48 |
| 1977 | Neochelys montolivensis Brion |
| 1988 | Pelomedusoides Broin p. 34 |
| 2001 | Pelomedusoides de la Fuente and Iturralde-Vinent |
| 2003 | Chrysemys montolivensis Lapparent de Broin |
| 2004 | Pelomedusoides Hutchison et al. p. 39 |
| 2004 | Pelomedusoides Joyce et al. |
| 2006 | Pelomedusoides Gaffney et al. p. 35 |
| 2007 | Pelomedusoides Oliveira and Kellner |
| 2012 | Pelomedusoides Cadena et al. p. 549 |
| 2012 | Pelomedusoides Cadena et al. |
| 2013 | Pelomedusoides Pérez-García and Lapparent de Broin |
| 2013 | Pelomedusoides Pérez-García et al. |
| 2013 | Pelomedusoides Pérez-García et al. |
| 2013 | Pelomedusoides Weems and Knight p. 290 |
| 2015 | Pelomedusoides Ferreira et al. |
| 2015 | Pelomedusoides Menegazzo et al. |
| 2016 | Pelomedusoides de Araujo Carvalho et al. |
| 2017 | Pelomedusoides Pérez-García p. 2 |
| 2017 | Pelomedusoides Pérez-García and Chapman p. 2 |
| 2018 | Pelomedusoides Ferreira et al. |
| 2018 | Pelomedusoides Pérez-García and Ortega |
| 2020 | Pelomedusoides Cadena |
| 2020 | Pelomedusoides Georgalis et al. p. 836 |
| 2020 | Pelomedusoides Pérez-Garcia |
| 2020 | Pelomedusoides Pérez-García |
| 2020 | Pelomedusoides de Lapparent de Broin et al. p. 466 |
| 2021 | Pelomedusoides López-Conde et al. |
| 2021 | Pelomedusoides Maniel et al. |
| 2024 | Pelomedusoides Antunes and Pérez-García |
| 2025 | Pelomedusoides Cadena et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Superfm. †Pelomedusoides Broin 1988
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Fm. †Araripemydidae Price 1973
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G. †Araripemys Price 1973
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G. †Laganemys Sereno and ElShafie 2013
G. †Taquetochelys Broin 1980
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†Taquetochelys decorata de Broin 1980
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Invalid names: Laganemys tenerensis Sereno and ElShafie 2013 [synonym]
Fm. †Peiropemydidae Gaffney et al. 2011
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Unr. †Pelomedusera Gaffney et al. 2006
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Fm. Pelomedusidae Cope 1868
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G. †Duerochelys Jimenez Fuentes 1975
G. Pelusios Wagler 1830
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Pelusios adansonii Schweigger 1812 [Adansons mud turtle]
Pelusios bechuanicus FitzSimons 1932 [Okavango mud turtle]
Pelusios broadleyi Bour 1986 [Turkana mud turtle]
Pelusios carinatus Laurent 1956 [African keeled mud turtle]
Pelusios castaneus Schweigger 1812 [West African mud turtle]
Pelusios castanoides Hewitt 1931 [Yellow-bellied mud turtle]
Pelusios chapini Laurent 1965 [Central African mud turtle]
Pelusios cupulatta Bour and Maran 2003 [Ivory Coast mud turtle]
Pelusios gabonensis Duméril 1856 [African forest turtle]
Pelusios marani Bour 2000 [Gabon mud turtle]
Pelusios nanus Laurent 1956 [African dwarf mud turtle]
Pelusios niger Duméril and Bibron 1835 [West African black mud turtle]
Pelusios rhodesianus Hewitt 1927 [Variable mud turtle]
†Pelusios rusingae Williams 1954
Pelusios sinuatus Smith 1838 [Serrated hinged terrapin]
Pelusios subniger Bonnaterre 1789 [East African black mud turtle]
Pelusios upembae Broadley 1981 [Upemba mud turtle]
Pelusios williamsi Laurent 1965 [Williams mud turtle]
Invalid names: Eusarkiidae Bergounioux 1952 [synonym], Pelomedusinae Zangerl 1948 [empty], Peltocephalidae Gray 1870 [synonym], Pelusiinae Zangerl 1948 [empty], Sternothaeridae Cope 1868 [synonym]
Unr. †Podocnemidera
