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Clypeodonta

Reptilia

Taxonomy
Clypeodonta was named by Norman (2014).

It was assigned to Neornithischia by Madzia et al. (2018); and to Ornithopoda by Norman (2014), Herne et al. (2019) and Dieudonné et al. (2020).

Subtaxa
Hypsilophodontidae (syn. Laosauridae, Macelognathidae, Kalodontidae), Hypsilophodontoidea, Iguanodontia, Proyandusaurus, Thescelosaurus (syn. Bugenasaura)
Sister taxa
Hypsilophodontidae (syn. Laosauridae, Macelognathidae, Kalodontidae), Iguanodontia

Synonyms
Synonymy list
YearName and author
1984Hypsilophodontia Russell p. 29
1985Hypsilophodontia Cooper p. 285
1986Hypsilophodontia Sereno
1989Hypsilophodontia Norman p. 63 fig. 1
2014Hypsilophodontia Alifanov and Saveliev p. 418
2014Clypeodonta Norman p. 29
2018Clypeodonta Madzia et al. p. 973 fig. 4
2019Clypeodonta Herne et al. p. 575
2020Clypeodonta Dieudonné et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
—Sauropsida
classReptilia
RankNameAuthor
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
informalAvemetatarsalia
—Ornithodira
—Dinosauromorpha
—Dinosauriformes
—Dinosauria()
—Ornithischia()
—Genasauria
—Cerapoda
—Ornithopoda()
—Clypeodonta
—Clypeodonta

