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Psocodea (bark louse)

Insecta - Psocodea

Synonyms
Synonymy list
YearName and author
1890Psocina Scudder p. 117
1926Psocoptera Martynov
1926Parapsocida Martynov p. 1364
1926Copeognatha Tillyard
1928Psocoptera Martynov
1935Copeognatha Tillyard
1966Psocodea Hennig p. 187
1992Psocoptera Carpenter p. 191
2004Psocodea Kukalová-Peck and Lawrence
2012Psocodea Nel et al. p. 385
2013Psocodea Mockford et al.
2014Psocodea Misof et al.
2017Psocodea Prokop et al. p. 2

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
orderPsocodea
orderPsocodea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Psocodea Hennig 1966 [bark louse]
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Subor. Psocomorpha Roesler 1944
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Infraor. Archipsocetae Yoshizawa 2002
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Fm. Archipsocidae Enderlein 1903 [ancient barklouse]
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Subfm. Archipsocinae Enderlein 1903
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G. Archipsocopsis Badonnel 1948
G. Archipsocus Hagen 1882
G. †Awightipsocus Azar 2014
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†Awightipsocus megaloi Azar 2014
†Awightipsocus minimus Azar 2014
Infraor. Caeciliusetae Pearman 1936
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Fm. Amphipsocidae Pearman 1936 [hairy-winged barklouse]
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Subfm. Amphipsocinae Pearman 1936
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Tr. Kolbiini Roesler 1940
Fm. Caeciliusidae Kolbe 1880 [lizard barklouse]
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Subfm. Caeciliusinae Kolbe 1880
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Tr. Caeciliusini Kolbe 1880
Tr. Coryphacini Mockford 2000
G. †Ptenolasia Enderlein 1911
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†Ptenolasia pilosa Hagen 1882
Invalid names: Caeciliusoidea Kolbe 1880 [invalid subgroup]
G. †Cretapsocus Vishnyakova 1975
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†Cretapsocus capillatus Vishnyakova 1975
Infraor. Epipsocetae Pearman 1936
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Fm. Epipsocidae Karny 1930 [elliptical bark louse]
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G. Epipsocus Hagen 1866
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†Epipsocus avus Roesler 1943
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Invalid names: Psocus ciliatus Pictet and Hagen 1856 [replaced]
G. Mesepipsocus Badonnell 1969
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†Mesepipsocus clarus Mockford 1969
Fm. Ptiloneuridae Enderlein 1910
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G. †Brisacia Azar et al. 2009
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†Brisacia colombiensis Azar et al. 2009
G. †Eotriplocania Zhang et al. 2016
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†Eotriplocania sinica Zhang et al. 2016
G. Euplocania Enderlein 1910
Infraor. Homilopsocidea Pearman 1936
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G. †Burmesopsocus Yoshizawa 2021
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†Burmesopsocus lienhardi Yoshizawa 2021
Fm. Ectopsocidae Roesler 1940 [outer barklouse]
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G. Ectopsocus McLachlan 1899
Fm. Elipsocidae Kolbe 1882 [damp barklouse]
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Subfm. Elipsocinae Kolbe 1882
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G. Elipsocus Hagen 1866
Fm. Lachesillidae Karny 1930 [fateful barklouse]
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G. †Archaelachesis Vishnyakova 1975
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†Archaelachesis granulosa Vishnyakova 1975
Subfm. Eolachesillinae Mockford and Sullivan 1986
