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Scarabaeoidea (scarab beetle)

Insecta - Coleoptera

Synonyms
Synonymy list
YearName and author
1802Scarabaeoidea Latreille p. 144
1847Onthophagus urus Heer p. 62 figs. Pl 2, fig 10
1862Gymnopleurus deperditus Heer p. 73 figs. Pl 6, fig 9
1862Copris subterranea Heer p. 74 figs. Pl 3, fig 4
1862Onthophagus urus Heer p. 76
1981Scarabaeoidea Crowson p. 695
1992Holcorobeini Nikolajev p. 102
1995Scarabaeoidea Lawrence and Newton, Jr.
2000Onthophagus urusheeri Krell p. 880
2000Holcorobeini Krell p. 889
2004Holcorobeini Nikolajev p. 35
2007Holcorobeini Nikolajev p. 142
2011Holcorobeini Bouchard et al.
2011Scarabaeoidea Bouchard et al.
2022Scarabaeoidea Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
RankNameAuthor
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyScarabaeoidea
superfamilyScarabaeoidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
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G. †Aliscarabaeus Hong 1984
+
Fm. †Alloioscarabaeidae Bai et al. 2012
+
G. †Alloioscarabaeus Bai et al. 2012
+
†Alloioscarabaeus cheni Bai et al. 2012
G. †Ateuchites Meunier 1898
+
†Ateuchites grandis Meunier 1898
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
+
Subfm. Athyreinae Lynch Arribálzaga 1878
+
G. †Amberathyreus Bai and Zhang 2017
+
†Amberathyreus beuteli Bai and Zhang 2017
Subfm. Bolboceratinae Mulsant 1842
+
Invalid names: Bolboceratini Mulsant 1842 [invalid subgroup]
G. †Cretobolbus Nikolajev 1996
+
†Cretobolbus rohdendorfi Nikolajev 1996
G. †Ceafornotensis Woolley 2016
+
G. †Cretaclopus Nikolajev 2004
+
†Cretaclopus longipes Ponomarenko 1986
G. †Cretonitis Nikolajev 2007
+
†Cretonitis copripes Nikolajev 2007
Fm. Diphyllostomatidae Holloway 1972
+
G. †Prodiphyllostoma Yamamoto 2024
+
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
+
Subfm. Geotrupinae Latreille 1802
+
G. Ceratophyus Fischer von Waldheim 1823
+
†Ceratophyus yatagaii Aiba and Král 2023
Tr. †Cretogeotrupini Nikolajev 1996
+
G. †Cretogeotrupes Nikolajev 1992
+
†Cretogeotrupes convexus Nikolajev 1992
†Cretogeotrupes nodusus Hong and Wang 1990
Tr. Geotrupini Latreille 1802
+
G. Geotrupes Latreille 1796
+
†Geotrupes atavus Oustalet 1874
†Geotrupes germari Heer 1862
†Geotrupes messelensis Meunier 1921
†Geotrupes rottensis Statz 1952
†Geotrupes vetustus Germar 1837
Invalid names: Obitiscarabaeus Hong 1984 [synonym]
G. †Lithogeotrupes Nikolajev 2008
+
†Lithogeotrupes confusus Nikolajev 2008
G. †Parageotrupes Nikolajev and Ren 2010
+
†Parageotrupes incanus Nikolajev and Ren 2010
G. †Orrhodomala Zhang 1989
+
†Orrhodomala protista Zhang 1989
G. Typhaeus Leach 1815
Invalid names: Lethrinae Mulsant and Rey 1871 [empty], Taurocerastinae [empty]
G. †Geotrupoides Handlirsch 1906
+
†Geotrupoides lithographicus Deichmüller 1886
†Geotrupoides saxosus Zhang 1997
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
+
Subfm. Amphicominae Blanchard 1845
+
G. Amphicoma Latreille 1807
Subfm. †Cretoglaphyrinae Nikolajev 2005
+
G. †Cretoglaphyrus Nikolajev 2005
+
†Cretoglaphyrus calvescens Nikolajev 2005
†Cretoglaphyrus leptopterus Nikolajev 2005
†Cretoglaphyrus olenguicus Nikolajev 2005
†Cretoglaphyrus rohdendorfi Nikolajev 2005
†Cretoglaphyrus zherikhini Nikolajev 2005
G. †Cretohypna Yan et al. 2012
+
†Cretohypna cristata Yan et al. 2012
†Cretohypna puncta Zhao et al. 2016
†Cretohypna robusta Zhao et al. 2016
†Cretohypna yixianensis Nikolajev and Ren 2015
Subfm. Glaphyrinae MacLeay 1819
+
G. Glaphyrus Latreille 1802
+
†Glaphyrus ancestralis Nikolajev and Ren 2011
†Glaphyrus antiquus Heer 1862
G. Lichnanthe Burmeister 1844
