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Worthenia
Taxonomy
Worthenia was named by de Koninck (1883) [Sepkoski's age data: D Fras J Plie-u Sepkoski's reference number: 362,797]. It is not extant. Its type is Turbo tabulatus.
It was reranked as Pleurotomaria (Worthenia) by Perner (1907).
It was assigned to Pleurotomariidae by Ulrich and Scofield (1897); to Pleurotomaria by Perner (1907); to Ruedemanninae by Knight et al. (1960); to Archaeogastropoda by Sepkoski (2002); to Lophospiridae by Szabo (1980), Kues and Batten (2001), Kaim et al. (2014), Nützel et al. (2018) and Ketwetsuriya et al. (2020); and to Phymatopleuridae by Karapunar and Nützel (2022).
It was reranked as Pleurotomaria (Worthenia) by Perner (1907).
It was assigned to Pleurotomariidae by Ulrich and Scofield (1897); to Pleurotomaria by Perner (1907); to Ruedemanninae by Knight et al. (1960); to Archaeogastropoda by Sepkoski (2002); to Lophospiridae by Szabo (1980), Kues and Batten (2001), Kaim et al. (2014), Nützel et al. (2018) and Ketwetsuriya et al. (2020); and to Phymatopleuridae by Karapunar and Nützel (2022).
Species
Pleurotomaria (Worthenia) constantini, W. (Humiliworthenia), W. (Worthenia), W. alticarinata, W. angulata, W. basifalcata, W. bialveozona, W. bicarinata, W. bieberi, W. bilineata, W. cainallensis, W. castlemanensis, W. cirriformis, W. constantini, W. contabulata, W. corrugata, W. crenilunula, W. crenulata, W. curvistriata, W. dharmaensis, W. dieneri, W. distincta, W. dregeri, W. duplicata, W. dyadica, W. eremita, W. escheri, W. extendia, W. furcata, W. gepidorum, W. gubleri, W. hoettingensis, W. humiligrada, W. humilis, W. indifferens, W. kingi, W. klamathensis, W. latialveozona, W. legrandi, W. liebeneri, W. loczyi, W. margaritacea, W. morovi, W. multicarinata, W. nana, W. pagoda, W. parvula, W. pilula, W. planalveozona, W. plutonis, W. praerupta, W. punctata, W. rhombifera, W. schirjaevensis, W. sculpta, W. sigaretoides, W. speciosa (syn. W. perizomata), W. strigata, W. strigillata, W. subplicata, W. substriata, W. subtilis, W. superstes, W. tabulata, W. tongxinensis, W. triton, W. vamosensis, W. whitehorsensis (syn. W. beedei), W. windowblindensis
Species lacking formal opinion data
Synonymy list
| Year | Name and author |
|---|---|
| 1883 | Worthenia de Koninck p. 267 |
| 1897 | Worthenia Ulrich and Scofield |
| 1907 | Pleurotomaria (Worthenia) Perner p. 63 |
| 1960 | Worthenia Knight et al. p. 209 |
| 1980 | Worthenia Szabo p. 56 |
| 2001 | Worthenia Kues and Batten |
| 2002 | Worthenia Sepkoski |
| 2014 | Worthenia Kaim et al. p. 168 |
| 2018 | Worthenia Nützel et al. p. 486 |
| 2020 | Worthenia Ketwetsuriya et al. p. 59 |
| 2022 | Worthenia Karapunar and Nützel p. 53 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Worthenia de Koninck 1883
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†Pleurotomaria (Worthenia) constantini Mansuy 1912
Subg. †Worthenia (Humiliworthenia) Yin and Yochelson 1983
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†Worthenia (Humiliworthenia) anisica Nützel et al. 2018
†Worthenia (Humiliworthenia) hausmanni Goldfuss 1841
†Worthenia (Humiliworthenia) nuda Koken 1900
Subg. †Worthenia (Worthenia) de Koninck 1883
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†Worthenia (Worthenia) conica Assmann 1937
†Worthenia (Worthenia) dobrogeana Nützel et al. 2018
†Worthenia (Worthenia) ecki Assmann 1924
†Worthenia (Worthenia) extensus Yin and Yochelson 1983
†Worthenia (Worthenia) xui Yin and Yochelson 1983
†Worthenia alticarinata Batten 1989
†Worthenia angulata Münster 1841
†Worthenia basifalcata Korner 1937
†Worthenia bialveozona Batten 1989
†Worthenia bicarinata Batten 1989
†Worthenia bieberi Kittl 1891
†Worthenia bilineata Klipstein 1843
†Worthenia cainallensis Kittl 1899
†Worthenia castlemanensis Lintz 1958
†Worthenia cirriformis Laube 1868
†Worthenia constantini Mansuy 1912
†Worthenia contabulata Costa 1964
†Worthenia corrugata Chronic 1952