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Tr. Eolachesillini Mockford and Sullivan 1986
Invalid names: Graphocaeciliini Roesler 1944 [empty]
Fm. Mesopsocidae Enderlein 1903
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G. †Mesopsocoides Azar et al. 2014
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†Mesopsocoides dupei Azar et al. 2014
G. Mesopsocus Kolbe 1880
Invalid names: Elipsocoidea Kolbe 1882 [invalid subgroup], Lachesilloidea Karny 1930 [invalid subgroup], Peripsocoidea Pearman 1936 [invalid subgroup]
Infraor. Philotarsetae Yoshizawa and Johnson 2014
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Fm. Philotarsidae Pearman 1936 [loving barklouse]
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Subfm. Philotarsinae Pearman 1936
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G. Philotarsus Kolbe 1880
Fm. Pseudocaeciliidae Pearman 1936 [false lizard barklouse]
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Subfm. †Electropsocinae Roesler 1940
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G. †Electropsocus Roesler 1940
Fm. Trichopsocidae Pearman 1936 [lash-faced psocid]
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G. †Palaeopsocus Kolbe 1883
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†Palaeopsocus tener Hagen 1856
G. Trichopsocus Kolbe 1882
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†Trichopsocus maculosus Mockford 1969
Invalid names: Pseudocaecilioidea Pearman 1936 [invalid subgroup]
Infraor. Psocetae Pearman 1936
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Fm. Myopsocidae Enderlein 1901 [mouse like barklouse]
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G. Myopsocus Hagen 1866
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†Myopsocus arthuri Nel et al. 2006
Fm. Psocidae Leach 1815 [common bark louse]
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Subfm. Psocinae Leach 1815
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Tr. Psocini Leach 1815
Tr. Ptyctini Mockford 1993
G. †Wightimicropsocus Azar 2014
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G. †Wightipsocus Azar 2014
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†Wightipsocus acourti Cockerell 1921
†Wightipsocus intasi Azar 2014
†Wightipsocus rossi Azar 2014
Invalid names: Hemipsocetae Yoshizawa 2002 [empty]
Subor. Troctomorpha Roesler 1940 [book louse]
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Infraor. Amphientometae Pearman 1936
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Superfm. Amphientomoidea Kolbe 1883
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Fm. Amphientomidae Kolbe 1883 [tropical barklouse]
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Subfm. Amphientominae Kolbe 1883
G. †Arcantipsocus Azar et al. 2009
G. †Proamphientomum Vishnyakova 1975
Invalid names: Arcantipsocidae Azar et al. 2009 [synonym]
Superfm. Electrentomoidea Enderlein 1911
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Fm. Compsocidae Mockford 1967
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G. †Burmacompsocus Nel and Waller 2007
G. †Paraelectrentomopsis Azar et al. 2016
Fm. †Electrentomidae Enderlein 1911
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G. †Electrentomum Enderlein 1911
G. †Manicapsocidus Baz and Ortuño 2001
G. †Paramesopsocus Azar et al. 2008
G. †Parelectrentomum Roesler 1940
Invalid names: Paramesopsocidae Azar et al. 2008 [synonym]
G. †Libanomphientomum Choufani et al. 2011
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†Libanomphientomum nudum Choufani et al. 2011
Fm. Troctopsocidae Mockford 1967
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G. Troctopsocoides Mockford and García Aldrete 2014
G. †Scocompus Azar et al. 2014
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†Scocompus atelisus Azar et al. 2014
Fm. Manicapsocidae Mockford 1967
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G. †Azarpsocus Maheu and Nel 2019
