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Hydrophiloidea (water scavenger beetle)

Insecta - Coleoptera

Synonymy list
YearName and author
1802Hydrophiloidea Latreille p. 136
1981Hydrophiloidea Crowson p. 694
1995Hydrophiloidea Lawrence and Newton, Jr.
2010Hydrophiloidea Fikáček et al.
2011Hydrophiloidea Bouchard et al. p. 153
2022Hydrophiloidea Cai et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderPolyphagaEmery 1886
infraorderStaphyliniformia
superfamilyHydrophiloidea
superfamilyHydrophiloidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
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G. †Cretotaenia Ponomarenko 1977
+
†Cretotaenia pallida Ponomarenko 1977
Fm. Helophoridae Leach 1815
+
G. Helophorus Fabricius 1775
+
+
+
Invalid names: Helophorus (Gephelophorus) lomnicki d'Orchymont 1927 [synonym], Helophorus (Trichelophorus) pleistocenicus Lomnicki 1894 [synonym]
Subg. Helophorus (Meghelophorus) Kuwert 1886
+
+
Invalid names: Helophorus (Praehelephorus) dzieduszyckii Lomnicki 1894 [synonym]
Invalid names: Helophorus (Praehelephorus) Lomnicki 1894 [synonym]
Subg. †Helophorus (Mesohelophorus) Ponomarenko 1977
+
+
Invalid names: Mesohelophorus sibiricus Ponomarenko 1977 [replaced]
Subg. †Helophorus (Mesosperchus) Ponomarenko 1977
+
†Helophorus (Mesosperchus) gracilis Prokin et al. 2010
†Helophorus (Mesosperchus) inceptivus Fikáček et al. 2012
+
Invalid names: Mesohelophorus elongatus Ponomarenko 1990 [synonym]
†Helophorus (Mesosperchus) yixianus Fikáček et al. 2012
Subg. Helophorus (Orphelophorus) d'Orchymont 1927
+
+
Invalid names: Helophorus (Orphelophorus) wandereri d'Orchymont 1927 [synonym]
Subg. Helophorus (Trichelophorus) Kuwert 1886
Helophorus auricollis Eschscholtz 1822
Helophorus browni McCorkle 1973
Helophorus eclectus d'Orchymont 1945
Helophorus lacustris Le Conte 1850
Helophorus nubilus Fabricius 1777
Helophorus oblongus Le Conte 1850
+
Invalid names: Helophorus (Rhopalhelophorus) kuwerti Lomnicki 1894 [synonym]
Helophorus orientalis Motschulsky 1860
Helophorus praenanus Lomnicki 1894
+
Invalid names: Helophorus polonicus Lomnicki 1894 [synonym]
Helophorus splendidus Sahlberg 1880
Helophorus tuberculatus Gyllenhal 1808
Fm. Hydrochidae Thomson 1859
+
G. †Ponohydrochus Prokin and Strelnikova 2021
+
†Ponohydrochus buryaticus Prokin and Strelnikova 2021
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
+
Subfm. Acidocerinae Zaitzev 1908
+
G. Helochares Mulsant 1846
+
†Helochares fog Arriaga-Varela et al. 2021
†Helochares mydaleus Zhang 1989
G. †Hydroyixia Fikáček et al. 2014
+
†Hydroyixia elongata Fikáček et al. 2014
†Hydroyixia latissima Fikáček et al. 2014
Invalid names: Horelophopsinae Hansen 1997 [synonym]
G. †Alegorius Fikáček et al. 2014
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†Alegorius yixianus Fikáček et al. 2014
G. †Aposphinctus Bode 1953
+
+
Invalid names: Spalacoides simplex Bode 1953 [synonym]
†Aposphinctus sibiricus Ponomarenko 1990
†Aposphinctus striatus Bode 1953
+
Invalid names: Hexameristus inflatus Bode 1953 [synonym]
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
Subfm. Chaetarthriinae Bedel 1881
+
Tr. Anacaenini Hansen 1991
+
G. Anacaena Thomson 1859
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†Anacaena morla Arriaga-Varela et al. 2021
†Anacaena paleodominica Fikáček and Engel 2011