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Clypeodonta Norman 2014
show all | hide all
Unr. †Hypsilophodontidae Dollo 1882 [hypsilophodontid]
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G. †Daurosaurus Alifanov and Saveliev 2014
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†Daurosaurus olovus Alifanov and Saveliev 2014
G. †Hypsilophodon Huxley 1870
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†Hypsilophodon foxii Huxley 1870
Unr. †Parksosauridae Buchholz 2002
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G. †Gasparinisaura Coria and Salgado 1996
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†Gasparinisaura cincosaltensis Coria and Salgado 1996
G. †Vectidromeus Longrich et al. 2024
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†Vectidromeus insularis Longrich et al. 2024
Invalid names: Dryosaurinae Cooper 1985 [empty], Hypsilophodontinae Abel 1919 [empty], Kalodontidae Nopcsa 1901 [synonym], Laosauridae Marsh 1895 [synonym], Macelognathidae Marsh 1884 [synonym], Nanosaurinae Marsh 1878 [empty]
Unr. †Iguanodontia Baur 1891
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G. †Choyrodon Gates et al. 2018
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†Choyrodon barsboldi Gates et al. 2018
G. †Convolosaurus Andrzejewski et al. 2019
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†Convolosaurus marri Andrzejewski et al. 2019
Unr. †Dryomorpha Sereno 1986
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Unr. †Ankylopollexia Sereno 1986
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G. †Bayannurosaurus Xu et al. 2018
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†Bayannurosaurus perfectus Xu et al. 2018
Fm. †Camptosauridae Marsh 1885 [camptosaurid]
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Invalid names: Camptonotidae Marsh 1881 [synonym]
G. †Camptosaurus Marsh 1885
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†Camptosaurus aphanoecetes Carpenter and Wilson 2008
†Camptosaurus dispar Marsh 1879
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Invalid names: Brachyrophus altarkansanus Cope 1878 [synonym], Camptosaurus browni Gilmore 1909 [synonym], Camptosaurus medius Marsh 1894 [synonym], Camptosaurus nanus Marsh 1894 [synonym], Symphyrophus musculosus Cope 1878 [synonym]
Invalid names: Brachyrophus Cope 1878 [synonym], Camptonotus Marsh 1879 [replaced], Symphyrophus Cope 1878 [synonym], Uteodon McDonald 2011 [synonym]
G. †Iguanodon Mantell 1825
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†Iguanodon bernissartensis Boulenger 1881
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Invalid names: Iguanodon seelyi Hulke 1882 [synonym]
†Iguanodon galvensis Verdú et al. 2015
†Iguanodon major Owen 1842
Invalid names: Heterosaurus Cornuel 1850 [synonym], Iguanodon anglicus Holl 1829 [nomen dubium], Iguanosaurus Anonymous 1824 [synonym], Struthopus Ballerstedt 1922 [synonym], Therosaurus Fitzinger 1843 [synonym]
Unr. †Neoiguanodontia Norman 2014
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G. †Altirhinus Norman 1998
Unr. †Hadrosauromorpha Norman 2014
G. †Hypselospinus Norman 2010
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Invalid names: Darwinsaurus Paul 2012 [synonym], Wadhurstia Carpenter and Ishida 2010 [objective synonym]
Fm. †Iguanodontidae Bonaparte 1850 [iguanodontid]
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Invalid names: Ornithopodidae Zittel 1895 [synonym]
G. †Ouranosaurus Taquet 1972
G. †Oblitosaurus Sánchez-Fenollosa et al. 2023
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†Oblitosaurus bunnueli Sánchez-Fenollosa et al. 2023
Fm. †Dryosauridae Milner and Norman 1984
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G. †Callovosaurus Galton 1980
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†Callovosaurus leedsi Lydekker 1889
G. †Dinehichnus Lockley et al. 1998
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†Dinehichnus socialis Lockley et al. 1998
G. †Dryosaurus Marsh 1894
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†Dryosaurus altus Marsh 1878
†Dryosaurus elderae Carpenter and Galton 2018
G. †Dysalotosaurus Virchow 1919
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G. †Elrhazosaurus Galton 2009
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†Elrhazosaurus nigeriensis Galton and Taquet 1982
G. †Valdosaurus Galton 1977
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G. †Eousdryosaurus Escaso et al. 2014
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†Eousdryosaurus nanohallucis Escaso et al. 2014
G. †Hesperonyx Rotatori et al. 2024
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†Hesperonyx martinhotomasorum Rotatori et al. 2024
G. †Owenodon Galton 2009
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†Owenodon hoggii Owen 1874
Unr. †Styracosterna Sereno 1986
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G. †Calvarius Prieto-Márquez and Sellés 2023
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†Calvarius rapidus Prieto-Márquez and Sellés 2023
G. †Cumnoria Seeley 1888
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†Cumnoria prestwichii Hulke 1880
G. †Dakotadon Paul 2008
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†Dakotadon lakotaensis Weishampel and Bjork 1989
G. †Delapparentia Ruiz-Omeñaca 2011
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†Delapparentia turolensis Ruiz-Omeñaca 2011
G. †Draconyx Mateus and Antunes 2001
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†Draconyx loureiroi Mateus and Antunes 2001
Unr. †Hadrosauriformes Sereno 1997
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G. †Brighstoneus Lockwood et al. 2021
G. †Comptonatus Lockwood et al. 2024
Unr. †Hadrosauroidea Huene 1954
G. †Proa McDonald et al. 2012
G. †Siamodon Buffetaut and Suteethorn 2011
G. †Hippodraco McDonald et al. 2010
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†Hippodraco scutodens McDonald et al. 2010
G. †Iguanacolossus McDonald et al. 2010
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†Iguanacolossus fortis McDonald et al. 2010
G. †Lanzhousaurus You et al. 2005
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†Lanzhousaurus magnidens You et al. 2005
G. †Magnamanus Fuentes Vidarte et al. 2016
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†Magnamanus soriaensis Fuentes Vidarte et al. 2016
G. †Napaisaurus Ji and Zhang 2022
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†Napaisaurus guangxiensis Ji and Zhang 2022
G. †Riabininohadros Ulansky 2015
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†Riabininohadros weberae Riabinin 1946
G. †Theiophytalia Brill and Carpenter 2007
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†Theiophytalia kerri Brill and Carpenter 2007
Invalid names: Dryosauroidea Norman 1984 [empty]
Unr. †Elasmaria Calvo et al. 2007
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G. †Anabisetia Coria and Calvo 2002
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†Anabisetia saldiviai Coria and Calvo 2002
G. †Atlascopcosaurus Rich and Vickers-Rich 1989
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†Atlascopcosaurus loadsi Rich and Vickers-Rich 1989
G. †Chakisaurus Nogueira et al. 2024
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†Chakisaurus nekul Nogueira et al. 2024
G. †Diluvicursor Herne et al. 2018
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†Diluvicursor pickeringi Herne et al. 2018
G. †Galleonosaurus Herne et al. 2019
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†Galleonosaurus dorisae Herne et al. 2019