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†Azarpsocus anjana Álvarez-Parra and Nel 2023
†Azarpsocus perreaui Maheu and Nel 2019
G. †Eomanicapsocus Nel et al. 2005
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†Eomanicapsocus melaniae Nel et al. 2005
G. †Eoprotroctopsocus Nel et al. 2005
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†Eoprotroctopsocus celinea Nel et al. 2005
G. †Palaeomanicapsocus Azar et al. 2016
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†Palaeomanicapsocus fouadi Azar et al. 2016
†Palaeomanicapsocus margoae Azar et al. 2016
G. †Paramanicapsocus Hakim et al. 2019
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†Paramanicapsocus longiantennatus Hakim et al. 2019
Infraor. Nanopsocetae Pearman 1936
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Fm. Liposcelididae Enderlein 1911 [booklouse]
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G. †Cretoscelis Grimaldi and Engel 2006
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†Cretoscelis burmitica Grimaldi and Engel 2006
Subfm. Embidopsocinae Karny 1930
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G. Belaphopsocus Badonnel 1955
G. Belaphotroctes Roesler 1943
G. Embidopsocus Hagen 1866
Subfm. Liposcelidinae Enderlein 1911
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G. Liposcelis Motschulsky 1852
Fm. Pachytroctidae Enderlein 1911 [thick barklouse]
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Subfm. †Libaneuphorinae Azar et al. 2015
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G. †Libaneuphoris Azar et al. 2015
Subfm. Tapinellinae Enderlein 1908
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G. †Atapinella Azar et al. 2015
G. †Burmipachytrocta Azar et al. 2015
G. Psylloneura Enderlein 1903
G. Tapinella Enderlein 1908
Unr. Phthiraptera Haeckel 1896 [parasitic louse]
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Superfm. Amblycera Kellogg 1896
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Fm. †Archimenoponidae Zhang et al. 2024
Fm. Menoponidae Mjoberg 1910 [bird louse]
Invalid names: Anoplura Leach 1815 [empty], Ischnocera Kellogg 1896 [empty]
Fm. Sphaeropsocidae Kolbe 1883
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G. †Asphaeropsocites Azar et al. 2010
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†Asphaeropsocites neli Azar et al. 2010
G. †Sphaeropsocites Grimaldi and Engel 2006
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†Sphaeropsocites lebanensis Grimaldi and Engel 2006
G. †Sphaeropsocoides Grimaldi and Engel 2006
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†Sphaeropsocoides canadensis Grimaldi and Engel 2006
G. Sphaeropsocus Hagen 1882
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†Sphaeropsocus kuenowii Hagen 1882
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Invalid names: Atropos succinica Hagen 1882 [synonym]
Invalid names: Palaeotroctes Enderlein 1911 [synonym]
Subor. Trogiomorpha Roesler 1940
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Infraor. Atropetae Pearman 1936
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Fm. †Archaeatropidae Baz and Ortuño 2000
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G. †Archaeatropos Baz and Ortuño 2000
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†Archaeatropos alavensis Baz and Ortuño 2000
†Archaeatropos randatae Azar and Nel 2004
G. †Bcharreglaris Azar and Nel 2004
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†Bcharreglaris amooni Kaddumi 2007
†Bcharreglaris amunobi Azar and Nel 2004
†Bcharreglaris haddadini Kaddumi 2007
G. †Heliadesdakruon Cumming and Le Tirant 2021
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†Heliadesdakruon morganae Cumming and Le Tirant 2021
G. †Libanoglaris Azar et al. 2003
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†Libanoglaris chehabi Azar and Nel 2004
†Libanoglaris hespericus Álvarez-Parra et al. 2022