G. Crenitis Bedel 1881
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†Crenitis profechuyi Arriaga-Varela et al. 2021
G. †Cretocrenis Fikáček et al. 2017
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†Cretocrenis burmanicus Fikáček et al. 2017
Invalid names: Horelophinae Hansen 1991 [synonym]
Tr. Chaetarthriini Bedel 1881
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G. Chaetarthria Stephens 1835
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G. †Creniphilites Wickham 1913
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†Creniphilites orpheus Wickham 1913
G. †Cretosperchus Ponomarenko 1997
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†Cretosperchus medievalis Ponomarenko 1997
Subfm. Enochrinae Short and Fikáček 2013
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G. Cymbiodyta Bedel 1881
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†Cymbiodyta exstincta Scudder 1900
†Cymbiodyta samueli Fikáček et al. 2023
G. Enochrus Thomson 1859
+
†Enochrus innovatus Theobald 1937
†Enochrus lapsus Zhang 1989
†Enochrus pleistocenicus Lomnicki 1894
†Enochrus primaevus Scudder 1876
†Enochrus scudderi Wickham 1909
†Enochrus striatus Theobald 1937
Invalid names: Philydrus Solier 1834 [replaced]
Subfm. Georissinae Laporte 1819
+
G. †Georissites Ponomarenko 2008
+
†Georissites magnus Haupt 1956
†Georissites negev Ponomarenko 2008
G. Georissus Latreille 1809
+
G. Hydrocharis Berthold 1827
Subfm. Hydrochinae Thomson 1859
+
G. Hydrochus Germar 1824
+
†Hydrochus amictus Scudder 1890
Hydrochus elongatus Schaller 1783
†Hydrochus relictus Scudder 1890
Subfm. Hydrophilinae Latreille 1802
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Tr. Berosini Mulsant 1844
+
G. Berosus Leach 1817
+
Subg. Berosus (Berosus) Leach 1817
†Berosus barclayi Ponomarenko and Soriano 2019
Berosus luridus Linnaeus 1761
†Berosus sexstriatus Scudder 1878
Berosus signaticollis Charpentier 1825
Berosus spinosus Steven 1808
†Berosus tenuis Scudder 1878
Tr. Hydrobiusini Mulsant 1844
+
G. †Baissalarva Fikáček et al. 2014
+
†Baissalarva hydrobioides Fikáček et al. 2014
G. †Cretoxenus Fikáček et al. 2014
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†Cretoxenus australis Fikáček et al. 2014
G. Hydrobius Linnaeus 1758
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†Hydrobius confixus Scudder 1890
Hydrobius convexus Brullé 1835
†Hydrobius couloni Heer 1862
†Hydrobius decineratus Scudder 1878
Hydrobius fuscipes Linnaeus 1758
†Hydrobius longicollis Heer 1847
†Hydrobius maceratus Scudder 1900
†Hydrobius prisconatator Wickham 1911
†Hydrobius titan Wickham 1913
G. Limnoxenus Motshulsky 1853
+
†Limnoxenus olenus Fikáček et al. 2010
Invalid names: Sperchopsini Hansen 1991 [synonym]
Tr. Hydrophilini Latreille 1802
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Invalid names: Hydrophilina Latreille 1802 [invalid subgroup], Hydrophilopsis Heer 1862 [nomen dubium], Hydrophilopsis elongata Heer 1862 [nomen dubium], Hydrous ovalis Heer 1862 [nomen dubium]
Tr. Laccobiini Houlbert 1922
+
G. Laccobius Erichson 1837
+
Laccobius ellipticus Le Conte 1855
†Laccobius elongatus Scudder 1878
†Laccobius flachii Lomnicki 1894
Laccobius minutus Linnaeus 1758
†Laccobius priscus Oustalet 1870
†Laccobius vetustus Oustalet 1874
G. Paracymus Thomson 1867
