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Geoemydidae
Taxonomy
Geoemydidae was named by Theobald (1868). It is extant. It was considered monophyletic by Joyce (2007) and Vlachos (2018).
It was reranked as the unranked clade Geoemydidae by Cadena et al. (2012).
It was assigned to Testudines by Hutchison (2006); to Cryptodira by Cadena et al. (2012); to Cryptodira by Vlachos et al. (2015), Vlachos and Delfino (2016) and Cadena et al. (2017); to Pangeoemydidae by Joyce (2007) and Vlachos (2018); and to Testudinoidea by Claude et al. (2012), Takahashi et al. (2013), Vremir (2013), van Dijk et al. (2014), de la Fuente et al. (2018), Collareta et al. (2020), Vlachos (2021) and Perez-Garcia et al. (2021).
It was reranked as the unranked clade Geoemydidae by Cadena et al. (2012).
It was assigned to Testudines by Hutchison (2006); to Cryptodira by Cadena et al. (2012); to Cryptodira by Vlachos et al. (2015), Vlachos and Delfino (2016) and Cadena et al. (2017); to Pangeoemydidae by Joyce (2007) and Vlachos (2018); and to Testudinoidea by Claude et al. (2012), Takahashi et al. (2013), Vremir (2013), van Dijk et al. (2014), de la Fuente et al. (2018), Collareta et al. (2020), Vlachos (2021) and Perez-Garcia et al. (2021).
Subtaxa
Bridgeremys, Clemmydopsis, Cuora, Cyclemys, Echmatemys, Geiselemys, Geoemyda, Geoemydinae, Guangdongemys, Isometremys, Mauremys (syn. Paralichelys), Ocadia, Orlitia, Palaeochelyinae, Palaeochelys, Palatochelydia, Ptychogasterinae, Ptychogastrini, Rhinoclemmydinae, Rhinoclemmys, Sacalia, Siebenrockiella (syn. Bellia)
Synonymy list
| Year | Name and author |
|---|---|
| 1868 | Geoemydidae Theobald |
| 2006 | Geoemydidae Hutchison |
| 2007 | Geoemydidae Joyce |
| 2012 | Geoemydidae Cadena et al. p. 540 |
| 2012 | Geoemydidae Claude et al. |
| 2013 | Geoemydidae Takahashi et al. p. 529 |
| 2013 | Geoemydidae Vremir |
| 2014 | Geoemydidae van Dijk et al. |
| 2015 | Geoemydidae Vlachos et al. |
| 2016 | Geoemydidae Vlachos and Delfino |
| 2017 | Geoemydidae Cadena et al. |
| 2018 | Geoemydidae Vlachos |
| 2018 | Geoemydidae de la Fuente et al. |
| 2020 | Geoemydidae Collareta et al. |
| 2021 | Geoemydidae Perez-Garcia et al. |
| 2021 | Geoemydidae Vlachos |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Geoemydidae Theobald 1868
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G. †Bridgeremys Hutchison 2006
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G. †Clemmydopsis Boda 1927
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†Clemmydopsis steinheimensis Staesche 1931
†Clemmydopsis turnauensis Meyer 1847
Invalid names: Chrysemys bisbalensis Bergounioux 1958 [nomen dubium]
G. Cuora Gray 1855 [Asian box turtle]
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Cuora amboinensis Riche 1801 [Southeast Asian box turtle]
Cuora aurocapitata Luo and Zong 1988 [Yellow-headed box turtle]
Cuora bourreti Obst and Reimann 1994 [Bourrets box turtle]
Cuora flavomarginata Gray 1863 [Yellow-margined box turtle]
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Invalid names: Terrapene culturalia Yeh 1961 [synonym]
Cuora galbinifrons Bourret 1939 [Indochinese box turtle]
Cuora mccordi Ernst 1988 [McCord's box turtle]
Cuora mouhotii Gray 1862 [Keeled box turtle]
Cuora pani Song 1984 [Pan's box turtle]
Cuora picturata Lehr et al. 1998 [Southern Vietnam box turtle]
†Cuora pitheca Yeh 1981
Cuora trifasciata Bell 1825 [Chinese three-striped box turtle]
†Cuora tungia Yeh 1963
Cuora yunnanensis Boulenger 1906 [Yunnan box turtle]
Cuora zhoui Zhao 1990 [Zhou's box turtle]
G. Cyclemys Bell 1834 [Asian leaf turtle]
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Cyclemys atripons Iverson and McCord 1997
Cyclemys dentata Gray 1831
Cyclemys enigmatica Fritz et al. 2008
Cyclemys fusca Fritz et al. 2008
Cyclemys gemeli Fritz et al. 2008
†Cyclemys miyatai Shikama 1949
Cyclemys oldhamii Gray 1863
Cyclemys pulchristriata Fritz et al. 1997
G. †Echmatemys Hay 1906
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†Echmatemys arethusa Hay 1908