G. †Isasicursor Novas et al. 2019
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†Isasicursor santacrucensis Novas et al. 2019
G. †Kangnasaurus Haughton 1915
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†Kangnasaurus coetzeei Haughton 1915
G. †Leaellynasaura Rich and Vickers-Rich 1989
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†Leaellynasaura amicagraphica Rich and Vickers-Rich 1989
G. †Macrogryphosaurus Calvo et al. 2007
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†Macrogryphosaurus gondwanicus Calvo et al. 2007
G. †Mahuidacursor Cruzado-Caballero et al. 2019
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†Mahuidacursor lipanglef Cruzado-Caballero et al. 2019
G. †Morrosaurus Rozadilla et al. 2016
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†Morrosaurus antarcticus Rozadilla et al. 2016
G. †Talenkauen Novas et al. 2004
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†Talenkauen santacrucensis Novas et al. 2004
G. †Tietasaura Bandeira et al. 2024
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†Tietasaura derbyiana Bandeira et al. 2024
G. †Trinisaura Coria et al. 2013
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†Trinisaura santamartaensis Coria et al. 2013
Superfm. †Iguanodontoidea Hay 1902
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G. †Craspedodon Dollo 1883
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†Craspedodon lonzeensis Dollo 1883
G. †Lurdusaurus Taquet and Russell 1999
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†Lurdusaurus arenatus Taquet and Russell 1999
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Invalid names: Gravisaurus tenerensis Chabli 1988 [objective synonym]
Invalid names: Gravisaurus Chabli 1988 [objective synonym]
Invalid names: Hadrosauria Horner 1990 [empty], Iguanodon orientalis Rozhdestvensky 1952 [nomen dubium], Lambeosauria Horner 1990 [empty]
G. †Iyuku Forster et al. 2022
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†Iyuku raathi Forster et al. 2022
G. †Mantellisaurus Paul 2007
+
+
Invalid names: Dollodon bampingi Paul 2008 [synonym], Heterosaurus neocomiensis Cornuel 1850 [synonym], Mantellodon carpenteri Paul 2012 [synonym]
Invalid names: Dollodon Paul 2008 [synonym], Mantellodon Paul 2012 [synonym], Proplanicoxa Carpenter and Ishida 2010 [nomen dubium]
G. †Osmakasaurus McDonald 2011
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†Osmakasaurus depressus Gilmore 1909
G. †Planicoxa DiCroce and Carpenter 2001
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†Planicoxa venenica DiCroce and Carpenter 2001
Unr. †Rhabdodontomorpha Dieudonné et al. 2016
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G. †Emiliasaura Coria et al. 2024
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†Emiliasaura alessandrii Coria et al. 2024
G. †Fostoria Bell et al. 2019
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†Fostoria dhimbangunmal Bell et al. 2019
G. †Iani Zanno et al. 2023
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†Iani smithi Zanno et al. 2023
G. †Muttaburrasaurus Bartholomai and Molnar 1981
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†Muttaburrasaurus langdoni Bartholomai and Molnar 1981
Fm. †Rhabdodontidae Weishampel et al. 2003 [rhabdodontid]
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G. †Matheronodon Godefroit et al. 2017
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†Matheronodon provincialis Godefroit et al. 2017
G. †Mochlodon Seeley 1881
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†Mochlodon priscus Matheron 1869
†Mochlodon vorosi Osi et al. 2012
†Rhabdodon suessi Bunzel 1871
G. †Pareisactus Párraga and Prieto-Márquez 2019
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†Pareisactus evrostos Párraga and Prieto-Márquez 2019
G. †Rhabdodon Matheron 1869
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†Rhabdodon priscus Matheron 1869
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Invalid names: Oligosaurus adelus Seeley 1881 [synonym], Ornithomerus gracilis Seeley 1881 [synonym]
†Rhabdodon septimanicus Buffetaut and Le Loeuff 1991
Invalid names: Oligosaurus Seeley 1881 [synonym], Ornithomerus Seeley 1881 [synonym]
G. †Transylvanosaurus Augustin et al. 2022
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†Transylvanosaurus platycephalus Augustin et al. 2022
G. †Zalmoxes Weishampel et al. 2003
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†Zalmoxes robustus Nopcsa 1900
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Invalid names: Camptosaurus inkeyi Nopcsa 1900 [synonym], Onychosaurus hungaricus Nopcsa 1902 [synonym]
†Zalmoxes shqiperorum Weishampel et al. 2003
Invalid names: Onychosaurus Nopcsa 1902 [synonym]
G. †Sektensaurus Ibiricu et al. 2019
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†Sektensaurus sanjuanboscoi Ibiricu et al. 2019
G. †Tenontosaurus Ostrom 1970
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†Tenontosaurus dossi Winkler et al. 1997
†Tenontosaurus tilletti Ostrom 1970
G. †Yueosaurus Zheng et al. 2012
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†Yueosaurus tiantaiensis Zheng et al. 2012
Invalid names: Albisaurus Fritsch 1905 [nomen dubium], Albisaurus scutifer Fritsch 1905 [nomen dubium], Anoplosaurus major Seeley 1879 [nomen dubium], Bihariosaurus Marinescu 1989 [nomen nudum], Camptosauroidea Cooper 1985 [empty], Eucercosaurus Seeley 1879 [nomen dubium], Eucercosaurus tanyspondylus Seeley 1879 [nomen dubium], Euiguanodontia Coria and Salgado 1996 [empty], Fulgurotherium Huene 1932 [nomen dubium], Iguanodon ottingeri Galton and Jensen 1979 [nomen dubium], Loncosaurus Ameghino 1899 [nomen dubium], Procerosaurus exogirarum Fritsch 1878 [nomen dubium], Sphenospondylus Seeley 1883 [nomen dubium], Syngonosaurus Seeley 1879 [nomen dubium], Syngonosaurus macrocercus Seeley 1869 [nomen dubium], Tichosteus Cope 1877 [nomen dubium], Vectisaurus Hulke 1879 [nomen dubium]
G. †Thescelosaurus Gilmore 1913
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†Thescelosaurus assiniboiensis Brown et al. 2011
†Thescelosaurus edmontonensis Sternberg 1940
†Thescelosaurus garbanii Morris 1976
†Thescelosaurus neglectus Gilmore 1913
Invalid names: Bugenasaura Galton 1995 [synonym], Bugenasaura infernalis Galton 1995 [nomen dubium]
Invalid names: Hypsilophodontia Cooper 1985 [synonym]
Diagnosis
ReferenceDiagnosis
V. R. Alifanov and S. V. Saveliev 2014 (Hypsilophodontia)Preorbital fenestrae expressed, often large. Supraoccipitals forming border of foramen magnum. Premaxillae toothed. Nasals and maxillae in contact. Frontals longer than, or equal to, nasals. Preacetabular process of pubes rodlike. Acetabular edge of these bones curving towards acetabulum and not divided into iliac and ischiadic rami. Descending ramus of pubes approximately as long as ischia. Scapulae not longer than humeri. In manus, digit III consisting of four phalanges and digit V including at most three phalanges. Femora always shorter than tibiotarsi. Digit I of foot usually enlarged. Integument having bristle like appendages.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Maximum body mass: 10000 kgg
Minimum body mass: 3 kgg
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: gregariouso
Environment: terrestrialo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: herbivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2009-01-03 20:40:05
Modified: 2009-01-03 22:40:05
Source: g = genus, subo = suborder, o = order
References: Marsh 1875, Peczkis 1995, Benton 1983

