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Athyridoidea

Rhynchonellata - Athyridida

Taxonomy
Athyridoidea was named by McCoy (1844).

It was assigned to Athyrididina by Alvarez and Brunton (1993) and Musteikis and Modzalevskaya (2002).

Synonymy list
YearName and author
1844Athyridoidea McCoy
1993Athyridoidea Alvarez and Brunton p. 310
2002Athyridoidea Musteikis and Modzalevskaya

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Spiralia
superphylumLophotrochozoa
Lophophorata
PanbrachiopodaCarlson and Cohen 2020
RankNameAuthor
phylumBrachiopodaCuvier 1805
subphylumRhynchonelliformeaWilliams et al. 1996
classRhynchonellataWilliams et al. 1996
orderAthyrididaBoucot et al. 1964
suborderAthyrididinaBoucot et al. 1964
superfamilyAthyridoideaMcCoy 1844
superfamilyAthyridoideaMcCoy 1844

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. †Athyridoidea McCoy 1844
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Fm. †Athyrididae McCoy 1841
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Subfm. †Athyridinae M'Coy 1844
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Actinoconchus planosulcatus Phillips 1836
G. †Athyris
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Subg. †Athyris (Cliothyris) King 1850
Subg. †Athyris (Seminula) M'Coy 1844
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Subg. †Spirigera (Didymospira) Salomon 1895
Spirigera borovacensis Kittl 1903
Spirigera forojulensis Bittner 1890
Spirigera kaihikuana Trechmann 1917
Spirigera kittlii Bittner 1902
Spirigera manzavinii Bittner 1891
Spirigera manzavinioides Trechmann 1917
Spirigera milesi Smith 1927
Spirigera misolica Krumbeck 1913
Spirigera monticola White 1874
Spirigera oppelii Laube 1865
Spirigera pseudodielasma Fliegel 1901
Spirigera sellaris Klipstein 1843
Spirigera siljanensis Kittl 1903
Spirigera stoliczkai Bittner 1899
Spirigera subindistincta Krumbeck 1924
Spirigera sufflata Verneuil 1855
Spirigera tricupii Bittner 1892
Spirigera wreyi Zittel 1864
Athyris ambiguaeformis Waagen 1883
Athyris angelica Hall 1861
Athyris angulata Mansuy 1912
Athyris bisulcata Jin and Hu 1982
Athyris bradyensis Carter 1967
Athyris cestriensis Snider 1915
Athyris concentrica von Buch 1834
Athyris damesi Fliegel 1901
Athyris globularis Phillips 1835
Athyris goniocolpos Frech 1912
Athyris lamellosa Léveillé 1835
Athyris mirabilis Zeng et al. 1995
Athyris persinuata Meek 1877
Athyris plicatiformis Jin and Hu 1982
Athyris simulans Reed 1931
Athyris spiriferoides Eaton 1831
Athyris subquadrata Yang and Xu 1966
Athyris xetra Diener 1915
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Atrythyris lacrimula Struve 1965
G. †Brimethyris Alvarez et al. 1998
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Brimethyris subconcentrica Verneuil and Archiac 1845
G. †Bruntonites Struve 1992
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G. †Eifyris
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Eifyris socia Godefroid and Mottequin 2005
G. †Johnsonathyris Savage et al. 1978
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Johnsonathyris adrianensis Savage et al. 1978
G. †Pachyplax
Subfm. †Athyrisininae Grabau 1931
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Invalid names: Athyrisinopsis [synonym], Kwangsia [synonym]
G. †Ikella
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Invalid names: Neoathyrisina Chen and Wan 1979 [synonym]
G. †Baliqliqia Chen and Shi 2006
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Baliqliqia baliqliqensis Chen and Shi 2006
Subfm. †Cleiothyridininae Alvarez et al. 1998
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G. †Bajtugania Grunt 1980
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Bajtugania boguchanica Solomina 1988
Bajtugania duseevi Grunt 1991
Bajtugania netschaevi Grunt 1980
G. †Carteridina Alvarez et al. 1998
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Carteridina prouti Swallow 1860
G. †Cleiothyridina Buckman 1906
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Cleiothyridina accola Reed 1944
Cleiothyridina anabathra Waterhouse 1968
Cleiothyridina attenuata Cooper 1957
Cleiothyridina bajkurica Chernyak 1963
Cleiothyridina baracoodensis Etheridge Jr. 1903
Cleiothyridina barbata Chronic 1949
Cleiothyridina capillata Waagen 1883
Cleiothyridina circularis Ustritsky 1960
Cleiothyridina ciriacksi Cooper and Grant 1976
Cleiothyridina corculum Lee and Su 1980
Cleiothyridina davidsoni Rigaux 1873
Cleiothyridina deroissyi Leveille 1835
Cleiothyridina echidniformis Waterhouse 1983
Cleiothyridina excavata Grabau 1931