+
†Lichnanthe defuncta Wickham 1910
G. †Lithohypna Nikolajev et al. 2011
+
†Lithohypna chifengensis Nikolajev et al. 2011
†Lithohypna laoningensis Nikolajev 2014
†Lithohypna lepticephala Nikolajev and Ren 2012
†Lithohypna longula Nikolajev and Ren 2012
†Lithohypna tuberculata Nikolajev and Ren 2012
†Lithohypna yuxiana Nikolajev and Ren 2012
G. †Mesohypna Nikolajev and Ren 2013
+
†Mesohypna lopatini Nikolajev and Ren 2013
†Mesohypna probata Nikolajev 2014
G. †Paraglaphyrus Nikolajev 2015
+
†Paraglaphyrus ovalis Nikolajev 2015
†Paraglaphyrus robustus Nikolajev 2015
†Paraglaphyrus subtilis Nikolajev 2015
†Paraglaphyrus yixianensis Nikolajev 2015
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
+
G. †Cretoglaresis Nikolajev 2007
+
†Cretoglaresis nana Nikolajev 2007
†Cretoglaresis ovalis Nikolajev 2009
G. Glaresis Erichson 1848
+
†Glaresis burmitica Cai and Huang 2018
†Glaresis cretacea Nikolajev 2007
†Glaresis orthochilus Bai et al. 2010
†Glaresis tridentata Bai et al. 2014
G. †Lithoglaresis Nikolajev 2007
+
†Lithoglaresis ponomarekoi Nikolajev 2007
Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Holcorobeus Nikritin 1977
+
†Holcorobeus atrosulcatus Nikolajev 1992
†Holcorobeus evittatus Zhang 1992
†Holcorobeus nigrovittatus Nikolajev 1992
†Holcorobeus vittatus Nikritin 1977
Invalid names: Holcorobeus (Holcorobeus) Nikritin 1977 [invalid subgroup]
G. †Hongscarabaeus Krell 2000
+
Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
+
Subfm. Anaidinae Nikolajev 1996
+
G. Anaides Westwood 1842
+
†Anaides howdeni Ocampo 2006
G. †Crassisorus Nikolajev et al. 2012
+
†Crassisorus fractus Nikolajev et al. 2012
G. †Cretanaides Nikolajev 1996
+
†Cretanaides trogopterus Nikolajev 1996
G. †Protanaides Nikolajev 2010
+
†Protanaides sibiricus Nikolajev 2010
Subfm. Ceratocanthinae Martinez 1968
+
Tr. Ceratocanthini Martinez 1968
+
G. Ceratocanthus White 1842
+
†Ceratocanthus huarongii Lu et al. 2022
G. Germarostes Paulian 1982
+
†Germarostes emarginatus Poinar 2014
G. Nesopalla Paulian and Howden 1982
+
†Nesopalla succini Lu et al. 2022
Invalid names: Acanthocerini [replaced]
G. †Palaeopycnus Lu et al. 2022
+
†Palaeopycnus circus Lu et al. 2022
†Palaeopycnus fushengii Lu et al. 2022
Invalid names: Ivieolini Howden and Gill 2000 [empty]
G. †Coprologus Heer 1847
+
†Coprologus gracilis Heer 1847
Subfm. Hybosorinae Erichson 1847
+
G. †Cretohybosorus Nikolajev 1999
+
†Cretohybosorus buryaticus Nikolajev 1999
†Cretohybosorus striatulus Nikolajev 1999
G. †Fortishybosorus Yan et al. 2013
+
†Fortishybosorus ericeusicus Yan et al. 2013
G. Hybosorus MacLeay 1819
+
†Hybosorus lividus Heer 1862
†Hybosorus ocampoi Bai and Zhang 2016
G. †Jurahybosorus Nikolajev 2005
+
†Jurahybosorus kazakhstanicus Nikolajev 2008
†Jurahybosorus mongolicus Nikolajev 2005
G. †Leptosorus Nikolajev 2006
+
†Leptosorus fortus Ren et al. 1995
†Leptosorus zherikhini Nikolajev 2006
G. Phaeochrous Laporte 1840
+
†Phaeochrous tertiarium Deichmüller 1881
G. †Protohybosorus Nikolajev 2010
+
†Protohybosorus grandissimus Nikolajev 2010
†Protohybosorus karatavicus Nikolajev 2010
†Protohybosorus mesoasiaticus Nikolajev 2010
Invalid names: Anaidini Nikolajev 1996 [empty], Hybosorini Erichson 1847 [empty]
Subfm. Liparochrinae Ocampo 2006
+
G. †Libanochrus Kirejtshuk et al. 2011
+
†Libanochrus calvus Kirejtshuk et al. 2011
Subfm. †Mimaphodiinae Nikolajev 2007
+
G. †Mimaphodius Nikolajev 2007
+
†Mimaphodius pusillus Nikolajev 2007
G. †Procoilodes Ocampo 2002
+
†Procoilodes adrastus Ocampo 2002
G. †Pulcherhybosorus Yan et al. 2012
+
†Pulcherhybosorus tridentatus Yan et al. 2012
