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Taxonomy
Aradiens was named by Brullé (1836). It is extant. Its type is Aradus.
It was corrected as Aradidae by Wappler (2003), Wappler and Heiss (2006), Cassis and Schuh (2010), Szwedo (2018) and Azar et al. (2020).
It was assigned to Heteroptera by Wappler (2003); and to Aradoidea by Wappler and Heiss (2006), Cassis and Schuh (2010), Szwedo (2018) and Azar et al. (2020).
It was corrected as Aradidae by Wappler (2003), Wappler and Heiss (2006), Cassis and Schuh (2010), Szwedo (2018) and Azar et al. (2020).
It was assigned to Heteroptera by Wappler (2003); and to Aradoidea by Wappler and Heiss (2006), Cassis and Schuh (2010), Szwedo (2018) and Azar et al. (2020).
Synonymy list
| Year | Name and author |
|---|---|
| 2003 | Aradidae Wappler |
| 2006 | Aradidae Wappler and Heiss p. 40 |
| 2010 | Aradidae Cassis and Schuh |
| 2018 | Aradidae Szwedo p. 113 |
| 2020 | Aradidae Azar et al. p. 7 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Aradidae Brullé 1836 [flat bug]
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Subfm. Aneurinae Douglas and Scott 1865
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G. Aneurus Curtis 1925
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Subg. Aneurus (Aneurodes) Heiss 1998
Subg. Aneurus (Aneurus) Curtis 1925
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†Aneurus (Aneurus) ancestralis Heiss 1997
†Aneurus (Aneurus) groehni Heiss 2001
Subg. †Aneurus (Paraneurosoma) Heiss 2012
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†Aneurus (Paraneurosoma) damzeni Heiss et al. 2021
†Aneurus (Paraneurosoma) ursulae Heiss 2012
†Aneurus incertus Heiss et al. 2015
†Aneurus opertus Zhang et al. 1994
Subfm. Aradinae Brullé 1836
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G. Aradus Fabricius 1803
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†Aradus andancensis Marchal et al. 2011
†Aradus antediluvianus Heer 1853
†Aradus assimilis Germar and Berendt 1856
†Aradus balticus Heiss 2002
†Aradus consimilis Germar and Berendt 1856
†Aradus creticus Kormilev and Popov 1986
†Aradus damzeni Heiss 2002
†Aradus frater Popov 1978
†Aradus frateroides Heiss 1998
†Aradus goellnerae Heiss 2002
†Aradus grabenhorsti Heiss 2013
†Aradus kotashevichi Heiss 2002
†Aradus lativentris Heiss 2002
†Aradus leptosomus Heiss 2014
†Aradus macrosomus Heiss 2014
†Aradus madagascariensis Bervoets 1909
†Aradus nicholasi Popov 1989
†Aradus penteneuros Heiss 2014
†Aradus popovi Heiss 1998
†Aradus rotundiventris Heiss 2014
†Aradus superstes Germar and Berendt 1856
†Aradus velteni Heiss 2002
†Aradus voigti Heiss 2002
†Aradus weitschati Heiss 2002
Subfm. †Archearadinae Heiss and Grimaldi 2002
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G. †Archearadus Heiss and Grimaldi 2001
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†Archearadus burmensis Heiss and Grimaldi 2001
†Archearadus elongatus Heiss 2016
G. †Cretopiesma Grimaldi and Engel 2008
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†Cretopiesma anticum Heiss and Poinar 2012
†Cretopiesma engelgrimaldii Azar et al. 2020
†Cretopiesma inexpectatum Azar et al. 2020
†Cretopiesma lini Azar et al. 2020
†Cretopiesma suukyiae Grimaldi and Engel 2008
Invalid names: Microaradus Heiss and Poinar 2012 [synonym]
Subfm. Calisiinae Stal 1873
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G. Calisiopsis Champion 1898
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†Calisiopsis azteca Heiss 2016
†Calisiopsis brodzinskyorum Froeschner 1992
G. Calisius Stal 1860
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†Calisius balticus Usinger 1941
†Calisius hoffeinsorum Heiss 2002
†Calisius rietscheli Heiss 2000
†Calisius vonholti Heiss 2000
†Calisius weitschati Heiss 2000
G. †Calisiomorpha Heiss 2016
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†Calisiomorpha herczeki Heiss 2023
†Calisiomorpha yuripopovi Heiss 2016
Subfm. Carventinae Usinger and Matsuda 1959
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Subfm. Mezirinae Oshanin 1908