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Invalid names: Platycheloides nyasae Haughton 1928 [nomen dubium]
Fm. Podocnemididae Cope 1868
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G. †Bairdemys Gaffney and Wood 2002
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†Bairdemys hartsteini Gaffney and Wood 2002
†Bairdemys healeyorum Weems and Knight 2013
†Bairdemys miocenica Collins and Lynn 1936
†Bairdemys sanchezi Gaffney et al. 2008
†Bairdemys thalassica Ferreira et al. 2015
†Bairdemys venezuelensis Wood and Diaz 1971
†Bairdemys winklerae Gaffney et al. 2008
G. †Bauruemys Suarez 1969
+
†Bauruemys elegans Suarez 1969
Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]
G. †Cambaremys França and Langer 2005
Subfm. †Erymnochelyinae Broin 1988
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G. †Eocenochelus Pérez-García et al. 2017
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†Eocenochelus eremberti Broin 1977
†Eocenochelus farresi Pérez-García et al. 2017
†Eocenochelus lacombianus Pérez-García et al. 2017
Tr. †Erymnochelyini de Broin 1988
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G. †Shetwemys Perez-Garcia 2021
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†Shetwemys fajumensis Andrews 1903
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Invalid names: Podocnemis blanckenhorni von Reinach 1903 [synonym]
G. †Neochelys Bergounioux 1954
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†Neochelys arenarum Broin 1977
†Neochelys bolcensis Bergounioux 1954
†Neochelys capellinii De Zigno 1890
†Neochelys eocaenica de Stefano 1902
†Neochelys franzeni Schleich 1993
†Neochelys laurenti Tong 1998
†Neochelys liriae Pérez-García and Lapparent de Broin 2013
†Neochelys mengaudi Bergounioux 1935
†Neochelys nicolisii Zigno 1890
†Neochelys salmanticensis Jimenez 1971
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Invalid names: Duerochelys arribasi Jimenez 1975 [synonym], Podocnemis armuniensis Jimenez 1970 [synonym], Podocnemis entodermica Jimenez 1970 [synonym]
†Neochelys zamorensis Jimenez-Fuentes 1992
Invalid names: Polysternum mengaudi Bergounioux 1935 [nomen dubium]
G. †Ragechelus de Lapparent de Broin et al. 2020
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†Ragechelus sahelica de Lapparent de Broin et al. 2020
G. †Stupendemys Wood 1976
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†Stupendemys geographica Wood 1976
†Stupendemys souzai Bocquetin and Melo 2006
G. †Papoulemys Tong 1998
G. Peltocephalus Duméril and Bibron 1835
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Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]
†Peltocephalus maturin Ferreira et al. 2024
Subfm. Podocnemidinae Cope 1868
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Unr. †Podocnemidodda Gaffney et al. 2011
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Invalid names: Erymnochelydand Gaffney et al. 2011 [empty], Podocnemidand Gaffney et al. 2011 [empty]
G. Podocnemis Wagler 1830 [river turtle]
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†Podocnemis argentinensis Cattoi and Freiberg 1958
†Podocnemis bassleri Williams 1956
†Podocnemis brasiliensis Staesche 1937
Podocnemis erythrocephala Spix 1824 [Red-headed river turtle]
Podocnemis expansa Schweigger 1812
†Podocnemis geologorum Simpson 1943
†Podocnemis harrisi Pacheco 1913
Podocnemis lewyana Dumeril 1852
†Podocnemis medemi Wood 1997
†Podocnemis negrii Carvalho et al. 2002
†Podocnemis pritchardi Wood 1997
Podocnemis sextuberculata Cornalia 1849
†Podocnemis stromeri von Reinach 1903
†Podocnemis tatacoensis Cadena and Vanegas 2023
