PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Azhdarchoidea

Reptilia - Pterosauria

Taxonomy

Azhdarchoidea was named by Unwin (1992). It was considered monophyletic by Unwin (2003).

It was reranked as the unranked clade Azhdarchoidea by Unwin and Lü (1997), Unwin (2003), Osi et al. (2005), Andres and Norell (2005), Martill and Naish (2006), Unwin and Martill (2007), Naish et al. (2013), Andres and Myers (2013), Rodrigues and Kellner (2013), Kellner et al. (2019).

It was assigned to Tapejaroidea by Andres and Norell (2005); to Pterosauria by Rodrigues and Kellner (2013); to Lophocratia by Unwin (2003), Martill and Naish (2006), Unwin and Martill (2007), Naish et al. (2013); to Ornithocheiroidea by Andres and Myers (2013), Andres et al. (2014); to Pterodactyloidea by Unwin and Lü (1997), Martill et al. (2013), Headden and Campos (2014), Pegas et al. (2016), Kellner et al. (2019); and to Pterodactyloidea by Osi et al. (2005), Agnolín et al. (2023).

Subtaxa

Azhdarchidae, Cretornis, Dsungaripteridae, Eopteranodon, Europejara, Leptostomia, Microtuban, Nemicolopterus, Neoazhdarchia, Ornithostoma, Palaeornis cliftii, Protazhdarchidae, Tapejaromorpha, Vectidraco

Synonyms

Palaeornis cliftii was named by Mantell (1844). Its type specimen is BMNH 2353 and 2353a, a limb element (Proximal and distal components of an almost complete left humerus), and it is a 3D body fossil. Its type locality is Cuckfield (general) [BMNH], which is in a Valanginian/Hauterivian terrestrial horizon in the Tunbridge Wells Sand Formation of the United Kingdom.

It was recombined as Ornithocheirus cliftii by Lydekker (1888), Brodkorb (1978) and Wellnhofer (1978); it was misspelled as Ornithocheirus clifti by Van Den Broeck (1900); it was considered a nomen dubium by Steel et al. (2005), Witton et al. (2009), Rodrigues and Kellner (2013).

Synonymy list

Year	Name and author
1844	Palaeornis cliftii Mantell
1888	Ornithocheirus cliftii Lydekker
1900	Ornithocheirus clifti Van Den Broeck p. 106
1978	Ornithocheirus cliftii Brodkorb p. 228
1978	Ornithocheirus cliftii Wellnhofer
1992	Azhdarchoidea Unwin
1997	Azhdarchoidea Unwin and Lü
2003	Azhdarchoidea Unwin p. 169
2005	Azhdarchoidea Andres and Norell pp. 3-4 fig. 3
2005	Azhdarchoidea Osi et al.
2006	Azhdarchoidea Martill and Naish p. 926
2007	Azhdarchoidea Unwin and Martill
2013	Azhdarchoidea Andres and Myers
2013	Azhdarchoidea Martill et al.
2013	Azhdarchoidea Naish et al.
2013	Azhdarchoidea Rodrigues and Kellner p. 3
2014	Azhdarchoidea Andres et al.
2014	Azhdarchoidea Headden and Campos
2016	Azhdarchoidea Pegas et al.
2019	Azhdarchoidea Kellner et al.
2023	Azhdarchoidea Agnolín et al. p. 310

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota

Rank	Name	Author
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Pterosauromorpha	Padian 1997
order	Pterosauria	Kaup 1834
suborder	Pterodactyloidea	Plieninger 1901
superfamily	Azhdarchoidea	Unwin 1992
superfamily	Azhdarchoidea	Unwin 1992