Cleiothyridina globulina Waagen 1883
Cleiothyridina grossula Waagen 1883
Cleiothyridina hayasakai Tazawa 2010
Cleiothyridina intonsa Chronic 1949
Cleiothyridina laqueata Waterhouse 1968
Cleiothyridina macleayana Etheridge Jr. 1889
Cleiothyridina maynci Dunbar 1955
Cleiothyridina milleri Savage et al. 1978
Cleiothyridina mulsa Cooper and Grant 1976
Cleiothyridina nana Cooper and Grant 1976
Cleiothyridina nielseni Dunbar 1955
Cleiothyridina obmaxima McChesney 1861
Cleiothyridina orbicularis McChesney 1859
Cleiothyridina ovalis Shi and Archbold 1993
Cleiothyridina perthensis Archbold 1997
Cleiothyridina pijaensis Waterhouse 1983
Cleiothyridina pilularis Cooper and Grant 1976
Cleiothyridina rara Cooper and Grant 1976
Cleiothyridina rectimarginata Cooper and Grant 1976
Cleiothyridina seriata Grant 1976
Cleiothyridina shenshuensis Lee and Su 1980
Cleiothyridina uralica Grabau 1934
Invalid names: Cleiothyridellina Waterhouse 1978 [synonym]
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Crinisarina angelicoides Merriam 1940
Crinisarina pseudoglobularis Serobyan et al. 2021
G. †Deltachania Waterhouse 1971
G. †Himathyris Waterhouse 1985
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Himathyris gerardi Diener 1899
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Leiothycridina expansa Phillips 1836
G. †Pinegathyris Grunt 1980
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Pinegathyris alata Grunt 1980
Pinegathyris kotlukovi Stepanov 1936
Pinegathyris royssiana Keyserling 1846
Pinegathyris stuckenbergi Netschajew 1911
G. †Rawdonia
Subfm. †Comelicaniinae Merla 1930
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G. †Comelicania Frech 1901
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Comelicania haueri Stache 1878
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Invalid names: Comelicania avis Merla 1930 [synonym], Comelicania dalpiazi Merla 1930 [synonym], Comelicania vespertilio Merla 1930 [synonym]
Comelicania megalotis Stache 1878
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Invalid names: Comelicania doryphora Merla 1930 [synonym], Comelicania rostrata Merla 1930 [synonym], Spirifer insanus Stache 1878 [synonym], Spirifer vultur Stache 1878 [synonym]
Comelicania merlai Posenato 1998
Invalid names: Alatothyris Waterhouse and Gupta 1986 [synonym], Spitispirifer Waterhouse and Gupta 1986 [synonym]
G. †Gruntallina Waterhouse and Gupta 1986
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Subfm. †Didymothyridinae Modzalevskaya 1979
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G. †Buchanathyris Talent 1956
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Buchanathyris pulchra Talent 1963
Buchanathyris westoni Talent 1956
G. †Collarothyris Modzalevskaya 1970
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Collarothyris canaliculata Wenjukow 1899
G. †Didymothyris Rubel and Modzalevskaya 1967
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Didymothyris didyma Dalman 1828
G. †Glassina Hall and Clarke 1893
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Glassina laeviuscula Sowerby 1839
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Leptathyris index Johnson 1971
G. †Pseudoprotathyris Modzalevskaya 1979
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Pseudoprotathyris infantilis Kozlowski 1929
G. †Svetlania Baranov 1982
Subfm. †Helenathyridinae Dagys 1974
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G. †Neptunathyris Mottequin 2008
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Neptunathyris buxi Mottequin 2008
Subfm. †Janicepsinae Posenato 2001
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G. †Comelicothyris Posenato 2001
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G. †Janiceps Frech 1901
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Janiceps bipartita Stache 1878
Janiceps cadorica Stache 1878
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Invalid names: Athyris globulus Merla 1930 [synonym], Athyris humeralis Merla 1930 [synonym], Athyris irregularis Merla 1930 [synonym], Athyris sagitta Merla 1930 [synonym], Spirigera archimedis Stache 1878 [synonym], Spirigera confinalis Stache 1878 [synonym], Spirigera pusilla Stache 1878 [synonym]
Janiceps lozovskyi Grunt 1986
Janiceps ogbinensis Grunt 1965
Janiceps papilio Stache 1878
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Invalid names: Spirigera aquilina Stache 1878 [synonym]
Janiceps peracuta Stache 1878
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Invalid names: Spirifer sextensis Stache 1878 [synonym], Spirigera janiceps Stache 1878 [synonym]