Podocnemis unifilis Troschel 1848
Podocnemis vogli Muller 1935
Invalid names: Podocnemis bramlyi Fourtau 1918 [nomen dubium], Podocnemis freibergi Agnolin 2004 [nomen dubium]
Tr. †Stereogenyini Gaffney et al. 2011
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Unr. †Stereogenyita Gaffney et al. 2011
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Invalid names: Mogharemydina Gaffney et al. 2011 [empty]
G. †Stereogenys Andrews 1901
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†Stereogenys cromeri Andrews 1901
Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]
Invalid names: Albertwoodemys Gaffney et al. 2011 [valid], Bairdemys Gaffney and Wood 2002 [valid], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemys Suarez 1969 [valid], Brontochelys Gaffney et al. 2011 [valid], Cambaremys França and Langer 2005 [valid], Carbonemys Cadena et al. 2012 [valid], Cerrejonemys Cadena et al. 2010 [valid], Cordichelys Gaffney et al. 2011 [valid], Dacquemys Williams 1954 [valid], Erymnochelyinae Broin 1988 [valid], Erymnochelys Baur 1888 [valid], Gestemys de la Fuente et al. 2022 [valid], Hamadachelys Tong and Buffetaut 1996 [valid], Kenyemys Wood 1983 [valid], Lapparentemys Gaffney et al. 2011 [valid], Latentemys Gaffney et al. 2011 [valid], Lemurchelys Gaffney et al. 2011 [valid], Mogharemys Gaffney et al. 2011 [valid], Papoulemys Tong 1998 [valid], Peiropemys Gaffney et al. 2011 [valid], Peltocephalus Duméril and Bibron 1835 [valid], Podocnemidinae Cope 1868 [valid], Podocnemoidea Cope 1868 [empty], Podocnemoidea Cope 1868 [empty], Shakiremys Cadena et al. 2025 [valid], Shweboemys Swinton 1939 [valid], Stereogenyina Gaffney et al. 2011 [valid], Stereogenyini Gaffney et al. 2011 [valid], Stereogenys Andrews 1901 [valid], Turkanemys Wood 2003 [valid]
Superfm. Podocnemidoidae Cope 1868
+
G. †Elkanemys Maniel et al. 2021
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†Elkanemys caelestis Sarda and Maniel 2025
†Elkanemys pritchardi Maniel et al. 2021
G. †Roxochelys Price 1953
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†Roxochelys vilavilensis de Broin 1971
†Roxochelys wanderleyi Price 1953
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Invalid names: Cambaremys langertoni França and Langer 2005 [synonym]
Invalid names: Roxochelys harrisi Pacheco 1913 [nomen dubium]
Invalid names: Brasilemydidae Lapparent de Broin 2000 [empty]
Superfm. †Podocnemidoidea Gaffney et al. 2006
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Fm. Podocnemididae Cope 1868
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G. †Bairdemys Gaffney and Wood 2002
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†Bairdemys hartsteini Gaffney and Wood 2002
†Bairdemys healeyorum Weems and Knight 2013
†Bairdemys miocenica Collins and Lynn 1936
†Bairdemys sanchezi Gaffney et al. 2008
†Bairdemys thalassica Ferreira et al. 2015
†Bairdemys venezuelensis Wood and Diaz 1971
†Bairdemys winklerae Gaffney et al. 2008
G. †Bauruemys Suarez 1969
+
†Bauruemys elegans Suarez 1969
Invalid names: Bauruemys brasiliensis Staesche 1937 [nomen dubium]
G. †Cambaremys França and Langer 2005
Subfm. †Erymnochelyinae Broin 1988
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G. †Eocenochelus Pérez-García et al. 2017
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†Eocenochelus eremberti Broin 1977
†Eocenochelus farresi Pérez-García et al. 2017
†Eocenochelus lacombianus Pérez-García et al. 2017
Tr. †Erymnochelyini de Broin 1988
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G. †Shetwemys Perez-Garcia 2021