Age range: base of the Early/Lower Jurassic to the top of the Edmontonian or 201.40000 to 66.00000 Ma

Collections (2370 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Early/Lower Jurassic201.4 - 174.7China (Gansu) Iguanodontidae indet. (131877)
Aalenian - Bajocian174.1 - 168.3China (Sichuan) Trachodontidae indet. (140257)
Early/Lower Bajocian - Early/Lower Bathonian171.6 - 164.7Mexico (Oaxaca) Ornithopoda indet. (36267)
Early/Lower Bathonian167.7 - 164.7United Kingdom (England) Alocodon sp. (31054)
Early/Lower Bathonian167.7 - 164.7France Hypsilophodontidae indet. (61378)
Late/Upper Bathonian - Late/Upper Jurassic167.7 - 145.0Russian Federation (Zabaykal'ye) Daurosaurus olovus (159014)
Middle Callovian164.7 - 161.2Germany (Nordrhein-Westfalen) Dryosauridae indet. (67698)
Middle Callovian164.7 - 161.2United Kingdom (England) Camptosaurus leedsi (59128) Reptilia indet. (109800)
Oxfordian163.5 - 157.3Chile (Tarapaca) Camptosauridae indet. (170681)
Late/Upper Jurassic161.5 - 145.0China (Yunnan) Dinehichnus sp. (229403)