+
†Paracymus excitatus von Heyden and von Heyden 1866
Invalid names: Amphiopini Kuwert 1890 [empty], Cymbiodyta austerus Statz 1939 [nomen dubium]
G. †Orphnospercheus Lin 1992
+
G. †Paraspercheus Ponomarenko 1977
+
†Paraspercheus asiaticus Ponomarenko 1977
†Paraspercheus vitimensis Ponomarenko 1977
G. †Penitus Zhang et al. 1994
+
†Penitus gemellus Zhang et al. 1994
G. Philhydrus Solier 1834
+
†Philhydrus scudderi Wickham 1909
G. †Prospercheus Prokin 2009
+
†Prospercheus cristatus Prokin 2009
G. †Protochares Fikáček et al. 2014
+
†Protochares brevipalpis Fikáček et al. 2014
G. Regimbartia Zaitzev 1908
+
Regimbartia attenuata Fabricius 1801
Subfm. Rygmodinae Orchymont 1916
+
Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
+
Tr. Coelostomatini Heyden 1891
+
G. Coelostoma Brullé 1835
+
Coelostoma orbiculare Fabricius 1775
Coelostoma stultum Walker 1858
Tr. Megasternini Mulsant 1844
+
G. Cercyon Leach 1817
+
Cercyon analis Paykull 1798
Cercyon herceus Smetana 1978
Cercyon luniger Mannerheim 1853
Cercyon marinus Thomson 1853
Cercyon melanocephalus Linnaeus 1758
Cercyon pygmaeus Illiger 1801
Cercyon sternalis Sharp 1918
†Cercyon terrigena Scudder 1879
Cercyon tristis Illiger 1801
Cercyon ustulatus Preyssler 1790
G. Cryptopleurum Mulsant 1844
+
Cryptopleurum crenatum Kugelann 1794
Cryptopleurum minutum Fabricius 1775
G. Megasternum Mulsant 1844
+
Megasternum obscurum Marsham 1802
Tr. Sphaeridiini Latreille 1802
+
G. Sphaeridium Fabricius 1775
+
Invalid names: Omicrini Smetana 1975 [empty], Protosternini Hansen 1991 [empty]
G. Sternolophus Solier 1834
+
Sternolophus rufipes Fabricius 1792
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
G. †Hydrophilopsia Ponomarenko 1987
+
†Hydrophilopsia longitarsalis Ponomarenko 1987
G. †Laetopsia Fikáček et al. 2012
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†Laetopsia baissensis Ponomarenko 1987
†Laetopsia bontsaganica Prokin 2009
†Laetopsia hydraenoides Prokin et al. 2010
†Laetopsia mongolica Ponomarenko 1987
†Laetopsia shatrovskiyi Prokin et al. 2010
G. †Sinosperchopsis Prokin et al. 2010
+
†Sinosperchopsis silinae Prokin et al. 2010
Fm. Spercheidae Erichson 1837
+
G. Spercheus Kugelann 1798
+
†Spercheus punctatus Ponomarenko 2019
†Spercheus wightensis Ponomarenko 2019
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Early/Lower Toarcian to the top of the Holocene or 184.20000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 182.9 Ma

Collections (198 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Toarcian184.2 - 182.9Germany (Niedersachsen) Hexameristus inflatus (105002) Spalacoides simplex (107555) Streblocardioides striatus, Aposphinctus conservatus (107343)
Toarcian - Aalenian184.2 - 170.9Russian Federation (Kemerovo) Hydrophilidae indet. (117989)
Bathonian168.2 - 165.3Mongolia Hydrophilidae indet. (119621)
Callovian - Oxfordian165.3 - 154.8Kazakhstan (Chimkent) Paraspercheus asiaticus (105426 106101)
Oxfordian - Kimmeridgian161.5 - 149.2China (Yunnan) Hydrophilidae indet. (164859)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Mesosperchus schultzi (123144)
Tithonian149.2 - 143.1Russian Federation (Chita) Aposphinctus sibiricus (152172) Mesohelophorus elongatus, Aposphinctus sibiricus (125407) Mesosperchus tarsalis (119424 123197 152174) Mesosperchus tarsalis, Hydrophilopsia longitarsalis, Mesohelophorus elongatus (114307)