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†Shetwemys fajumensis Andrews 1903
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Invalid names: Podocnemis blanckenhorni von Reinach 1903 [synonym]
G. †Neochelys Bergounioux 1954
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†Neochelys arenarum Broin 1977
†Neochelys bolcensis Bergounioux 1954
†Neochelys capellinii De Zigno 1890
†Neochelys eocaenica de Stefano 1902
†Neochelys franzeni Schleich 1993
†Neochelys laurenti Tong 1998
†Neochelys liriae Pérez-García and Lapparent de Broin 2013
†Neochelys mengaudi Bergounioux 1935
†Neochelys nicolisii Zigno 1890
†Neochelys salmanticensis Jimenez 1971
+
Invalid names: Duerochelys arribasi Jimenez 1975 [synonym], Podocnemis armuniensis Jimenez 1970 [synonym], Podocnemis entodermica Jimenez 1970 [synonym]
†Neochelys zamorensis Jimenez-Fuentes 1992
Invalid names: Polysternum mengaudi Bergounioux 1935 [nomen dubium]
G. †Ragechelus de Lapparent de Broin et al. 2020
+
†Ragechelus sahelica de Lapparent de Broin et al. 2020
G. †Stupendemys Wood 1976
+
†Stupendemys geographica Wood 1976
†Stupendemys souzai Bocquetin and Melo 2006
G. †Papoulemys Tong 1998
G. Peltocephalus Duméril and Bibron 1835
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Peltocephalus dumerilianus Schweigger 1812 [Bighead Amazon River turtle]
†Peltocephalus maturin Ferreira et al. 2024
Subfm. Podocnemidinae Cope 1868
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Unr. †Podocnemidodda Gaffney et al. 2011
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Invalid names: Erymnochelydand Gaffney et al. 2011 [empty], Podocnemidand Gaffney et al. 2011 [empty]
G. Podocnemis Wagler 1830 [river turtle]
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†Podocnemis argentinensis Cattoi and Freiberg 1958
†Podocnemis bassleri Williams 1956
†Podocnemis brasiliensis Staesche 1937
Podocnemis erythrocephala Spix 1824 [Red-headed river turtle]
Podocnemis expansa Schweigger 1812
†Podocnemis geologorum Simpson 1943
†Podocnemis harrisi Pacheco 1913
Podocnemis lewyana Dumeril 1852
†Podocnemis medemi Wood 1997
†Podocnemis negrii Carvalho et al. 2002
†Podocnemis pritchardi Wood 1997
Podocnemis sextuberculata Cornalia 1849
†Podocnemis stromeri von Reinach 1903
†Podocnemis tatacoensis Cadena and Vanegas 2023
Podocnemis unifilis Troschel 1848
Podocnemis vogli Muller 1935
Invalid names: Podocnemis bramlyi Fourtau 1918 [nomen dubium], Podocnemis freibergi Agnolin 2004 [nomen dubium]
Tr. †Stereogenyini Gaffney et al. 2011
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Unr. †Stereogenyita Gaffney et al. 2011
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Invalid names: Mogharemydina Gaffney et al. 2011 [empty]
G. †Stereogenys Andrews 1901
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†Stereogenys cromeri Andrews 1901
Invalid names: Stereogenys podocnemoides Reinach 1903 [nomen dubium]
Invalid names: Albertwoodemys Gaffney et al. 2011 [valid], Bairdemys Gaffney and Wood 2002 [valid], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemydinae Gaffney et al. 2011 [empty], Bauruemys Suarez 1969 [valid], Brontochelys Gaffney et al. 2011 [valid], Cambaremys França and Langer 2005 [valid], Carbonemys Cadena et al. 2012 [valid], Cerrejonemys Cadena et al. 2010 [valid], Cordichelys Gaffney et al. 2011 [valid], Dacquemys Williams 1954 [valid], Erymnochelyinae Broin 1988 [valid], Erymnochelys Baur 1888 [valid], Gestemys de la Fuente et al. 2022 [valid], Hamadachelys Tong and Buffetaut 1996 [valid], Kenyemys Wood 1983 [valid], Lapparentemys Gaffney et al. 2011 [valid], Latentemys