G. †Sinochaetodus Lu et al. 2018
+
†Sinochaetodus tridentatus Lu et al. 2018
G. †Sinohybosorus Nie et al. 2018
+
†Sinohybosorus cheni Nie et al. 2018
G. †Tyrannasorus Ratcliffe and Ocampo 2001
+
†Tyrannasorus rex Ratcliffe and Ocampo 2001
G. †Hybosorites Nikolajev 1996
+
†Hybosorites fissuratus Nikolajev 1996
G. †Juraclopus Nikolajev 2004
+
†Juraclopus rohdendorfi Nikolajev 2004
Fm. Lucanidae Latreille 1804 [stag beetle]
+
Subfm. Aesalinae MacLeay 1819
+
Tr. Aesalini MacLeay 1819
+
G. †Cretaesalus Nikolajev 1993
+
†Cretaesalus ponomarenkoi Nikolajev 1993
G. †Juraesalus Nikolajev et al. 2011
+
†Juraesalus atavus Nikolajev et al. 2011
G. †Sinaesalus Nikolajev et al. 2011
+
†Sinaesalus curvipes Nikolajev et al. 2011
†Sinaesalus longipes Nikolajev et al. 2011
†Sinaesalus tenuipes Nikolajev et al. 2011
Tr. Ceratognathini Sharp 1899
+
G. †Cretognathus Yamamoto 2023
+
†Cretognathus minutissimus Yamamoto 2023
G. †Oncelytris Li and Cai 2023
+
†Oncelytris esquamatus Li and Cai 2023
Tr. Nicagini LeConte 1861
+
G. †Protonicagus Cai et al. 2017
+
†Protonicagus mandibularis Yamamoto 2023
†Protonicagus tani Cai et al. 2017
Subfm. †Ceruchitinae Nikolajev 2006
+
G. †Ceruchites Statz 1952
+
†Ceruchites hahnei Statz 1952
G. †Cretolucanus Nikolajev 2007
+
†Cretolucanus longus Nikolajev 2007
†Cretolucanus ordinarius Nikolajev 2007
†Cretolucanus sibericus Nikolajev 2007
G. †Dorcasoides Motschulsky 1856
+
†Dorcasoides bilobus Motschulsky 1856
Subfm. Lampriminae MacLeay 1819
+
G. †Electraesalopsis Bai et al. 2017
+
†Electraesalopsis beuteli Bai et al. 2017
G. †Prostreptocerus Yu and Cai 2024
+
†Prostreptocerus burmiticus Yu and Cai 2024
G. †Protognathinus Chalumeau and Brochier 2001
+
†Protognathinus spielbergi Chalumeau and Brochier 2001
Subfm. †Litholampriminae Nikolajev 2015
+
G. †Litholamprima Nikolajev and Ren 2015
+
†Litholamprima longimana Nikolajev and Ren 2015
†Litholamprima qizhihaoi Jiang et al. 2022
Subfm. Lucaninae Latreille 1804
+
G. †Anisoodontus Wu et al. 2022
+
†Anisoodontus qizhihaoi Wu et al. 2022
†Anisoodontus xiafangyuani Wu et al. 2022
Tr. Lucanini Latreille 1804
+
G. Dorcus MacLeay 1819
+
†Dorcus primigenius Deichmüller 1881
G. Lucanus Scopoli 1763
+
Lucanus cervus Linnaeus 1758
G. Penichrolucanus Deyrolle 1863
+
†Penichrolucanus kabakovi Nikolajev 2010
Invalid names: Figulidae Burmeister 1847 [synonym]
G. †Miocenidorcus Riou 1999
+
Tr. Platycerini Mulsant 1842
+
G. Platycerus Geoffroy 1762
+
†Platycerus zherichini Nikolayev 1990
G. †Succiniplatycerus Nikolayev 1990
+
G. †Prolucanus Qi et al. 2022
+
†Prolucanus beipiaoensis Qi et al. 2022
Invalid names: Chiasognathini Burmeister 1847 [empty], Lucanus fossilis Wickham 1913 [nomen dubium], Penichrolucaninae Arrow 1950 [synonym], Platycerus sepultus Germar 1837 [nomen dubium]
G. †Paleognathus Waga 1883
+
†Paleognathus succini Waga 1883
Subfm. †Paralucaninae Nikolajev 2000
+
G. †Paralucanus Nikolajev 2000
+
†Paralucanus mesozoicus Nikolajev 2000
G. †Prosinodendron Bai et al. 2012
+
†Prosinodendron krelli Bai et al. 2012
Subfm. †Protolucaninae Nikolajev 2007
+
G. †Protolucanus Nikolajev 2007
+
†Protolucanus jurassicus Nikolajev 2007
Subfm. Syndesinae MacLeay 1819
+
G. Ceruchus MacLeay 1819
+
†Ceruchus fuchsii Wickham 1911
G. Syndesus Macleay 1819
+
†Syndesus ambericus Woodruff 2009
G. †Mahengea Strümpher et al. 2023
+
†Mahengea mckayi Strümpher et al. 2023
G. †Mesaclopus Nikolajev 1992
+
†Mesaclopus mongolicus Nikolajev 1992
G. †Mesoscarabaeus Hong 1982
+
G. †Mongolrobeus Nikolajev 2004
+
†Mongolrobeus zherikhini Nikolajev 2004
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
+