†Echmatemys callopyge Hay 1908
†Echmatemys cibollensis Cope 1877
†Echmatemys haydeni Leidy 1870
†Echmatemys jeanesi Leidy 1870
†Echmatemys lativertebralis Cope 1877
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Invalid names: Chrysemys inornata Loomis 1904 [synonym], Echmatemys douglassi Gilmore 1915 [synonym], Echmatemys hollandi Gilmore 1915 [synonym], Echmatemys rivalis Hay 1909 [synonym], Emys shaughnessiana Cope 1882 [synonym], Graptemys cordifera Clark 1937 [synonym]
†Echmatemys stevensoniana Leidy 1870
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Invalid names: Echmatemys aegle Hay 1908 [synonym], Echmatemys cyane Hay 1908 [synonym], Echmatemys ocyrrhoe Hay 1908 [synonym], Emys septarius Cope 1873 [synonym]
Invalid names: Echmatemys depressa Gilmore 1915 [nomen dubium], Palaeotheca polycypha Cope 1872 [nomen dubium], Palaeotheca terrestris Cope 1872 [nomen dubium]
G. †Geiselemys Khosatzky and Mlynarski 1966
G. Geoemyda Gray 1834
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Subg. †Geoemyda (Geoliemys) Matsumoto 1929
†Geoemyda amamiensis Takahashi et al. 2007
†Geoemyda eureia Wegner 1913
†Geoemyda headonensis Hooley 1905
Geoemyda japonica Fan 1931
†Geoemyda malustensis Macarovici and Vanca 1960
†Geoemyda mossoczyi Mlynarski 1964
†Geoemyda pidoplickai Khosatzky 1946
†Geoemyda ptychogastroides Hummel 1935
Geoemyda spengleri Gmelin 1789
†Geoemyda striata Deraniyagala 1969
Subfm. Geoemydinae Theobald 1868
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G. Heosemys Gray 1831 [Forest turtles]
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Heosemys annandalii Boulenger 1903 [Yellow-headed temple turtle]
Heosemys depressa Anderson 1875 [Arakan forest turtle]
Heosemys grandis Gray 1860 [Giant Asian pond turtle]
Heosemys leytensis Tayor 1920 [Philippine pond turtle]
Heosemys spinosa Gray 1831 [Spiny turtle]
G. Leucocephalon McCord et al. 2000 [Sulawesi forest turtle]
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Leucocephalon yuwonoi McCord et al. 1995 [Sulawesi forest turtle]
G. Malayemys Lindholm 1931 [Malayan snail-eating turtles]
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Malayemys macrocephala Gray 1859 [Malayan snail-eating turtle]
Malayemys subtrijuga Schlegel and Muller 1844 [Malayan snail-eating turtle]
G. Melanochelys Schweigger 1812 [Indian black turtles]
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†Melanochelys sivalensis Lydekker 1885
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Invalid names: Bellia theobaldi Lydekker 1885 [synonym], Clemmys hydaspica Lydekker 1885 [synonym], Clemmys punjabiensis Lydekker 1885 [synonym]
†Melanochelys tapani Garbin et al. 2020
Melanochelys tricarinata Blyth 1856 [Tricarinate hill turtle]
Melanochelys trijuga Schweigger 1812 [Indian black turtle]
G. Morenia Gray 1870 [Eyed turtles]
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Morenia ocellata Dumeril and Bibron 1835 [Burmese eyed turtle]
Morenia petersi Anderson 1879 [Indian eyed turtle]
G. Vijayachelys Praschag et al. 2006 [Cochin forest cane turtle]
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Vijayachelys silvatica Henderson 1912 [Cochin forest cane turtle]
G. Mauremys Gray 1870 [pond turtle]
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Mauremys annamensis Siebenrock 1903 [Vietnamese pond turtle]
†Mauremys aristotelica Vlachos et al. 2019
Mauremys caspica Gmelin 1774 [Caspian turtle]
†Mauremys fenhoense Gray 1931
†Mauremys gaudryi Depéret 1885
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Invalid names: Clemmys romani Bergounioux 1935 [synonym], Emys etrusca Portis 1890 [synonym], Paralichelys carinata Bergounioux 1935 [synonym]
†Mauremys guntiana Roger 1902
†Mauremys isoclina Dubois 1908
Mauremys japonica Temminck and Schlegel 1835 [Japanese pond turtle]
Mauremys leprosa Schweigger 1812 [Mediterranean pond turtle]
†Mauremys massiliensis Bergounioux 1936
Mauremys mutica Cantor 1842 [yellow pond turtle]