†Libanoglaris mouawadi Azar et al. 2003
G. †Longiantennum Liang et al. 2022
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†Longiantennum fashengi Liang et al. 2022
G. †Proprionoglaris Perrichot et al. 2003
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†Proprionoglaris axioperierga Azar et al. 2014
†Proprionoglaris guyoti Perrichot et al. 2003
G. †Prospeleketor Perrichot et al. 2003
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†Prospeleketor albianensis Perrichot et al. 2003
G. †Setoglaris Azar and Nel 2004
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†Setoglaris reemae Azar and Nel 2004
Fm. †Empheriidae Kolbe 1883
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G. †Burmempheria Li et al. 2020
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†Burmempheria curvatavena Li et al. 2022
†Burmempheria densuschaetae Li et al. 2020
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Invalid names: Latempheria kachinensis Li et al. 2022 [synonym]
†Burmempheria raruschaetae Li et al. 2020
Invalid names: Latempheria Li et al. 2022 [synonym]
G. †Empheria Hagen 1856
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Subg. †Empheria (Bebiosis) Enderlein 1911
†Empheria reticulata Hagen 1856
G. †Empherium Hakim et al. 2021
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†Empherium rasnitsyni Hakim et al. 2021
G. †Empheropsocus Baz and Ortuño 2001
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†Empheropsocus arilloi Baz and Ortuño 2001
†Empheropsocus margineglabrus Baz and Ortuño 2001
G. †Eoempheria Nel et al. 2005
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†Eoempheria intermedia Nel et al. 2005
G. †Jerseyempheria Azar et al. 2010
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†Jerseyempheria grimaldii Azar et al. 2010
G. †Paralellopsocus Hakim et al. 2023
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†Paralellopsocus elongatus Hakim et al. 2023
G. †Preempheria Baz and Ortuño 2001
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†Preempheria antiqua Baz and Ortuño 2001
G. †Santonipsocus Álvarez-Parra et al. 2024
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†Santonipsocus mimeticus Álvarez-Parra et al. 2024
G. †Trichempheria Enderlein 1911
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†Trichempheria villosa Hagen 1882
Fm. Lepidopsocidae Enderlein 1903 [scaly-winged bark louse]
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Subfm. Perientominae Kolbe 1882
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G. †Nepticulomima Enderlein 1906
G. Perientomum Hagen 1865
Subfm. Thylacinae Enderlein 1903
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G. Thylacella Enderlein 1911
G. †Thylax Hagen 1866
Invalid names: Thylacellinae Roesler 1944 [synonym]
Fm. Psoquillidae Kolbe 1882
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G. †Eorhyopsocus Nel et al. 2005
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†Eorhyopsocus magnificus Nel et al. 2005
Fm. Trogiidae Enderlein 1911 [granary booklouse]
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G. †Cretolepinotus Cockx et al. 2020
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†Cretolepinotus tankei Cockx et al. 2020
G. †Eolepinotus Vishnyakova 1975
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†Eolepinotus pilosus Vishnyakova 1975
†Eolepinotus zherikhini Hakim et al. 2021
G. †Paralepinotus Azar et al. 2018
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†Paralepinotus fushunensis Azar et al. 2018
G. †Brachyantennum Liang and Liu 2022
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†Brachyantennum spinosum Liang and Liu 2022
Fm. †Cormopsocidae Yoshizawa and Lienhard 2020
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G. †Cormopsocus Yoshizawa and Lienhard 2020
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†Cormopsocus baleoi Hakim et al. 2021