Subfm. Chaetocanthinae Scholtz 1988
+
Tr. Synochodaeini Scholtz 1988
+
G. †Yixianochodaeus Nikolajev 2015
+
†Yixianochodaeus horridus Nikolajev 2015
Subfm. †Cretochodaeinae Nikolajev 1995
+
G. †Cretochodaeus Nikolajev 1995
+
†Cretochodaeus mongolicus Nikolajev 1995
†Cretochodaeus striatus Nikolajev 1995
G. †Mioochodaeus Nikolajev 1995
+
†Mioochodaeus proaevus Germar 1849
G. †Mesochodaeus Nikolajev and Ren 2010
+
†Mesochodaeus daohugouensis Nikolajev and Ren 2010
Subfm. Ochodaeinae Streubel 1846
+
G. †Lithochodaeus Nikolajev 2007
+
†Lithochodaeus cretaceus Nikolajev 2007
G. †Opiselleipon Bode 1953
+
†Opiselleipon gravis Bode 1953
Fm. Passalidae Leach 1815 [bess beetle]
+
Subfm. Aulacocyclinae Kaup 1868
+
Tr. Cylindrocaulini Zang 1905
+
Invalid names: Ceracyclini Boucher and Bai 2017 [synonym]
Subfm. Passalinae Leach 1815
+
Tr. Macrolinini Kaup 1871
+
G. Macrolinus Kaup 1868
G. †Serrulus Hong 1984
+
†Serrulus sinicus Hong 1984
Tr. Passalini Leach 1815
+
G. Passalus Fabricius 1792
+
Subg. Passalus (Passalus) Fabricius 1792
Tr. †Protopassalini Santos et al. 2020
+
G. †Protopassalus Santos et al. 2020
+
†Protopassalus araripensis Santos et al. 2020
Fm. †Passalopalpidae Boucher and Bai 2016
+
G. †Mesoceratocanthus Nikolajev et al. 2010
+
†Mesoceratocanthus tuberculifrons Nikolajev et al. 2010
G. †Passalopalpus Boucher and Bai 2016
+
†Passalopalpus cheni Boucher et al. 2016
Fm. Pleocomidae Leconte 1861 [rain beetle]
+
Subfm. †Archescarabaeinae Nikolajev 2010
+
G. †Archescarabaeus Nikolajev 2010
+
†Archescarabaeus mesozoicus Nikolajev 2010
Subfm. †Cretocominae Nikolajev 2002
+
G. †Cretocoma Nikolajev 2002
+
†Cretocoma tologoica Nikolajev 2002
Subfm. Pleocominae Leconte 1861
+
G. Pleocoma Le Conte 1856
+
†Pleocoma dolichophylla Nikolajev and Ren 2012
G. †Proteroscarabaeus Grabau 1923
+
†Proteroscarabaeus magnus Nikolajev 1996
†Proteroscarabaeus nikritini Nikolajev 1996
†Proteroscarabaeus yeni Grabau 1923
G. †Prionocephale Lin 1980
+
G. †Prophaenognatha Bai et al. 2011
+
†Prophaenognatha robusta Bai et al. 2011
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
+
Subfm. Aclopinae Blanchard 1850
+
Invalid names: Aclopini Blanchard 1850 [empty]
Subfm. Aegialiinae Laporte 1840
+
G. Aegialia Latreille 1807
+
Aegialia cylindrica Eschscholtz 1822
Aegialia lacustris Le Conte 1850
+
Invalid names: Loricera lutosa Scudder 1890 [synonym]
†Aegialia rupta Scudder 1890
Aegialia sibirica Stebnicka 1977
G. †Archeopsammoporus Minkina 2020
+
G. †Cretaegialia Nikolayev 1993
+
†Cretaegialia aphodiiformis Nikolayev 1993
†Cretaegialia rhypariformis Nikolayev 1993
G. †Antiqusolidus Lu et al. 2023
+
†Antiqusolidus maculatus Lu et al. 2023
Subfm. Aphodiinae Leach 1815
+
Tr. Aphodiini Leach 1815
+
G. Agoliinus Schmidt 1913
+
Subtr. Aphodiina Leach 1815
+
G. Aphodius Hellwig 1798
G. Diapterna Horn 1891
G. Heptaulacus Mulsant 1842
G. Caligodorus Gordon and Skelley 2007
+
Caligodorus opacus Le Conte 1872
G. Dialytes Harold 1869
G. Dialytodius Gordon and Skelley 2007
+
G. Pardalosus Gordon and Skelley 2007
+
Pardalosus pardalis Le Conte 1857
G. Planolinoides Dellacasa and Dellacasa 2005
+
Planolinoides duplex LeConte 1878
G. †Cretorabaeus Nikolajev 1995
+
†Cretorabaeus elongatus Nikolajev 1995
Tr. Eupariini Schmidt 1910
+
G. Airapus Stebnicka and Howden 1996
+
†Airapus lithuanicus Tamutis et al. 2017
G. Ataenius Harold 1867
+
†Ataenius damzeni Bukejs and Alekseev 2018
†Ataenius europaeus Quiel 1911
†Ataenius patescens Scudder 1900
†Ataenius restructus Wickham 1912
G. Oxyomus Kirby and Spence 1828
+
†Oxyomus nearcticus Wickham 1914
G. Saprosites Redtenbacher 1858
+