G. †Kintathyris Shi and Waterhouse 1991
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Kintathyris malayaensis Shi and Waterhouse 1991
Subfm. †Lochengiinae Ching and Yang 1977
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G. †Cryptospirifer Grabau 1931
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Invalid names: Cryptospirifer iranica Nakamura and Golshani 1981 [synonym]
Cryptospirifer shawanensis Jin et al. 1974
Cryptospirifer transversa Liu and Zhao 1982
G. †Lochengia Yoh 1929
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Invalid names: Flexathyris [synonym]
G. †Titanothyris Jin and Hu 1982
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Titanothyris striata Huang 1933
Titanothyris striatus Huang 1933
Titanothyris subplicata Hu and Jin 1982
G. †Mayangella Xiong 1990
G. †Pachyplaxoides Grunt and Racki 1998
Subfm. †Plicathyridinae Alvarez 1990
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G. †Anathyris
G. †Plicathyris Khalfin 1946
Subfm. †Pradoiinae Garcia-Alcadle 1986
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G. †Guaxa
G. †Pradoia
G. †Quadriloba Alvarez et al. 1999
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Quadriloba collettii Verneuil 1850
Invalid names: Tetraloba Alvarez et al. 1998 [replaced]
Subfm. †Spirigerellinae Grunt 1965
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G. †Araxathyris Grunt 1965
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Araxathyris abichi Arthaber 1900
Araxathyris beipeiensis Xu and Grant 1994
Araxathyris bruntoni Angiolini and Carabelli 2010
Araxathyris costalis Zeng et al. 1995
Araxathyris dilatatus Shen et al. 1992
Araxathyris felina Arthaber 1900
Araxathyris glossexserta Zeng et al. 1995
Araxathyris indentatus Zeng et al. 1995
Araxathyris jalaidensis Lee and Su 1980
Araxathyris longa Zhang and Jin 1961
Araxathyris minor Wu et al. 2022
Araxathyris protea Abich 1878
Araxathyris rhombiformis Zeng et al. 1995
Araxathyris sinensis Shen et al. 2004
Araxathyris timorensis Rothpletz 1892
Araxathyris undulata Shen et al. 1992
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Invalid names: Araxathyris pentagonalis Zeng et al. 1995 [synonym]
Invalid names: Spirigera subtilita armeniaca Arthaber 1900 [synonym]
G. †Composita Brown 1849
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Composita affinis Girty 1909
Composita apheles Cooper and Grant 1976
Composita apsidata Cooper and Grant 1976
Composita argentea Shepard 1838
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Invalid names: Terebratula titicacensis Gabb 1877 [synonym]
Composita bamberi Shi and Waterhouse 1996
Composita biforma Yang 1991
Composita bucculenta Cooper and Grant 1976
Composita costata Cooper and Grant 1976
Composita cracens Cooper and Grant 1976
Composita crassa Cooper and Grant 1976
Composita depressa Sun 1991
Composita discina Cooper and Grant 1976
Composita elongata Dunbar and Condra 1932
Composita emarginata Girty 1909
Composita enormis Cooper and Grant 1976
Composita gigantea Branson 1930
Composita girtyi Raymond 1911
Composita globosa Weller 1914
Composita grandis Cooper 1953
Composita hapsida Stehli and Grant 1970
Composita idahoensis Butts 2007
Composita imbricata Cooper and Grant 1976
Composita insulcata Zeng et al. 1995
Composita jogensis Yanagida and Nishikawa 1984
Composita laevis Weller 1914
Composita lewisensis Weller 1914
Composita magnicarina Campbell 1961
Composita mexicana Hall 1857
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Invalid names: Composita arizonica McKee 1938 [synonym], Composita guadalupensis Girty 1909 [synonym], Composita parva Branson 1930 [synonym]
Composita minuscula Chronic 1949
Composita mira Girty 1899
Composita misriensis Reed 1944
Composita nucella Cooper and Grant 1976
Composita obesa Easton et al. 1958
Composita ovata Mather 1915
Composita parasulcata Cooper and Grant 1976
Composita pikensis Carter 1968
Composita pilula Cooper and Grant 1976
Composita plana Branson 1930
Composita prospera Cooper and Grant 1976
Composita pyriformis Cooper and Grant 1976
Composita quadrirotunda Lee and Su 1980
Composita quantilla Cooper and Grant 1976
Composita rotunda Snider 1915
Composita sigma Gordon 1975
Composita stalagmium Cooper and Grant 1976
Composita strongyle Cooper and Grant 1976
Composita subquadrata Hall 1858
Composita subtilita Hall 1852
Composita sulcata Weller 1914
Composita tareica Chernyak 1963
Composita tetralobata Hoare 1960
Composita tobaensis Wang 1985
Composita trinuclea Hall 1856