Gaffney et al. 2011 [valid], Lemurchelys Gaffney et al. 2011 [valid], Mogharemys Gaffney et al. 2011 [valid], Papoulemys Tong 1998 [valid], Peiropemys Gaffney et al. 2011 [valid], Peltocephalus Duméril and Bibron 1835 [valid], Podocnemidinae Cope 1868 [valid], Podocnemoidea Cope 1868 [empty], Podocnemoidea Cope 1868 [empty], Shakiremys Cadena et al. 2025 [valid], Shweboemys Swinton 1939 [valid], Stereogenyina Gaffney et al. 2011 [valid], Stereogenyini Gaffney et al. 2011 [valid], Stereogenys Andrews 1901 [valid], Turkanemys Wood 2003 [valid]
Unr. †Podocnemidinura Gaffney et al. 2006
+
Unr. †Sahonachelyidae Joyce et al. 2021
+
Invalid names: Chrysemys montolivensis Roman 1897 [invalid subgroup], Naiadochelys Hay 1908 [nomen dubium], Panpodocnemidae Joyce et al. 2004 [empty], Taphrosphys dares Hay 1908 [nomen dubium]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| O. A. López-Conde et al. 2021 | IGM-11517, can be attributed to Pleurodira considering the following character combination: a pair of reduced mesoplastra, not meeting in midline; a single intergular scute; absence of inframarginals and pelvis sutured to the shell. |
Measurements
No measurements are available
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| Source: subo = suborder, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Uetz 2005, Carroll 1988, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Tithonian to the top of the Holocene or 145.06000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 143.1 Ma
Minimum age of oldest fossil (stem group age): 143.1 Ma
Collections (376 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Tithonian | Ethiopia (Shewa) | Pelomedusidae indet. (48832) | |
| Late/Upper Tithonian - Early/Lower Berriasian | Morocco (Oriental) | Araripemydidae indet., Taquetochelys sp. (45726) | |
| Berriasian - Hauterivian | Brazil (Paraíba) | Araripemydidae indet. (58790) | |
| Early/Lower Cretaceous - Aptian | Brazil (Ceará) | Pelomedusidae indet. (59355) | |
| Late/Upper Valanginian | Colombia | Pelomedusoides indet. (137831) Podocnemidoidea indet. (118389) | |
| Valanginian | Colombia (Santander) | Pelomedusoides indet. (213310) | |
| Hauterivian - Barremian | Libya (Yafran) | Pelomedusidae indet. (69791) | |
| Late/Upper Barremian | Brazil (Alagoas) | Pelomedusoides indet. (152472) | |
| Late/Upper Barremian | Spain (Cuenca) | Pelomedusidae indet. (61319) | |
| Aptian | Malawi (Northern) | Platycheloides nyasae (38911 89853) | |
| Aptian | Niger (Agadez) | Teneremys lapparenti, Pelomedusidae indet. (37200) | |
| Aptian - Albian | Brazil (Ceará) | Pelomedusoides indet. (132359) | |
| Aptian - Albian | Niger (Agadez) | Laganemys tenerensis (133668) | |
| Late/Upper Aptian | Niger (Agadez) | Araripemys sp., Teneremys lapparenti, Taquetochelys decorata (77189) | |
| Late/Upper Aptian | Brazil (Ceará) | Araripemys barretoi (128565) Araripemys barretoi, Caririemys violetae (67712) Araripemys barretoi, Euraxemys essweini (83306) Brasilemys josai (92290) | |
| Late/Upper Aptian | Brazil (Ceara) | Araripemys sp., Araripemys arturi (77891) | |
| Late/Upper Aptian - Middle Albian | Brazil (Maranhão) | Chelonia indet. (67219) | |
| Albian | Niger (Agadez) | Araripemydidae indet., Teneremys lapparenti (228823 228824) Pelomedusidae indet. (28569) | |
| Late/Upper Albian - Early/Lower Cenomanian | Morocco (Drâa-Tafilalet) | Araripemys sp., Dirqadim schaefferi, Hamadachelys escuilliei (155306) Dirqadim schaefferi, Hamadachelys escuilliei (13274) | |
| Late/Upper Albian - Early/Lower Cenomanian | Algeria (Adrar) | Araripemydidae indet. (59578) | |