Tithonian149.2 - 143.1Australia (New South Wales) Protochares brevipalpis (128121)
Tithonian149.2 - 143.1Russian Federation Mesosperchus tarsalis (125408 138150)
Tithonian149.2 - 143.1Mongolia Helophorus spp., Helophorus costalis (104748) Helophorus spp., Prospercheus cristatus, Helophorus inceptivus, Helophorus costalis (121394) Prospercheus cristatus, Helophorus inceptivus (131803)
Berriasian143.1 - 137.05Israel Cretosperchus medievalis (84472)
Hauterivian132.6 - 125.77Russian Federation (Chita) Aposphinctus sibiricus (125409) Mesosperchus tarsalis (151559)
Barremian125.77 - 121.4Russian Federation (Chita) Mesosperchus tarsalis (151642)
Late/Upper Barremian125.77 - 121.4China (Nei Mongol) Alegorius yixianus (125655)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Alegorius sp., Alegorius yixianus, Hydroyixia elongata, Hydroyixia latissima (111710) Hydrophilopsia hydraenoides, Hydrophilopsia shatrovskiyi, Hydrophilopsia gracilis, Helophorus yixianus, Sinosperchopsis silinae (111716)
Early/Lower Aptian121.4 - 119.57Russian Federation (Chita) Hydrophilopsia longitarsalis (119499) Mesosperchus tarsalis (152173)
Aptian121.4 - 113.2Russian Federation (Buryatia) Cretotaenia pallida, Mesohelophorus sibiricus (105536) Hydrochidae indet., Ponohydrochus buryaticus (113350) Mesohelophorus sibiricus, Paraspercheus vitimensis, Hydrophilopsia baissensis, Baissalarva hydrobioides (105969) Ponohydrochus buryaticus (113349)
Aptian121.4 - 113.2Mongolia Hydrophilopsia bontsaganica (113355) Hydrophilopsia mongolica (103655)
Aptian121.4 - 113.2China (Jilin) Orphnospercheus lonjingensis (172391)
Aptian121.4 - 113.2China (Nei Monggol) Hydrophilidae indet. (152245)
Late/Upper Aptian119.57 - 113.2Brazil (Ceará) Hydrophilidae indet. (122557)
Late/Upper Aptian119.57 - 113.2Australia (Victoria) Hydrophilidae indet., Hydrophilidae larva indet., Cretoxenus australis (77192)
Albian113.2 - 100.5Jordan Hydrophilidae indet. (124847)
Early/Lower Cenomanian100.5 - 93.9Myanmar Cretocrenis burmanicus (161726)
Early/Lower Cenomanian100.5 - 93.9Myanmar (Kachin) Cretocrenis burmanicus (221886)
Middle Turonian93.9 - 89.8Israel Georissites negev (156985)
Danian - Selandian66.0 - 59.24Svalbard and Jan Mayen Elytridium undecimstriatum (122011)
Ypresian56.0 - 48.07Canada (British Columbia) Cercyon terrigena (122190)
Bridgerian50.5 - 46.2USA (Wyoming) Hydrochus relictus, Hydrobius confixus (122090) Laccobius elongatus, Philhydrus primaevus (121306) Tropisternus saxialis, Tropisternus sculptilis, Berosus tenuis, Berosus sexstriatus, Philhydrus sp., Hydrobius decineratus (121322)
Lutetian48.07 - 41.03Germany (Sachsen-Anhalt) Georyssus magnus (114060)
MP 1148.07 - 41.03Germany (Hessen) Hydrobiomorpha eopalpalis (165522) Hydrophilus sp., Hydrobiomorpha eopalpalis (120036) Hydrophilus sp., Hydrobiomorpha sp. 1, Hydrochara sp., Hydrophilina indet., Hydrobiomorpha eopalpalis (120798)
Priabonian37.71 - 33.9Russian Federation Anacaena morla (123419)
Priabonian37.71 - 33.9Lithuania Cymbiodyta samueli (230326)
Priabonian37.71 - 33.9United Kingdom (Isle of Wight) Berosus barclayi (156628)
Priabonian37.71 - 33.9France Enochrus striatus (113709)