Mauremys nigricans Gray 1834 [Chinese red-necked turtle]
†Mauremys portisi Sacco 1889
†Mauremys pygolopha Peters 1868
Mauremys rivulata Valenciennes 1833 [Western Caspian turtle]
†Mauremys sarmatica Purschke 1885
Mauremys sinensis Gray 1834 [Chinese stripe-necked turtle]
†Mauremys sophiae Ammon 1911
†Mauremys thanhinensis Claude et al. 2007
†Mauremys yabei Shikama 1949
Invalid names: Clemmys precaspica de Stefano 1902 [nomen dubium], Emys prosigriz Pallary 1901 [nomen dubium], Paralichelys Bergounioux 1935 [synonym]
G. Ocadia Gray 1870
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†Ocadia nicoleti Pictet and Humbert 1856
†Ocadia nipponica Hirayama et al. 2007
†Ocadia perplexa Gilmore 1931
†Ocadia tanegashimensis Takahashi et al. 2013
†Ocadia turgaica Kuznetsov and Chkhikvadze 1977
G. †Palaeochelys Meyer 1847
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†Palaeochelys astrei Bergounioux 1935
†Palaeochelys bussenensis Meyer 1847
†Palaeochelys elongata Gilmore 1931
†Palaeochelys gabuniai Chkhikvadze 1973
Invalid names: Palaeochelys haslachensis Meyer 1851 [nomen nudum]
Unr. Palatochelydia Joyce and Lyson 2010
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G. Batagur Gray 1855
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Batagur affinis Cantor 1847 [Southern River terrapin]
Batagur baska Gray 1831 [River terrapin]
Batagur borneoensis Schlegel and Muller 1845 [Painted terrapin]
Batagur dhongoka Gray 1834 [Three-striped roofed turtle]
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Invalid names: Batagur durandi Lydekker 1885 [synonym]
Batagur kachuga Gray 1831 [Red-crowned roofed turtle]
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Invalid names: Batagur bakeri Lydekker 1885 [synonym], Batagur ellioti Gray 1962 [synonym]
Batagur trivittata Duméril and Bibron 1835 [Burmese roofed turtle]
Invalid names: Batagur cautleyi Lydekker 1885 [nomen dubium]
G. Geoclemys Gray 1855
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†Geoclemmys matuuraensis Shikama 1956
†Geoclemmys yudaensis Shikama 1956
Geoclemys hamiltonii Gray 1831 [Spotted pond turtle]
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Invalid names: Clemmys palaeindica Lydekker 1885 [synonym], Geoclemys sivalensis Tewari and Badam 1969 [synonym]
†Geoclemys matuuraensis Shikama 1956
G. Hardella Gray 1870
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†Hardella siamensis Claude et al. 2007
Hardella thurjii Gray 1831
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Invalid names: Batagur falconeri Lydekker 1885 [synonym], Clemmys watsoni Lydekker 1886 [synonym]
G. Pangshura Gray 1856
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Pangshura smithii Gray 1863
Pangshura sylhetensis Jerdon 1870
†Pangshura tatrotia Joyce and Lyson 2010
Pangshura tecta Gray 1830
Pangshura tentoria Gray 1834
Subfm. †Ptychogasterinae De Stefano 1903
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G. †Ptychogaster Pomel 1847
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Subg. †Ptychogaster (Temnoclemmys) Bergounioux 1957
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†Ptychogaster (Temnoclemmys) bardenensis Murelaga et al. 1999
†Temnoclemmys mazedonica Pasic and Klincarski 1959
†Ptychogaster bardenensis Murellaga et al. 1999
†Ptychogaster batalleri Bergounioux 1931
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Invalid names: Emys egarensis Bataller 1967 [synonym], Temnoclemmys elongata Bergounioux 1957 [synonym], Temnoclemmys vallesensis Bergounioux 1957 [synonym], Testudo celonica Bergounioux 1958 [synonym]
†Ptychogaster cayluxensis Lydekker 1889
†Ptychogaster emydoides Pomel 1847
†Ptychogaster kinkelini Reinach 1900
†Ptychogaster pomeli Lydekker 1889
†Ptychogaster ronheimensis Groessens-Van Dyck and Schleich 1985
Tr. †Ptychogastrini De Stefano 1903
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Subfm. Rhinoclemmydinae Gray 1873
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G. Rhinoclemmys Fitzinger 1835