Youngest occurrences
Time interval Ma Country or state PBDB collection number
Late/Upper Maastrichtian70.6 - 66.0USA (Colorado) Agathaumas milo, Cionodon arctatus (49524) Hadrosauridae indet. (46210 49530 49539 49540 49546 49547 49549 86892 184700 219683 219684)
Late/Upper Maastrichtian70.6 - 66.0USA (New Mexico) Hadrosauridae indet. (47015 56045 59068 97795 138836 171032) Kritosaurus navajovius (76747) Lambeosaurini indet. (138833 138834) Trachodontidae indet. (62486 62488)
Late/Upper Maastrichtian70.6 - 66.0Ukraine (Crimea) Orthomerus weberi (53157) Styracosterna indet. (165811)
Late/Upper Maastrichtian70.6 - 66.0Canada (Nunavut) Hadrosauridae indet. (201443) Lambeosaurinae indet. (118411)
Late/Upper Maastrichtian70.6 - 66.0Belgium (Liège) Hadrosauridae indet. (67130 67131 67132 155974) Orthomerus dolloi (48877 48878)
Late/Upper Maastrichtian70.6 - 66.0Germany (Bayern) Hadrosauridae indet. (67197)
Late/Upper Maastrichtian70.6 - 66.0Spain (Lleida) Calvarius rapidus (230459)
Late/Upper Maastrichtian70.6 - 66.0Romania (Hunedoara) Hadrosauroidea indet. (27894) Rhabdodontidae indet. (219176) Telmatosaurus sp. (221784) Telmatosaurus sp., Rhabdodontidae indet., Zalmoxes sp., Zalmoxes shqiperorum, Iguanodontia indet. (26369) Telmatosaurus sp., Zalmoxes sp. (14454) Transylvanosaurus platycephalus (230092) Zalmoxes sp. (221769) Zalmoxes sp., Hadrosauroidea indet. (193675) Zalmoxes sp., Telmatosaurus transsylvanicus (219186)
Late/Upper Maastrichtian70.6 - 66.0Russian Federation (Amur) Hadrosauridae indet., Lambeosauridae indet., Gryposaurus asiaticus (38134) Kerberosaurus manakini, Hadrosauridae indet. (47029)
Late/Upper Maastrichtian - Danian70.6 - 61.6Slovenia (Obalno-Kraska) Iguanodontidae indet., Hadrosauridae indet. (67198)