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G. Mezira Amyot and Audinet-Serville 1843
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†Mezira crassifemur Wappler and Heiss 2006
†Mezira eckfeldensis Wappler and Heiss 2006
†Mezira eocenica Wappler and Heiss 2006
†Mezira parapetrificata Heiss et al. 2015
†Mezira petrificata Heiss et al. 2015
†Mezira scheveni Heiss 2000
†Mezira succinica Usinger 1941
G. Neuroctenus Fieber 1860
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†Neuroctenus enspelensis Wappler and Heiss 2006
†Neuroctenus incompletus Wappler and Heiss 2006
†Neuroctenus kotejai Wappler and Heiss 2006
†Neuroctenus messelensis Wappler and Heiss 2006
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Aptian to the top of the Holocene or 125.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 113.0 Ma
Minimum age of oldest fossil (stem group age): 113.0 Ma
Collections (46 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Aptian | Mongolia | Aradus nicholasi (103657) | |
| Early/Lower Cenomanian | Myanmar | Archemezira nuoxichenae (229441) Cretopiesma anticum, Cretopiesma engelgrimaldii, Cretopiesma lini, Cretopiesma inexpectatum (161726) Cretopiesma suukyiae, Archearadus burmensis (111688) Cretopiesma suukyiae, Microaradus anticus, Calisiomorpha yuripopovi, Ellenbergeria oviventris, Aradoleptus birmanus, Archearadus elongatus, Kachinocoris brevipennis, Archecalisius longiventris, Archeaneurus neli, Cretopiesma anticum, Cretopiesma lini, Cretopiesma inexpectatum, Pachytylaradus cretaceous, Cretozemira elongata, Calisiomorpha herczeki (153736) | |
| Early/Lower Cenomanian | Myanmar (Kachin) | Myanmezira longicornis, Microaradus anticus (104053) | |
| Early/Lower Campanian | Russian Federation (Tenkinskii) | Aradus creticus (105375) | |
| Ypresian | France (Oise) | Calisius balticus (123850) | |
| MP 11 | Germany (Hessen) | Aneurus incertus (173355) Mezira eocenica (125466 144629) Mezira parapetrificata (68475) Mezira petrificata (166830 173356) Neuroctenus kotejai (144627) Neuroctenus messelensis (144628) | |
| MP 13 | Germany (Eifel) | Aradidae indet. (135641) Mezira crassifemur, Aradidae indet. (138912) Neuroctenus incompletus, Aradidae indet. (135643) | |
| Priabonian | Poland | Aradus frater, Aradus popovi (120813) Aradus superstes, Aradus assimilis, Aradus consimilis, Aradidae larva indet. (109625) | |
| Priabonian | Russian Federation (Kaliningrad) | Aradus damzeni, Calisius weitschati (123764) Aradus goellnerae, Aradus lativentris (123221) Aradus grabenhorsti (123770) Calisius balticus, Aneurus groehni (110735) Calisius balticus, Mezira succinica (124205) Calisius sp., Calisius balticus, Aneurus kotashevichi, Aneurus ancestralis, Allocalisius spiniventris, Calisius vonholti, Aradus weitschati, Aradus kotashevichi, Aradus velteni, Aradus voigti, Aradus balticus, Aradus frateroides, Calisius rietscheli, Aradus grabenhorsti, Aradus macrosomus, Aneurus ursulae, Quilnus rectinotus, Aradus leptosomus, Aradus rotundiventris, Aradus penteneuros, Aneurus damzeni (123862) | |
| Priabonian | Germany (Halle) | Aneurus goitschenus (169965) Aradus grabenhorsti (147606) Calisius balticus (124273 149475 169966) | |
| Priabonian | Lithuania | Aradus frateroides (123767) | |
| Priabonian | Russian Federation | Allocalisius spiniventris, Calisius hoffeinsorum, Aradus kotashevichi (123419) | |
| MP 28 | Germany (Rheinland-Pfalz) | Neuroctenus enspelensis (160097) | |
| Aquitanian | New Zealand | Aneurus sp. (50035) | |
| Early/Lower Miocene | Mexico (Chiapas) | Calisiopsis azteca (178678) | |
| Early/Lower Miocene - Middle Miocene | Thailand | Neuroctenus sp. (160098) | |
| Burdigalian | China (Shandong) | Aneurus opertus (138946) | |
| Burdigalian - Langhian | Dominican Republic | Brevisensoria incrustata, Neoproxius latocanus, Acaricoris robertae, Proneoproxius cornutus (122472) Calisiopsis brodzinskyorum (122473) Carventus bechlyi (122474) Mezira scheveni (124229) | |
| Sarmatian | Croatia | Aradus antediluvianus (117246) | |
| Turolian | France (Ardèche) | Aradus andancensis (113893) | |
| Holocene | Madagascar | Aradus madagascariensis (151372) |