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Rhinoclemmys annulata Gray 1860
Rhinoclemmys areolata Dumeril and Bibron 1851
Rhinoclemmys diademata Mertens 1954
Rhinoclemmys funerea Cope 1876
Rhinoclemmys melanosterna Gray 1861
Rhinoclemmys nasuta Boulenger 1902
†Rhinoclemmys panamaensis Cadena et al. 2012
Rhinoclemmys pulcherrima Gray 1856
Rhinoclemmys punctularia Daudin 1801
Rhinoclemmys rubida Cope 1870
G. Sacalia Gray 1870 [eyed turtle]
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Sacalia bealei Gray 1831
Sacalia quadriocellata Siebenrock 1903
G. Siebenrockiella Lindholm 1929 [black marsh turtle]
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Siebenrockiella crassicollis Gray 1830
Siebenrockiella leytensis Taylor 1920
Invalid names: Bellia Gray 1869 [replaced]
Invalid names: Palaeochelyinae Karl and Wettlaufer 2011 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: subo = suborder, c = class, subp = subphylum | |||||
| References: Uetz 2005, Carroll 1988, Hendy et al. 2009 | |||||
Age range
Maximum range based only on fossils: base of the Lostcabinian to the top of the Holocene or 55.80000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 50.3 Ma
Minimum age of oldest fossil (stem group age): 50.3 Ma
Collections (455 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Ypresian | France (Oise) | Palaeochelys sp. (68174) | |
| Ypresian | France (Champagne Ardenne) | Palaeochelys sp. (177753) | |
| Ypresian | France (Marne) | Merovemys ploegi (35533) Palaeochelys sp. (35535 35540 181653) | |
| Ypresian - Lutetian | France (Occitanie) | Hummelemys sp. (195457) | |
| Ypresian - Lutetian | France (Languedoc-Roussillon Region) | Chelonia indet. (31717) | |
| Lostcabinian | USA (Wyoming) | Echmatemys wyomingensis (15997) | |
| Lysitean | USA (Colorado) | Echmatemys stevensoniana (15654) | |
| Wasatchian | USA (New Mexico) | Echmatemys lativertebralis (15443 176872 176873 176877 176879 176880 176882 176885 176886 176887 176888 176890 176891 176894 176898 176899 176900 176902 176909 176911 176912 176914) Emys cibollensis, Emys lativertebralis (131867) | |
| Wasatchian | USA (Colorado) | Echmatemys lativertebralis (176837) | |
| Wasatchian | USA (Wyoming) | Echmatemys lativertebralis (46718) Echmatemys rivalis (177320) Echmatemys sp. (15458 15563 15566 15587 15759 46650 46772 59904 59905) | |
| MP 10 | France (Picardie) | Palaeochelys sp. (68440) | |
| Wasatchian - Bridgerian | USA (New Mexico) | Echmatemys sp. (25762) | |
| Wasatchian - Bridgerian | USA (Wyoming) | Echmatemys sp. (16134) | |
| Bridgerian | USA (Colorado) | Echmatemys stevensoniana (16242) | |
| Bridgerian | USA (Wyoming) | Bridgeremys pusilla (16243 16418 168275 168278 168279 168280 168281 176700 176701 176702 176703 176704 176705 176706 176707 176708 176709 176710 176711 176712 176713 176714 176715 176716 176717 176718 176719 176720 176721 176722 176723 176724 176725 176726 176727 176728 176729 176730 176731 176732 176733 176735) Clemmys morrisiae, Echmatemys pusilla, Echmatemys arethusa, Echmatemys aegle, Echmatemys cyane (16228) Echmatemys callopyge, Echmatemys wyomingensis (74504) Echmatemys cibollensis (70264) Echmatemys naomi (16255 206092) Echmatemys ocyrrhoe (16201) Echmatemys septaria (16261 16262 16270 16273) Echmatemys septaria, Echmatemys wyomingensis (15837 16265 16267) Echmatemys septaria, Rhinoclemys terrestris (16405) Echmatemys sp. (16209 16296) Emys haydeni, Palaeotheca polycypha, Palaeotheca terrestris, Emys shaugnessiana (130809) Emys jeanesi, Emys haydeni, Emys stevensonianus (128447) Emys septarius (191004) Rhinoclemys terrestris, Echmatemys septaria, Bridgeremys pusilla (55207) | |
| Bridgerian | USA (Utah) | Echmatemys sp. (16396) | |
| MP 11 | Germany (Hessen) | Geoemydidae indet. (120798) | |
| MP 13 | Germany (Eifel) | Geiselemys ptychogastroides (24201) | |
| Lutetian | France ( Île-de-France) | Hummelemys sp. (167224) | |
| Lutetian | France (Picardy) | Hummelemys sp. (167226) | |