†Cormopsocus groehni Yoshizawa and Lienhard 2020
†Cormopsocus neli Hakim et al. 2021
†Cormopsocus perantiqua Cockerell 1919
G. †Longiglabellus Wang et al. 2021
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†Longiglabellus edentatus Wang et al. 2021
†Longiglabellus pedhyalinus Wang et al. 2021
G. †Stimulopsocus Liang and Liu 2021
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†Stimulopsocus jiewenae Liang and Liu 2021
G. †Empheriopsis Vishnyakova 1975
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†Empheriopsis vulnerata Vishnyakova 1975
G. †Parapsyllipsocus Perrichot et al. 2003
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†Parapsyllipsocus vergereaui Perrichot et al. 2003
Infraor. Prionoglaridetae Smithers 1972
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Fm. Prionoglarididae Karny 1930
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Subfm. Prionoglaridinae Karny 1930
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Tr. Siamoglaridini Azar et al. 2017
Infraor. Psyllipsocetae Smithers 1972
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Fm. Psyllipsocidae Kolbe 1884 [cave barklouse]
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G. †Annulipsyllipsocus Hakim et al. 2017
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†Annulipsyllipsocus andreneli Hakim et al. 2017
†Annulipsyllipsocus inexspectatus Hakim et al. 2017
G. †Concavapsocus Wang et al. 2019
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†Concavapsocus parallelus Wang et al. 2019
G. †Globopsocus Azar and Engel 2008
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†Globopsocus aquilonius Azar and Engel 2008
G. †Khatangia Vishnyakova 1975
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†Khatangia inclusa Vishnyakova 1975
G. †Libanopsyllipsocus Azar and Nel 2011
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G. Psyllipsocus Selys-Longchamps 1872
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†Psyllipsocus eocenicus Nel et al. 2005
†Psyllipsocus myanmarensis Jouault et al. 2021
†Psyllipsocus yangi Liang and Liu 2021
†Psyllipsocus yoshizawai Álvarez-Parra et al. 2020
G. †Sinopsyllipsocus Zhang et al. 2016
+
†Sinopsyllipsocus fushunensis Zhang et al. 2016
Invalid names: Psocatropetae Pearman 1936 [invalid subgroup]
Invalid names: Psocoptera [replaced]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Hettangian to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 190.8 Ma

Collections (106 total)


Time interval Ma Country or state Original ID and collection number
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Psococicadellopsis primitiva (105087)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Paramesopsocus adibi (31311 105426 105970 105971 106101 106102 113439 114238)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) Pseudopsocus parvus (154730)
Early/Lower Barremian130.0 - 125.45Lebanon Asphaeropsocites neli, Libanoglaris chehabi, Setoglaris reemae, Paramesopsocus lu, Libanoglaris mouawadi, Libanopsyllipsocus alexanderasnitsyni, Libanomphientomum nudus, Palaeosiamoglaris hammanaensis (122471) Bcharreglaris amunobi (123017) Libaneuphoris jantopi (146535) Libanoglaris randatae (123000) Sphaeropsocites lebanensis, Libanoglaris randatae (122359)
Aptian125.0 - 113.0Mongolia (Govi-Altai) Psocoptera indet. (136020)
Albian113.0 - 100.5Jordan Psocodea indet., Troctomorpha indet., Trogiomorpha indet., Bcharreglaris haddadini, Bcharreglaris amooni (124847)
Early/Lower Albian112.03 - 109.0Spain (Teruel) Psocomorpha indet., Libanoglaris hespericus, Azarpsocus anjana (225779)
Early/Lower Albian112.03 - 109.0Myanmar (Sagaing) Palaeosiamoglaris hkamtiensis (215115)
Early/Lower Albian112.03 - 109.0Spain Archaeatropos alavensis, Libanoglaris hespericus, Azarpsocus anjana, Burmacompsocus ojancano (109438)
Late/Upper Albian105.3 - 99.6France (Poitou-Charentes) Arcantipsocus courvillei, Proprionoglaris guyoti, Prospeleketor albianensis, Parapsyllipsocus vergereaui, Santonipsocus mimeticus (99114)