Priabonian37.71 - 33.9United Kingdom (England) Berosus sp., Spercheus wightensis, Hydrochara woodwardi, Berosus barclayi (123959) Hydrochara woodwardi, Berosus barclayi (21976) Spercheus wightensis, Spercheus punctatus, Hydrochara woodwardi, Berosus barclayi (123960)
Priabonian37.71 - 33.9Poland Crenitis profechuyi, Helochares fog (110737)
Chadronian37.0 - 33.9USA (Colorado) Hydrobius prisconatator (106381 123119) Hydrobius titan, Creniphilites orpheus (113749) Philydrus scudderi (124002) Tropisternus limitatus, Tropisternus vanus, Philhydrus scudderi, Hydrobius titan, Hydrocharis extricatus, Hydrobius maceratus (106384)
MP 2333.9 - 27.3France Hydrophilidae indet., Philydrus innovatus, Laccobius vetustus (122535)
Rupelian33.9 - 27.3Germany Hydrobius convexus (122407)
Rupelian33.9 - 27.3France (Auvergne) Laccobius priscus (127630)
Chattian27.3 - 23.04Germany (North Rhine-Westphalia) Berosus capitatus, Paracymus excitatus, Hydrophilus fraternus, Hydrophilus neptunus, Hydrous rottensis, Cymbiodyta austerus, Berosus morticinus (122010)
Chattian27.3 - 23.04Brazil Hydrophilidae indet. (153054)
Chattian27.3 - 23.04France (Provence-Alpes-Côte d'Azur) Hydrophilopsis incerta (122487)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Laccobius vetustus, Hydrophilus antiquus (123944) Limnoxenus olenus (127415)
Late/Upper Oligocene27.3 - 23.04France (Puy-de-Dome) Hydrophilus magnificus (155204)
MP 3027.3 - 23.04Germany (Nordrhein-Westfalen) Hydrophilus fraternus, Hydrous miserandus (127628)
MP 3027.3 - 23.04Germany Coccinella protogaeae (115802) Philydrus morticinus, Laccobius excitatus, Hydrous neptunus (123945)
Chattian - Aquitanian27.3 - 20.45Germany (Sachsen) Hydrobiomorpha enspelense (26765)
Aquitanian23.04 - 20.45Switzerland Hydrophilus gaudini, Hydrophilus ruminianus (154053)
Aquitanian23.04 - 20.45Germany Hydrous sp., Hydrophilus sp. (123872)
Aquitanian23.04 - 20.45New Zealand Hydrophilidae indet. (50035)
Aquitanian - Burdigalian23.04 - 15.98Russian Federation Hydrophilidae indet., Helophorus sibiricus (117611)
Early/Lower Miocene23.04 - 15.98Mexico Anacaena sp. (21977)
Burdigalian20.45 - 15.98China (Shandong) Enochrus lapsus, Helochares mydaleus, Hydrous pergrandis (128552) Penitus gemellus (138945)
Burdigalian20.45 - 15.98Greece Hydrophilus vexatorius (144526)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Anacaena paleodominica (122982)
MN 320.0 - 16.9Czechia Hydrophilus pistaceus (105508)
MN 515.98 - 13.82Germany (Baden-Württemberg) Hydrophilidae indet. (109058)
Badenian13.82 - 12.8Austria (Steiermark) Hydrophilus carbonarius (115306)
Sarmatian12.8 - 11.63Germany (Wurttemberg) Hydrophilus kuntzeni, Hydrophilus caraboides (113752)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Hydrobiomorpha heeri (144245) Hydrophilus rehmanni (117338) Hydrophilus spectabilis (172253) Hydrophilus spectabilis, Hydrous escheri, Hydrobiomorpha heeri, Hydrophilus giganteus, Hydrophilopsis elongata, Hydrous ovalis, Escheria bella, Hydrous rehmanni, Hydrous braunii, Hydrobius couloni (117252) Hydrophilus vexatorius, Hydrophilus knorrii, Hydrophilus braunii (115305) Hydrophilus vexatorius, Hydrophilus spectabilis, Hydrophilus noachicus, Escheria ovalis (115304)
Sarmatian12.8 - 11.63Croatia Hydrobius longicollis (117246)
Late/Upper Miocene11.63 - 5.333Russian Federation Helophorus tuberculatus (153434) Hydrophilidae indet., Coelostoma orbiculare (113713)