G. †Densalvus
G. †Gruntea Shi et al. 1999
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Gruntea grunti Shi and Shen 1997
G. †Nordathyris Grunt 1977
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Nordathyris ganelini Grunt 1977
G. †Planalvus Carter 1971
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Planalvus cailliaudi Barrois 1889
Planalvus densa Hall and Clarke 1894
Planalvus gibberosa Carter 1971
Planalvus rufus Racheboeuf et al. 1994
G. †Posicomta Grunt 1986
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Posicomta advena Grant 1976
Posicomta dolabrata Grant 1976
Posicomta gundarensis Grunt 1986
Posicomta subsolana Grant 1976
Posicomta zaalaica Grunt 1986
G. †Rectambitus Xu and Grant 1994
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Rectambitus spondomarginata Shen et al. 2004
Rectambitus subpentangulata Xu and Grant 1994
Rectambitus xui Shen et al. 2004
G. †Spirigerella Waagen 1883
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Spirigerella alata Waagen 1883
Spirigerella alataeformis Schréter 1963
Spirigerella bellistriata Zeng et al. 1995
Spirigerella derbyi Waagen 1883
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Spirigerella discors Reed 1944
Spirigerella fusiformis Waagen 1883
Spirigerella gigas Reed 1944
Spirigerella grandis Davidson 1862
Spirigerella media Waagen 1883
Spirigerella minuta Waagen 1883
Spirigerella numismalis Waagen 1883
Spirigerella obovata Reed 1944
Spirigerella ovoidalis Waagen 1883
Spirigerella pertumida Diener 1897
Spirigerella praelonga Waagen 1883
Spirigerella proxima Reed 1944
Spirigerella subtriquetra Jin et al. 1974
Spirigerella teseroi Chen 2006
Invalid names: Athyrella Renz 1912 [objective synonym]
G. †Tongzithyris Jin et al. 1974
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Tongzithyris episulcata Jin et al. 1974
Tongzithyris extensa Liao 1980
G. †Tarimathyris Chen and Shi 2006
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Tarimathyris discusella Xu and Grant 1994
Tarimathyris hotanensis Chen and Shi 2006
Tarimathyris minima Ustritsky 1960
Tarimathyris ovaloides Xu and Grant 1994
Tarimathyris postambigua Ustritsky 1960
Subfm. †Transcaucasathyriinae Angiolini and Carabelli 2010
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G. †Juxathyris Liang 1990
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Juxathyris abrupta Liang 1990
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Invalid names: Araxathyris zhinanensis Liang 1990 [synonym], Composita corpuscula Liang 1990 [synonym], Juxathyris parorbicularis Liang 1990 [synonym], Juxathyris placodepressa Liang 1990 [synonym]
Juxathyris apionucula Liang 1990
Juxathyris bisulcata Liao 1980
Juxathyris doliformis Liang 1990
Juxathyris elongata Jin and Ye 1979
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Invalid names: Spirigerella gaojiaoensis Lu 1982 [objective synonym]
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Invalid names: Spirigerella simplex Liao 1987 [synonym]
Juxathyris linguformis Liang 1990
Juxathyris mongoliensis Grabau 1931
Juxathyris obesa Huang 1933
Juxathyris pinguis Tong 1978
Juxathyris rectimarginalis Shen et al. 1992
Juxathyris subcircularis Wu et al. 2018
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Invalid names: Athyris triangularis Shen et al. 1992 [synonym]
Invalid names: Athyris rhynchonelloides Jin and Ye 1979 [nomen dubium], Spirigerella disulcata Lu 1978 [nomen dubium], Spirigerella gulingensis Lu 1978 [nomen dubium], Spirigerella leshanensis Lu 1978 [nomen dubium]
G. †Transcaucasathyris Shen et al. 2004
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Transcaucasathyris kandevani Fantini Sestini and Glaus 1966
Subfm. †Xenosariinae Cooper and Grant 1976
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G. †Xenosaria Cooper and Grant 1976
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Xenosaria exotica Cooper and Grant 1976
Xenosaria tenuis Shen and Clapham 2009
Invalid names: Athyris naliensis Reed 1944 [nomen nudum], Protathyridinae Boucot et al. 1964 [empty]
Fm. †Nucleospiridae Davidson 1881
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G. †Nucleospira Hall 1859
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Nucleospira bellornata Talent 1963
Nucleospira concinna Hall 1843
Nucleospira cunctata Cooper and Grant 1976
Nucleospira hecetensis Kirk and Amsden 1952
Nucleospira lentilca Havlicek 1990
Nucleospira pisum Sowerby 1839
Nucleospira subsphaerica Johnson 1970
Nucleospira superata Easton 1962
Diagnosis
No diagnoses are available