†Saprosites cascus Britton 1960
†Saprosites succini Zang 1905
Tr. †Psammaegialiini Nikolajev et al. 2014
+
G. †Psammaegialia Nikolajev et al. 2014
+
†Psammaegialia abdita Nikolajev et al. 2014
†Psammaegialia zebrina Nikolajev et al. 2014
Tr. Psammodiini Mulsant 1842
+
Subtr. Psammodiina Mulsant 1842
+
G. Pleurophorus Mulsant 1842
G. Psammodius Fallen 1807
Invalid names: Pleurophorina Mulsant 1842 [synonym]
Subtr. Rhyssemina Pittinno and Marianni 1986
+
G. Rhyssemus Mulsant 1842
Tr. Rhyparini Schmidt 1910
+
G. Leptorhyparus Howden 2003
+
G. †Priscyparus Skelley 2021
+
†Priscyparus quisqueyensis Skelley 2021
Subfm. Aulonocneminae Janssens 1946
+
G. †Protopalnia Nikolajev 2007
+
†Protopalnia magadanica Nikolajev 2007
G. Caccobius Thompson 1859
+
Caccobius schreberi Linnaeus 1767
Subfm. Cetoniinae Leach 1815 [flower chafer]
+
Tr. Cetoniini Leach 1815
+
Subtr. Cetoniina Leach 1815
+
G. Cetonia Fabricius 1775
Tr. Schizorhinini Burmeister 1842
+
G. †Genonota Zhang 1989
+
†Genonota mochthera Zhang 1989
†Genonota scola Zhang 1989
G. Heterosoma Schaum 1845
Subtr. Lomapterina Burmeister 1842
+
G. Macronota Hoffmansegg 1817
G. †Paleotrichius Poinar 2011
+
Tr. Trichiini Fleming 1821
+
G. Gnorimus Lepeletier and Serville 1828
+
†Gnorimus aedilis Heer 1862
Subtr. Trichiina Fleming 1821
+
G. Trichius Fabricius 1775
Tr. Valgini Mulsant 1842
+
Subtr. Valgina Mulsant 1842
+
G. Valgus Scriba 1790
Subfm. †Cretoscarabaeinae Nikolajev 1995
+
G. †Cretoscarabaeus Nikolajev 1995
+
†Cretoscarabaeus gibbosus Nikolajev 1995
†Cretoscarabaeus lentiginosus Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
+
G. †Calcitoryctes Krell 2011
+
G. Cyclocephala Dejean 1821
+
G. Ligyrus Burmeister 1847
+
†Ligyrus compositus Wickham 1911
†Ligyrus effetus Wickham 1914
Tr. Oryctini Mulsant 1842
+
G. Oryctes Hellwig 1798
+
Oryctes nasicornis Linnaeus 1758 [European rhinoceros beetle]
G. Strategus Kirby 1828
+
†Strategus cessatus Wickham 1914
G. †Oryctoantiquus Ratcliffe and Smith 2005
+
†Oryctoantiquus borealis Ratcliffe and Smith 2005
Tr. Pentodontini Mulsant 1842
+
G. Diloboderus Reiche 1859
+
Subtr. Pentodontina Mulsant 1842
+
G. Pentodon Hope 1837
Invalid names: Pentodon proserpinae Heer 1862 [nomen dubium]
Subfm. †Electrorubesopsinae Bai and Wang 2018
+
G. †Electrorubesopsis Bai and Wang 2018
+
†Electrorubesopsis beuteli Bai and Wang 2018
Subfm. Eremazinae Stebnicka 1977
+
Subfm. †Yixianscarabaeinae Nikolajev 2015
+
G. †Yixianscarabaeus Nikolajev 2015
+
†Yixianscarabaeus sulcatus Nikolajev 2015
G. Liothorax Motschulsky 1859
+
Liothorax kraatzi Harold 1868
Subfm. †Lithoscarabaeinae Nikolajev 1992
+
G. †Baisarabaeus Nikolajev 2005
+
†Baisarabaeus rugosus Nikolajev 2005
†Baisarabaeus yixianensis Nikolajev et al. 2013
G. †Lithoscarabaeus Nikolajev 1992
+
†Lithoscarabaeus baissensis Nikritin 1977
Subfm. Melolonthinae Leach 1819
+
Tr. †Cretomelolonthini Nikolajev 1998
+
G. †Cretomelolontha Nikolajev 1998
+
Tr. Diplotaxini Kirby 1837
+
G. Diplotaxis Kirby 1837
+
†Diplotaxis aurora Wickham 1913
†Diplotaxis simplicipes Wickham 1912
G. †Eophyllocerus Haupt 1950
+
Tr. Euchirini Hope 1840
+
G. Cheirotonus Hope 1840
+
†Cheirotonus otai Ueda 1989
Tr. Hopliini Latreille 1829
+
Subtr. Hopliina Latreille 1829
+
G. Hoplia Illiger 1803
Subtr. Pachycnemina Laporte 1840
+
G. Lepitrix Lepeletier and Serville 1828
G. Listrochelus Blanchard 1851
+
†Listrochelus puerilis Wickham 1914
Tr. Macrodactylini Kirby 1837
+
G. Macrodactylus Latreille 1827
+
†Macrodactylus pluto Wickham 1912
†Macrodactylus propheticus Wickham 1912
Tr. Melolonthini Leach 1819
+
Subtr. Melolonthina Leach 1819
+
G. Melolontha Fabricius 1775