Messinian7.246 - 5.333USA (Alaska) Helophorus sp., Georyssus sp. (123352)
Zanclean5.333 - 3.6Japan Coelostoma sp., Regimbartia attenuata (165737)
Pliocene5.333 - 2.58Japan Coelostoma sp., Hydrochara libera, Sternolophus rufipes (165744) Coelostoma sp., Regimbartia attenuata (165743)
Piacenzian3.6 - 2.58USA (Alaska) Georyssus sp. (162796)
Piacenzian3.6 - 2.58Japan Coelostoma sp. (165742) Hydrophilidae indet., Hydrophilus sp., Hydrochara sp., Coelostoma sp., Coelostoma stultum, Regimbartia attenuata, Sternolophus rufipes (165738)
Gelasian2.58 - 1.8Greenland Helophorus khnzoriani (160143) Helophorus spp., Helophorus khnzoriani (160144) Laccobius sp., Helophorus spp., Helophorus tuberculatus, Helophorus khnzoriani (160109) Megasternum obscurum (160166)
Gelasian2.58 - 1.8France Georyssus nigrobrunneus (113996)
Early/Lower Pleistocene2.58 - 0.774Japan Coelostoma sp. (153617)
Early/Lower Pleistocene2.58 - 0.774Russian Federation Hydrophilidae indet. (163089) Laccobius sp., Cercyon sp., Helophorus splendidus (163069)
Calabrian1.8 - 0.774Japan Hydrochara sp., Coelostoma orbiculare (160539)
Chibanian0.774 - 0.129Japan Hydrophilidae indet. (165705) Hydrophilidae indet., Coelostoma sp. (165704) Hydrophilidae indet., Hydrophilus sp., Hydrochara sp., Coelostoma sp., Hydrochara libera (165700)
Chibanian0.774 - 0.129Russian Federation Helophorus obscurellus (163093) Helophorus splendidus (163094 163097)
Chibanian0.774 - 0.129Japan (Kumamoto) Coelostoma sp. (164901)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Cercyon sp., Cryptopleurum crenatum (169391) Helophorus sp., Cercyon sp. (169389)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Coelostoma sp. (165716)
Middle Pleistocene0.774 - 0.129United Kingdom Enochrus sp., Cercyon sp., Cercyon sternalis (168272) Enochrus sp., Helophorus sp., Coelostoma orbiculare, Hydrobius fuscipes, Sphaeridium scarabaeoides, Cercyon melanocephalus, Megasternum boletophagum, Chaetarthria seminulum (168266) Enochrus sp., Helophorus sp., Georissus crenulatus, Hydrochus elongatus, Cercyon pygmaeus, Cercyon sternalis (168270) Enochrus sp., Laccobius sp., Helophorus spp., Coelostoma orbiculare, Cercyon sp., Hydrobius fuscipes, Cercyon ustulatus, Cercyon sternalis, Megasternum boletophagum, Chaetarthria seminulum (168264) Enochrus spp., Laccobius sp., Helophorus sp., Coelostoma orbiculare, Cercyon sp., Sphaeridium scarabaeoides, Georissus crenulatus, Megasternum boletophagum, Chaetarthria seminulum (168265) Helophorus sp., Coelostoma orbiculare, Sphaeridium scarabaeoides, Hydrophilus caraboides, Cercyon melanocephalus, Georissus crenulatus, Megasternum boletophagum (168267) Helophorus sp., Hydrochus elongatus, Cercyon sternalis, Megasternum boletophagum (168271) Laccobius sp., Coelostoma orbiculare, Cryptopleurum minutum, Cercyon analis, Megasternum boletophagum (168268) Paracymus sp., Helophorus spp., Coelostoma orbiculare, Hydrobius fuscipes, Cercyon analis (168269)
Late/Upper Pleistocene0.129 - 0.0117Germany Helophorus lomnicki, Helophorus wandereri (156167)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Cercyon tristis (164904 164906 164908 164921) Cercyon tristis, Helophorus brevipalpis, Hydrobius fascipes (164920) Cercyon tristis, Hydrobius fascipes (164905) Helophorus brevipalpis (164903)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Hydrophilidae indet., Enochrus sp., Hydrochara sp., Anacaena sp., Cercyon marinus, Cercyon tristis, Hydrochara caraboides, Hydrochara flavipes, Berosus luridus, Berosus spinosus, Enochrus quadripunctatus, Berosus signaticollis (125794)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Cercyon sp., Cercyon herceus (169406)