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. †Azhdarchoidea Unwin 1992

show all | hide all

Fm. †Azhdarchidae Nessov 1984

G. †Aerotitan Novas et al. 2012

†Aerotitan sudamericanus Novas et al. 2012

G. †Alanqa Ibrahim et al. 2010

†Alanqa saharica Ibrahim et al. 2010

G. †Albadraco Solomon et al. 2020

†Albadraco tharmisensis Solomon et al. 2020

G. †Aralazhdarcho Averianov 2007

†Aralazhdarcho bostobensis Averianov 2007

G. †Arambourgiania Nessov et al. 1987

†Arambourgiania philadelphiae Arambourg 1959

G. †Azhdarcho Nessov 1981

†Azhdarcho lancicollis Nessov 1984

Invalid names: Azhdarcho imparidens Nessov 1981 [replaced]

G. †Bakonydraco Osi et al. 2005

†Bakonydraco galaczi Osi et al. 2005

G. †Bennettazhia Nessov 1991

†Bennettazhia oregonensis Gilmore 1928

G. †Bogolubovia Nessov and Yarkov 1989

†Bogolubovia orientalis Bogoliubow 1914

G. †Chaoyangopterus Wang and Zhou 2003

†Chaoyangopterus zhangi Wang and Zhou 2003

G. †Cryodrakon Hone et al. 2019

†Cryodrakon boreas Hone et al. 2019

G. †Doratorhynchus Seeley 1875

G. †Eoazhdarcho Lü and Ji 2005

†Eoazhdarcho liaoxiensis Lü and Ji 2005

G. †Eurazhdarcho Vremir et al. 2013

†Eurazhdarcho langendorfensis Vremir et al. 2013

G. †Hatzegopteryx Buffetaut et al. 2002

†Hatzegopteryx thambema Buffetaut et al. 2002

G. †Mistralazhdarcho Vullo et al. 2018

†Mistralazhdarcho maggii Vullo et al. 2018

G. †Montanazhdarcho Padian et al. 1995

†Montanazhdarcho minor Padian et al. 1995

G. †Navajodactylus Sullivan and Fowler 2011

†Navajodactylus boerei Sullivan and Fowler 2011

G. †Phosphatodraco Pereda Suberbiola et al. 2003

†Phosphatodraco mauritanicus Pereda Suberbiola et al. 2003

Subfm. †Quetzalcoatlinae Andres et al. 2014

G. †Infernodrakon Thomas et al. 2025

†Infernodrakon hastacollis Thomas et al. 2025

G. †Quetzalcoatlus Lawson 1975

†Quetzalcoatlus lawsoni Andres and Langston Jr 2021

†Quetzalcoatlus northropi Lawson 1975

G. †Thanatosdrakon Ortiz David et al. 2022

†Thanatosdrakon amaru Ortiz David et al. 2022

G. †Wellnhopterus Andres and Langston Jr 2021

†Wellnhopterus brevirostris Andres and Langston Jr 2021

G. †Titanopteryx Arambourg 1959

G. †Zhejiangopterus Cai and Wei 1994

†Zhejiangopterus linhaiensis Cai and Wei 1994

G. †Cretornis Fritsch 1881

†Cretornis hlavaci Fritsch 1881

Fm. †Dsungaripteridae Young 1964

G. †Banguela Headden and Campos 2014

†Banguela oberlii Headden and Campos 2014

G. †Domeykodactylus Martill et al. 2000

†Domeykodactylus ceciliae Martill et al. 2000

G. †Dsungaripterus Young 1964

†Dsungaripterus weii Young 1964

G. †Lonchognathosaurus Maisch et al. 2004

†Lonchognathosaurus acutirostris Maisch et al. 2004

G. †Noripterus Young 1973

†Noripterus complicidens Young 1973

†Noripterus parvus Bakhurina 1982

Invalid names: Phobetor Bakhurina 1986 [synonym]