G. Phyllophaga Harris 1826
+
†Phyllophaga avus Cockerell 1921
†Phyllophaga disrupta Cockerell 1927
†Phyllophaga extincta Wickham 1916
Invalid names: Lachnosterna Hope 1837 [synonym]
Subtr. Rhizotrogina Burmeister 1855
+
G. Rhizotrogus Latreille 1825
Invalid names: Schizonychina Burmeister 1855 [empty]
G. †Melolonthites Heer 1847
+
†Melolonthites collinsi Wickham 1929
†Melolonthites interemtus Cockerell 1926
†Melolonthites kollari Heer 1847
Tr. Sericini Kirby 1837
+
G. †Cretoserica Nikolajev 1998
+
†Cretoserica latitibialis Nikolajev 1998
G. †Lithanomala Nikolajev 1992
+
†Lithanomala crassa Ponomarenko 1990
†Lithanomala oblonga Ponomarenko 1990
†Lithanomala sibirica Ponomarenko 1990
G. Maladera Mulsant 1871
+
†Maladera spinitibialis Statz 1952
G. †Mioserica Zhang 1989
+
†Mioserica margelis Zhang 1989
Subtr. Sericina Kirby 1837
+
G. Serica MacLeay 1819
G. †Mesydra Ponomarenko 1977
+
†Mesydra elongata Ponomarenko 1977
Subfm. †Prototroginae Nikolajev 2000
+
G. †Prototrox Nikolajev 2000
+
†Prototrox transbaikalicus Nikolajev 2000
Subfm. Rutelinae MacLeay 1819
+
Tr. Adoretini Burmeister 1844
+
Subtr. Adoretina Burmeister 1844
+
G. Adoretus Dejean 1833
Tr. Anomalini Streubel 1839
+
Subtr. Anomalina Streubel 1839
+
G. Anomala Samouelle 1819
+
Invalid names: Magniscarabaeus Hong 1984 [synonym]
G. Phyllopertha Stephens 1830
G. †Anomalites Frič 1885
+
†Anomalites fugitivus Frič 1885
Tr. Anoplognathini MacLeay 1819
+
Subtr. Anoplognathina MacLeay 1819
+
G. Anoplognathus Leach 1815
G. Holotrichia Hope 1837
+
†Holotrichia cressona Zhang et al. 1994
†Holotrichia spatha Zhang 1989
G. Mimela Kirby 1823
+
†Mimela rhenana von Heyden 1862
Mimela testaceipes Motschulsky 1860
Invalid names: Rhombonyx Hope 1837 [synonym]
G. †Miolachnosterna Wickham 1914
+
G. †Pelidnotites Cockerell 1920
+
†Pelidnotites atavus Cockerell 1920
G. †Petraeianus Zhang 1989
+
†Petraeianus ruderatus Zhang 1989
G. †Petulantis Zhang et al. 1994
+
†Petulantis yimengensis Zhang et al. 1994
G. †Zhangsunia Krell 2000
+
†Zhangsunia extumida Zhang et al. 1994
Invalid names: Dolichopoda Zhang et al. 1994 [replaced]
Subfm. Scarabaeinae Latreille 1802
+
G. Anachalcos Hope 1837
+
†Anachalcos mfwangani Paulian 1976
Tr. Ateuchini Perty 1830
+
Subtr. Ateuchina Perty 1830
+
G. Ateuchus Weber 1801
+
Invalid names: Choeridium Lepeletier de Saint-Fargeau and Audinet-Serville 1828 [synonym]
G. †Lobateuchus Montreuil et al. 2010
+
†Lobateuchus parisii Montreuil et al. 2010
G. Canthochilum Chapin 1934
+
†Canthochilum alleni Tarasov and Vaz-de-Mello 2016
†Canthochilum philipsivieorum Tarasov and Vaz-de-Mello 2016
Tr. Coprini Leach 1815
+
G. Copris Geoffroy 1762
+
Subg. Copris (Copris) Geoffroy 1762
†Copris druidum Heer 1862
Copris incertus Say 1835
†Copris leakeyorum Paulian 1976
Copris lunaris Linnaeus 1758 [horned dung beetle]
†Copris pristinus Pierce 1946
Tr. Deltochilini Lacordaire 1856
+
G. Canthon Hoffmannsegg 1817
+
Subg. Canthon (Boreocanthon) Halffter 1958
G. Deltochilum Eschscholtz 1822
G. Scybalophagus Martínez 1953
+
†Scybalophagus brellenthinae Tello et al. 2023
Tr. Gymnopleurini Lacordaire 1856
+
G. Gymnopleurus Illiger 1803
+
†Gymnopleurus sisyphus Heer 1847
G. Heliocopris Hope 1837
+
†Heliocopris antiquus Fujiyama 1968
G. Metacatharsius Paulian 1939
+
†Metacatharsius rusingae Paulian 1976
Tr. Oniticellini Kolbe 1905
+
Subtr. Drepanocerina van Lansberge 1875
+
G. Eodrepanus Barbero et al. 2009
Subtr. Oniticellina Kolbe 1905
+
G. Oniticellus Dejean 1821
Tr. Onitini Laporte 1840
+
G. Onitis Fabricius 1798
Tr. Onthophagini Burmeister 1846
+
G. Onthophagus Latreille 1802
+