†Worthenia crenilunula Yoo 1994
†Worthenia crenulata Batten 1989
†Worthenia curvistriata Koken and Perner 1925
†Worthenia dharmaensis Blaschke 1907
†Worthenia dieneri Blaschke 1907
†Worthenia distincta Kittl 1894
†Worthenia dregeri Kittl 1891
†Worthenia duplicata Kittl 1891
†Worthenia dyadica Kittl 1903
†Worthenia eremita Koken 1897
†Worthenia escheri Stoppani 1865
†Worthenia extendia Tong and Erwin 2001
†Worthenia furcata Kittl 1891
†Worthenia gepidorum Kittl 1900
†Worthenia gubleri Delpey 1941
†Worthenia hoettingensis Alma 1926
†Worthenia humiligrada Ketwetsuriya et al. 2020
†Worthenia humilis Böhm 1895
†Worthenia indifferens Kittl 1894
†Worthenia kingi Batten 1989
†Worthenia klamathensis Smith 1927
†Worthenia latialveozona Batten 1989
†Worthenia legrandi Kues and Batten 2001
†Worthenia liebeneri Laube 1868
†Worthenia loczyi Kittl 1900
†Worthenia margaritacea Laube 1868
†Worthenia morovi Mazaev 2020
†Worthenia multicarinata Mansuy 1912
†Worthenia nana Grabau 1931
†Worthenia pagoda Mansuy 1914
†Worthenia parvula Hoare et al. 1997
†Worthenia pilula Batten 1989
†Worthenia planalveozona Batten 1989
†Worthenia plutonis Kittl 1894
†Worthenia praerupta Böhm 1895
†Worthenia punctata Wanner 1922
†Worthenia rhombifera Korner 1937
†Worthenia schirjaevensis Stuckenberg 1905
†Worthenia sculpta Phillips 1836
†Worthenia sigaretoides Kittl 1894
†Worthenia speciosa Meek and Worthen 1860
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Invalid names: Worthenia perizomata White 1881 [synonym]
†Worthenia strigata Böhm 1895
†Worthenia strigillata Herrick 1888
†Worthenia subplicata Klipstein 1843
†Worthenia substriata Klipstein 1843
†Worthenia subtilis Kittl 1891
†Worthenia superstes Szabo 1980
†Worthenia tabulata Conrad 1835
†Worthenia tongxinensis Guo 1983
†Worthenia triton d'Orbigny 1849
†Worthenia vamosensis Kittl 1900
†Worthenia whitehorsensis Saunders and Yochelson 1964
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Invalid names: Worthenia beedei Knight 1940 [replaced]
†Worthenia windowblindensis Batten and Stokes 1986
Diagnosis
No diagnoses are available
Measurements
| part | N | mean | minimum | maximum | median | s.d. | |
|---|---|---|---|---|---|---|---|
| height | 1 | 5.40 | - | - | - | - | |
| shell height | 99 | 8.89 | 1.70 | 81.0 | 9.00 | 10.8 | |
| shell diameter | 99 | 9.55 | 3.10 | 70.1 | 8.10 | 15.1 |
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| Reference: Bambach et al. 2007 | |||||
Collections (361 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Hastarian | Australia (New South Wales) | W. crenilunula (43901) | |
| Kinderhookian | Australia (New South Wales) | W. sp. (75614) | |
| Tournaisian | Australia (New South Wales) | W. sp. (43950 44080 44088) W. sp., W. crenilunula (43904 44079) | |
| Tournaisian | USA (Iowa) | W. sp. (3309) | |
| Carboniferous | USA (Oklahoma) | W. sp. (163) | |
| Osagean | USA (Ohio) | W. strigillata (93067 155894) | |
| Meramecian | USA (Nevada) | W. sp. (62674) | |
| Brigantian | United Kingdom (England) | Ptychomphalus sculptus (67670) | |
| Brigantian - Pendleian | USA (Pennsylvania) | W. sp. (2774) | |
| Chesterian | USA (Oklahoma) | W. sp. (584) | |
Youngest occurrences |
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| Time interval | Ma | Country or state | PBDB collection number |
| Sevatian | USA (Idaho) | W. sp., W. rhombifera (38377) | |
| Rhaetian | Peru | W. basifalcata (68116) W. n. sp. 1, W. rhombifera, W. basifalcata (68096) | |
| Rhaetian | Hungary (Veszprem) | W. escheri (203351) | |
| Raricostatum - Davoei | Hungary | W. superstes (78690) | |
| Pliensbachian | Hungary | W. superstes (91406) | |
| Davoei | Hungary | W. superstes (108954 119137) | |
| Ibex | Hungary | W. superstes (108945 108949) | |
| Margaritatus | Spain (Alicante ) | W. superstes (130881) | |
| Margaritatus | Hungary | W. superstes (108960) | |
| Early/Lower Pliensbachian - Middle Toarcian | Hungary | W. superstes (108969) | |