Late/Upper Pleistocene0.129 - 0.0117USA (Alaska) Helophorus splendidus (169385)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Enochrus hamiltoni (169353) Enochrus sp., Helophorus auricollis (169350) Helophorus eclectus (169347) Hydrobius fuscipes (169333 169336)
Late/Upper Pleistocene0.129 - 0.0117USA (Ohio) Hydrochus amictus (152574)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Helophorus sp., Helophorus sibiricus, Helophorus aquaticus, Helophorus praenanus, Helophorus grandis, Helophorus glacialis, Helophorus aspericollis, Helophorus orientalis, Helophorus browni, Helophorus discrepans (113753) Hydrobius fuscipes, Philhydrus pleistocenicus, Laccobius flachii, Helophorus pleistocenicus, Helophorus dzieduszyckii, Helophorus praenanus, Helophorus polonicus, Helophorus kuwerti, Sphaeridium scarabaeoides, Helophorus aquaticus, Helophorus lomnicki (153476)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Helophorus sempervarians (137210 137211 137213) Helophorus sp., Helophorus sempervarians (137208) Helophorus tuberculatus (137209) Helophorus tuberculatus, Helophorus sempervarians (137212) Hydrophilidae indet., Helophorus sempervarians (137207)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Helophorus lacustris (169359 169363) Helophorus oregonensis (169370 169372 169374 169376) Helophorus oregonensis, Helophorus lacustris (169366) Helophorus oregonensis, Helophorus sp., Helophorus lacustris (169361) Helophorus sp. (169362) Helophorus sp., Helophorus lacustris (169360) Helophorus spp., Hydrobius fuscipes, Laccobius ellipticus, Cercyon luniger (167662) Hydrophilidae indet. (169378) Hydrophilidae indet., Helophorus spp. (169375 169379) Laccobius sp., Cymbiodyta sp., Helophorus spp., Cercyon sp., Georissus sp., Hydrobius fuscipes (157092)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Helophorus splendidus (153437 153439 153443 153446 173208) Hydrobius fuscipes (153435 163102 173216)
Late/Upper Pleistocene0.129 - 0.0117USA (California) Hydrophilus sp. (44954)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Cymbiodyta exstincta (152577)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Cercyon spp., Hydrobius fuscipes, Sphaeridium sp., Helophorus aquaticus, Cercyon tristis, Helophorus nubilus, Helophorus brevipalpis, Cercyon melanocephalus, Cercyon ustulatus, Cryptopleurum minutum, Laccobius minutus, Georissus crenulatus (163070) Hydrobius fuscipes (163075)
Late/Upper Pleistocene0.129 - 0.0117Japan Coelostoma sp., Coelostoma orbiculare, Hydrochara libera (166082) Hydrophilidae indet. (164902)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Helophorus sp. (167675)
Late/Upper Pleistocene0.129 - 0.0117Peru Hydrophilus sp., Gyrinidae indet. (71346)
Tarantian0.129 - 0.0117USA (Pennsylvania) Helophorus sp. (137201) Hydrophilidae indet., Enochrus sp., Helophorus sp. (137204) Hydrophilidae indet., Enochrus sp., Helophorus sp., Helophorus sempervarians (137205 137206)
Holocene0.0117 - 0.0Brazil Sphaeridium melanarium (155717)