†Onthophagus arvernus Piton and Théobald 1939
†Onthophagus crassus Heer 1862
†Onthophagus everestae Pierce 1946
Onthophagus fracticornis Preyssler 1790
Onthophagus janus Panzer 1794
Onthophagus lenzii Harold 1874
†Onthophagus ovatulus Heer 1847
†Onthophagus pilauco Tello et al. 2021
†Onthophagus prodromus Heer 1862
†Onthophagus statzi Krell 1990
+
Invalid names: Onthophagus muelleri Statz 1952 [replaced]
Onthophagus taurus Schreber 1759
G. Phalops Erichson 1848
Tr. Phanaeini Hope 1838
+
G. Phanaeus Macleay 1819
+
†Phanaeus labreae Pierce 1946
†Phanaeus violetae Zunino 2013
Invalid names: Palaeocopris Pierce 1946 [synonym], Phanaeus antiquus Horn 1876 [nomen dubium]
Tr. Scarabaeini Latreille 1802
+
G. Scarabaeus Linnaeus 1758
Invalid names: Onthophagus spitsbergeniensis Krell 2010 [nomen dubium]
G. Triodonta Mulsant 1842
Invalid names: Allidiostomatinae Arrow 1940 [empty], Coprophaginae [empty], Dynamopodinae Arrow 1911 [empty], Oniticellus amplicollis Heer 1862 [nomen dubium], Orphninae Erichson 1847 [empty], Pachypodinae Erichson 1840 [empty], Phaenomeridinae Erichson 1847 [empty]
G. †Scelocopris Zhang 1989
+
†Scelocopris enertheus Zhang 1989
Fm. †Septiventeridae Bai et al. 2013
+
G. †Septiventer Bai et al. 2013
+
†Septiventer quadridentatus Bai et al. 2013
Fm. Trogidae MacLeay 1819 [hide beetle]
+
Subfm. †Kresnikinae Tihelka et al. 2019
+
G. †Kresnikus Tihelka et al. 2019
+
†Kresnikus beynoni Tihelka et al. 2019
Subfm. Omorginae Nikolajev 2005
+
G. †Cretomorgus Nikolajev 2007
+
†Cretomorgus ikhbogdensis Nikolajev 2007
Subfm. Troginae MacLeay 1819
+
Tr. †Avitortorini Nikolajev 2007
+
G. †Avitortor Ponomarenko 1977
+
†Avitortor dolichodactylus Nikolajev 2007
†Avitortor leptoscelis Nikritin 1977
†Avitortor ovalis Nikolajev 2007
†Avitortor parallelus Nikolajev 2007
†Avitortor primitivus Ponomarenko 1977
G. †Paratrox Nikolajev 2009
+
†Paratrox medvedevi Nikolajev 2009
G. Trox Fabricius 1775
+
†Trox antiquus Wickham 1909
†Trox cretaceus Nikolajev 2007
†Trox minutus Nikolajev 2008
†Trox sibericus Nikolajev 2007
Invalid names: Trox oustaleti Scudder 1879 [nomen dubium]
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Angulata to the top of the Holocene or 201.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 196.5 Ma

Collections (310 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Angulata201.3 - 196.5Switzerland Aphodiites protogaeus (131766)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Opiselleipon gravis (105002)
Bajocian - Bathonian170.3 - 166.1Mongolia (Uver-Khangay Aymag) Jurahybosorus mongolicus (118011)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Jurahybosorus kazakhstanicus (105971) Protohybosorus grandissimus (112846) Protohybosorus karatavicus (113439) Protohybosorus mesoasiaticus (106102)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Alloioscarabaeus cheni (113366) Juraesalus atavus (113371) Mesochodaeus daohugouensis (113365)
Tithonian152.1 - 145.0Mongolia Paralucanus mesozoicus (104748)
Early/Lower Tithonian150.8 - 145.0Germany (Bayern) Geotrupes lithographicus (133578)
Valanginian - Barremian139.8 - 125.0Mongolia (Dornogov) Holcorobeus mongolicus (113407)
Early/Lower Barremian130.0 - 125.45Lebanon Libanochrus calvus (123416)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Lithohypna chifengensis, Baisarabaeus yixianensis, Crassisorus fractus (125655) Mesoceratocanthus tuberculifrons, Sinaesalus tenuipes, Sinaesalus longipes, Sinaesalus curvipes, Psammaegialia abdita, Psammaegialia zebrina (103654) Pulcherhybosorus tridentatus, Cretohypna cristata, Cretohypna puncta, Cretohypna robusta, Antiqusolidus maculatus (125656)