Late/Upper Albian105.3 - 99.6Spain (Álava) Empheropsocus arilloi, Empheropsocus margineglabrus, Preempheria antiqua, Archaeatropos alavensis, Manicapsocidus enigmaticus (108769)
Late/Upper Albian105.3 - 99.6Spain (Aragón) Preempheria antiqua, Archaeatropidae indet., Archaeatropos alavensis (91626)
Late/Upper Albian105.3 - 99.6Spain (Teruel) Archaeatropos alavensis (225780)
Late/Upper Albian - Early/Lower Cenomanian105.3 - 93.5France (Charente-Maritime) Psocoptera indet. (107076)
Cenomanian100.5 - 93.9Russian Federation Eolepinotus zherikhini (226224)
Early/Lower Cenomanian99.6 - 93.5Myanmar Archimenopon myanmarensis (178121) Azarpsocus perreaui, Burmacompsocus pouilloni (189531) Burmacompsocus coniugans (189783) Burmacompsocus perreaui (123055) Burmesopsocus lienhardi (222694) Cormopsocus groehni (172900) Heliadesdakruon morganae (220289) Longiantennum fashengi, Concavapsocus parallelus, Burmempheria densuschaetae, Burmempheria raruschaetae, Longiglabellus pedhyalinus, Longiglabellus edentatus, Latempheria kachinensis, Burmempheria curvatavena (165561) Longiantennum fashengi, Psyllipsocus yangi, Brachyantennum spinosum, Paramanicapsocus xingyuei (196491) Psocodea morphotype PI (212006) Psocodea morphotype PI, Psocodea morphotype PII, Psocodea morphotype PIII, Longiantennum fashengi, Burmipachytrocta singularis, Atapinella garroustei, Paraelectrentomopsis chenyangcaii, Palaeomanicapsocus margoae, Palaeomanicapsocus fouadi, Palaeosiamoglaris lienhardi, Palaeosiamoglaris burmica, Palaeosiamoglaris inexpectata, Annulipsyllipsocus andreneli, Annulipsyllipsocus inexspectatus, Paramanicapsocus longiantennatus, Burmempheria densuschaetae, Burmempheria raruschaetae, Psyllipsocus myanmarensis, Cormopsocus neli, Cormopsocus baleoi, Paralellopsocus elongatus (161726) Psocoptera indet., Cretoscelis burmitica (111688) Psyllipsocus banksi, Psylloneura perantiqua (123020) Psyllipsocus myanmarensis (179227) Psyllipsocus yoshizawai (205558) Stimulopsocus jiewenae (226881)
Early/Lower Cenomanian99.6 - 93.5France Psocoptera indet. (123019)
Late/Upper Cenomanian99.6 - 93.5Russian Federation Globopsocus aquilonius, Empherium rasnitsyni (35051)
Turonian93.9 - 89.8USA (New Jersey) Jerseyempheria grimaldii (110895)
Turonian93.9 - 89.8Botswana Psocoptera indet. (34851)
Turonian93.9 - 89.8France Psocodea indet., Mesopsocidae indet., Proprionoglaris axioperierga, Scocompus atelisus, Mesopsocoides dupei (164386)
Late/Upper Santonian85.8 - 83.5Russian Federation Psocomorpha indet. 1, Cretapsocus capillatus (122590) Psocoptera indet. 1, Psocoptera indet. 2, Psocoptera indet. 3, Psocomorpha indet. 2, Psocomorpha indet. 3, Eolepinotus pilosus, Khatangia inclusa, Proamphientomum cretaceum, Archaelachesis granulosa, Cretapsocus capillatus, Empheriopsis vulnerata (35056)
Late/Upper Campanian83.5 - 70.6Canada (Alberta) Cretolepinotus tankei (207633)
Late/Upper Campanian83.5 - 70.6Myanmar Lepidopsocidae indet. (195455)
Middle Campanian83.5 - 70.6Canada (Alberta) Sphaeropsocoides canadensis (122383)
Ypresian56.0 - 47.8Canada (British Columbia) Psocodea sp. (123795)
Ypresian56.0 - 47.8France (Oise) Psyllipsocus eocenicus, Thylacella eocenica, Eoempheria intermedia, Eorhyopsocus magnificus, Embidopsocus eocenicus, Tapinella eocenica, Eomanicapsocus melaniae, Amphientomum parisiensis, Eolachesilla eocenica, Archipsocus puber, Eoprotroctopsocus celinea, Parathylacella oisensis (123850)
Ypresian56.0 - 47.8China (Liaoning) Eopsocites fushunensis, Stenopterites guchengziensis, Stenopterites longus, Fushunopsocus latus (124820) Sinopsyllipsocus fushunensis, Eotriplocania sinica, Paralepinotus fushunensis (125934)
Bridgerian50.3 - 46.2USA (Utah) Paropsocus disjunctus (121147)
MP 1348.6 - 40.4Germany (Eifel) Megamenopon rasnitsyni (135640)
Priabonian38.0 - 33.9Poland Psocus picteti, Psocus affinis, Psocus ciliatus, Psocus debilis, Psocus proavus, Psocus tener, Psocus abnormis, Amphientomum paradoxum (109625)
Priabonian38.0 - 33.9Germany (Halle) Embidopsocus saxonicus (129227)
Priabonian38.0 - 33.9United Kingdom (Isle of Wight) Wightipsocus acourti, Wightipsocus intasi, Awightipsocus jarzembowskii, Awightipsocus megaloi, Awightipsocus minimus, Wightimicropsocus inexpectatus (157666)
Priabonian38.0 - 33.9United Kingdom (England) Psocus acourti, Wightipsocus intasi, Awightipsocus jarzembowskii, Awightipsocus megaloi, Awightipsocus minimus (123960) Wightipsocus intasi, Awightipsocus megaloi (21976) Wightipsocus intasi, Wightipsocus rossi, Awightipsocus jarzembowskii (123959)
Priabonian38.0 - 33.9Ukraine Archipsocus puber, Trichempheria villosa, Sphaeropsocus kuenowii, Psocidus multiplex (113621) Sphaeropsocus kuenowii (122996) Sphaeropsocus kuenowii, Epipsocus avus (122997) Valenzuela prometheus (122999)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Archipsocus puber, Psocus sparsipennis, Psocus trigonoscenea, Psocus electricus, Psocus picteti, Copostigma affinis, Epipsocus ciliatus, Kolbea ava, Caecilius debilis, Caecilius scenepipedus, Caecilius proavus, Caecilius prometheus, Caecilius klebsi, Caecilius sucinicaptus, Ptenolasia pilosa, Elipsocus abnormis, Philotarsus antiquus, Philotarsus bullicornis, Amphientomum paradoxum, Amphientomum leptolepis, Amphientomum colpolepis, Electrentomum klebsianum, Empheria reticulata, Bebiosis pertinens, Sphaeropsocus kunowi (124073) Archipsocus puber, Psocus trigonoscenea, Psocus picteti, Psocus affinis, Epipsocus ciliatus, Caecilius debilis, Caecilius proavus, Caecilius prometheus, Caecilius pilosus, Elipsocus abnormis, Amphientomum paradoxum, Amphientomum leptolepis, Amphientomum colpolepis, Empheria reticulata, Empheria villosa, Atropos succinica, Sphaeropsocus kuenowii, Atropos succinica (123962) Caecilius debilis (123993) Epipsocus ciliatus, Caecilius debilis, Caecilius proavus, Caecilius prometheus, Amphientomum paradoxum (124088) Parelectrentomum priscum, Electropsocus unguidens (124903) Psocus picteti (124493) Psocus picteti, Caecilius proavus (124494) Psocus trigonoscenea, Psocus picteti, Copostigma affinis, Caecilius proavus, Elipsocus kuhli, Philotarsus antiquus, Amphientomum paradoxum, Palaeotroctes succinicus (123221) Sphaeropsocus kuenowii, Embidopsocus pankowskiorum (124370)
Orellan33.9 - 33.3USA (Colorado) Psocoptera indet. (149289)
Early/Lower Miocene23.03 - 15.97Mexico Psyllipsocus sp., Archipsocus sp. 1, Archipsocus sp. 2, Archipsocus sp. 3, Belaphotroctes sp., Myopsocus sp. (21977)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Amphientomum elongatum, Epipsocus clarus (124470) Amphientomum elongatum, Myopsocidae indet. (124623) Archipsocus antiguus (124472) Archipsocus sp., Archipsocus antiguus (124471) Belaphotroctes similis (124469) Ectopsocus sp. (124473) Liposcelis sp. (124810) Psocomorpha indet. (124474) Psyllipsocus sp. (124467) Trichopsocus maculosus (140313)
Early/Lower Miocene23.03 - 15.97Spain (Castellon) Mesopsocus sp. (140798)
Early/Lower Miocene23.03 - 15.97Ethiopia Psocoptera indet. (151981 151982)
Burdigalian20.44 - 15.97New Zealand (Otago) Pseudocaeciliidae indet. (191650)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Belaphopsocus dominicus (122982) Myopsocus arthuri (124475 227086) Troctopsocoides gracilis (162585)
Hemingfordian20.43 - 15.97USA (California) Miotroctes rousei (144521) Miotroctes sp. (106431) Trogiidae indet. (138761)
Langhian15.97 - 13.82China (Fujian) Belaphotroctes grimaldii (220273)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Phthiraptera indet. (169336)
Holocene0.0117 - 0.0Tanzania Amphientomum incultum, Perientomum mortuum, Thylax fimbriatum, Atropos resinata (124495) Liposcelididae indet., Amphientomum knorrei (234540) Thylacella eversiana (140788)
Holocene0.0117 - 0.0Colombia (Santander) Brisacia colombiensis, Euplocania sp. (151348)