Late/Upper Barremian130.0 - 125.45China (Hebei) Geotrupoides songyingziensis (155165)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Antiqusolidus maculatus (140491) Fortishybosorus ericeusicus (124759) Glaphyrus ancestralis, Pleocoma dolichophylla, Mesohypna lopatini, Lithohypna laoningensis, Mesohypna probata, Lithohypna tuberculata, Lithohypna lepticephala, Lithohypna yuxiana, Lithohypna longula, Paraglaphyrus yixianensis, Paraglaphyrus robustus, Paraglaphyrus ovalis, Paraglaphyrus subtilis, Litholamprima longimana, Yixianscarabaeus sulcatus, Yixianscarabaeus tenuistriatus, Cretohypna yixianensis, Yixianochodaeus horridus (111712) Glaresis orthochilus (123396) Litholamprima qizhihaoi (226159 226160) Litholamprima sp. 1 (156389) Litholamprima sp. 2 (226161) Mesoceratocanthus tuberculifrons, Prolucanus beipiaoensis (111725) Parageotrupes incanus (123472) Prophaenognatha robusta, Prosinodendron krelli (111716) Septiventer quadridentatus, Glaresis tridentata (141615) Sinohybosorus cheni, Sinochaetodus tridentatus (182836)
Barremian129.4 - 125.0Russian Federation (Chita) Proteroscarabaeus crassus, Proteroscarabaeus oblongus, Proteroscarabaeus sibiricus (151642)
Aptian125.0 - 113.0China (Jilin) Geotrupoides jiaoheense (172386) Geotrupoides saxosus, Proteroscarabaeus robustus (151876)
Aptian125.0 - 113.0China (Nei Monggol) Geotrupoides fortus (152245)
Aptian125.0 - 113.0Russian Federation (Buryatia) Aphodius cretaceous, Penichrolucanus kabakovi (118216) Baisarabaeus rugosus, Leptosorus zherikhini (125791) Cretanaides trogopterus (113354) Cretobolbus rohdendorfi, Cretorabaeus elongatus, Baisarabaeus rugosus, Lithoglaresis ponomarekoi, Avitortor ovalis, Geotrupoides sulcatus (119498) Cretomelolontha transbaikalica, Cretoserica latitibialis, Holcorobeus nigrovittatus, Cretohybosorus striatulus, Cretoglaresis nana, Lithogeotrupes confusus (106103) Cretoscarabaeus lentiginosus (125413) Glaresidae indet. (113349) Hybosorites fissuratus, Cretaegialia aphodiiformis, Cretoscarabaeus gibbosus (119441) Lucanidae indet., Glaphyridae indet., Scarabaeidae indet. (113350) Mesydra elongata, Proteroscarabaeus baissensis, Proteroscarabaeus yeni, Holcorobeus vittatus, Holcorobeus atrosulcatus, Geotrupoides sulcatus (105536) Protanaides sibiricus, Avitortor primitivus, Geotrupoides leptoscelis, Proteroscarabeus baissensis, Holcorobeus vittatus, Cretoscarabaeus gibbosus, Trox sibericus, Cretonitis copripes (105969) Proteroscarabaeus baissensis, Proteroscarabaeus yeni, Holcorobeus vittatus, Avitortor leptoscelis (113403) Proteroscarabaeus magnus (125786) Proteroscarabaeus nikritini, Cretoglaresis ovalis, Cretomelolontha transbaikalica, Holcorobeus atrosulcatus, Cretohybosorus buryaticus, Cretoglaphyrus rohdendorfi, Glaresis cretacea, Mimaphodius pusillus (122549)
Aptian125.0 - 113.0Mongolia Cretocoma tologoica, Holcorobeus atrosulcatus, Paratrox medvedevi, Cretomorgus ikhbogdensis, Archescarabaeus mesozoicus (103273) Protolucanus jurassicus (105796) Prototrox transbaikalicus, Mongolrobeus zherikhini, Cretochodaeus mongolicus, Cretochodaeus striatus, Lithochodaeus cretaceus, Trox cretaceus, Avitortor parallelus, Avitortor dolichodactylus (103657)
Aptian125.0 - 113.0China (Gansu) Mesoscarabaeus corneus (155034) Proscarabaeus brunneus, Mesoscarabaeus morulosus (155045)
Aptian125.0 - 113.0Mongolia (Kobdo aimak) Geotrupoides longipes (104054 136025)
Aptian125.0 - 113.0China (Shandong) Holcorobeus evittatus (124369) Proteroscarabaeus baissensis, Proteroscarabaeus yeni, Geotrupoides nodusus (154887) Proteroscarabaeus yeni (154897 156598)