G. †Eopteranodon Lü and Zhang 2005

†Eopteranodon lii Lü and Zhang 2005

G. †Europejara Vullo et al. 2012

†Europejara olcadesorum Vullo et al. 2012

G. †Leptostomia Smith et al. 2020

†Leptostomia begaaensis Smith et al. 2020

G. †Microtuban Elgin and Frey 2011

†Microtuban altivolans Elgin and Frey 2011

G. †Nemicolopterus Wang et al. 2008 [forest dweller wing]

†Nemicolopterus crypticus Wang et al. 2008

Unr. †Neoazhdarchia Unwin 2003

Fm. †Chaoyangopteridae Lü et al. 2008

G. †Apatorhamphus McPhee et al. 2020

†Apatorhamphus gyrostega McPhee et al. 2020

G. †Lacusovagus Witton 2008

†Lacusovagus magnificens Witton 2008

G. †Shenzhoupterus Lü et al. 2008

†Shenzhoupterus chaoyangensis Lü et al. 2008

G. †Radiodactylus Andres and Myers 2013

†Radiodactylus langstoni Andres and Myers 2013

Superfm. †Tapejaroidea Kellner 1989

Fm. †Tapejaridae Kellner 1989

G. †Afrotapejara Martill et al. 2020

†Afrotapejara zouhri Martill et al. 2020

Unr. †Chaoyangopterinae Pinheiro et al. 2011

G. †Jidapterus Dong et al. 2003

G. †Huaxiapterus Lü and Yuan 2005

†Huaxiapterus benxiensis Lü et al. 2007

†Huaxiapterus corollatus Lü et al. 2006

†Huaxiapterus jii Lü and Yuan 2005

G. †Ingridia Unwin and Martill 2007

G. †Sinopterus Wang and Zhou 2002

†Sinopterus dongi Wang and Zhou 2002

†Sinopterus gui Li et al. 2003

G. †Tapejara Kellner 1989

†Tapejara wellnhoferi Kellner 1989

Subfm. †Tapejarinae Kellner 1989

G. †Aymberedactylus Pegas et al. 2016

G. †Caiuajara Manzig et al. 2014

G. †Caupedactylus Kellner 2013

Subfm. †Thalassodrominae Kellner and Campos 2007

G. †Kariridraco Cerqueira et al. 2021

G. †Tupandactylus Kellner and Campos 2007

†Tupandactylus imperator Campos and Kellner 1997

†Tupandactylus navigans Frey et al. 2003

G. †Wightia Martill et al. 2020

†Wightia declivirostris Martill et al. 2020

Fm. †Thalassodromidae Kellner and Campos 2007

G. †Thalassodromeus Kellner and Campos 2002

†Thalassodromeus sethi Kellner and Campos 2002

G. †Tupuxuara Kellner and Campos 1988

†Tupuxuara deliradamus Witton 2009

†Tupuxuara leonardii Kellner and Campos 1994

†Tupuxuara longicristatus Kellner and Campos 1988

G. †Xericeps Martill et al. 2018

†Xericeps curvirostris Martill et al. 2018

G. †Ornithostoma Seeley 1871

†Ornithostoma sedgwicki Seeley 1871

Invalid names: Ornithostoma seeleyi Lydekker 1904 [synonym]

Fm. †Protazhdarchidae Frey et al. 2011

G. †Aurorazhdarcho Frey et al. 2011

†Aurorazhdarcho primordius Frey et al. 2011

Unr. †Tapejaromorpha Andres et al. 2014

G. †Keresdrakon Kellner et al. 2019

†Keresdrakon vilsoni Kellner et al. 2019

G. †Vectidraco Naish et al. 2013

†Vectidraco daisymorrisae Naish et al. 2013

Invalid names: Palaeornis cliftii Mantell 1844 [nomen dubium]

Diagnosis

Reference	Diagnosis
D. M. Unwin 2003	Tapejara wellnhoferi, Quetzalcoatlus northropi, their most recent common ancestor, and all its descendants

Measurements

No measurements are available

Composition:

phosphaticsubp

Maximum body mass:

544 kgs

Minimum body mass:

259 kgs

Environment:

marineo

Locomotion:

actively mobilec

Life habit:

volanto

Diet:

piscivoreo

Created:

2009-01-04 11:38:30

Modified:

2017-04-17 10:39:23

Source: s = species, o = order, c = class, subp = subphylum

References: Witton 2008, Padian 1985, Carroll 1988, Hendy et al. 2009

Age range: base of the Late/Upper Kimmeridgian to the top of the Early/Lower Maastrichtian or 152.21000 to 66.00000 Ma

Collections (136 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Kimmeridgian	152.21 - 149.2	Germany (Niedersachsen)	Dsungaripteridae indet. (51858)
Early/Lower Tithonian	149.2 - 145.06	Germany (Bayern)	Aurorazhdarcho primordius (138219)
Early/Lower Tithonian	149.2 - 145.06	France (Poitou-Charentes)	Azhdarchidae indet. (164377)
Tithonian	149.2 - 143.1	Tanzania (Lindi)	Tapejaroidea indet. (89996)
Tithonian - Berriasian	149.2 - 137.05	Thailand (Kalasin)	Azhdarchoidea indet. (157421)
Berriasian	143.1 - 137.05	Romania (Judetul Bihor)	Azhdarchidae indet., Ornithocheiridae indet. (13335)
Berriasian - Valanginian	143.1 - 132.6	China (Shandong)	Theropoda indet. (47070)
Early/Lower Cretaceous	143.1 - 100.5	Chile (Antofagasta)	Pterodaustro sp. (92228)
Early/Lower Cretaceous	143.1 - 100.5	Mongolia (Ovorkhangai)	Tapejaroidea indet. (70140)
Early/Lower Cretaceous	143.1 - 100.5	United Kingdom (Southeast England)	Azhdarchoidea indet. (232501)
Late/Upper Berriasian	139.1 - 137.05	United Kingdom (England)	Azhdarchidae indet. (80391)
Valanginian	137.05 - 132.6	China (Xinjiang)	Dsungaripterus sp. (220472)
Late/Upper Valanginian - Early/Lower Hauterivian	137.05 - 125.77	United Kingdom (England)	Palaeornis cliftii (73272)
Valanginian - Barremian	137.05 - 121.4	Mongolia (Uvs)	Noripterus complicidens (91411) Noripterus complicidens, Dsungaripterus weii (91412)
Valanginian - Barremian	137.05 - 121.4	Mongolia (Ovorhangai)	Dsungaripteridae indet., Phobetor parvus, Dsungaripterus parvus (55238)
Valanginian - Albian	137.05 - 100.5	China (Xinjiang)	Lonchognathosaurus acutirostris (89993) Noripterus complicidens (49817 49818) Noripterus complicidens, Dsungaripterus weii (49047 49819) Ornithurae indet. (71847)
Hauterivian - Barremian	132.6 - 121.4	Japan (Gifu)	Dsungaripteridae indet. (89998)
Late/Upper Hauterivian - Aptian	132.6 - 113.2	Japan (Gifu)	Dsungaripteridae indet. (50310)
Barremian	125.77 - 121.4	United Kingdom (England)	Neoazhdarchia indet. (90213) Wightia declivirostris (101091)
Late/Upper Barremian	125.77 - 121.4	Spain (Castilla-La Mancha)	Europejara olcadesorum (28006)
Late/Upper Barremian - Early/Lower Aptian	125.77 - 119.57	United Kingdom (England)	Vectidraco daisymorrisae (90306)
Late/Upper Barremian - Early/Lower Aptian	125.77 - 119.57	China (Liaoning)	Eopteranodon lii (92903) Eopteranodon yixianensis (79522)
Early/Lower Aptian	121.4 - 119.57	China (Shandong)	Azhdarchidae indet. (100364)
Aptian	121.4 - 113.2	China (Shandong)	Pterosauria indet. (55475)
Aptian	121.4 - 113.2	China (Liaoning)	Chaoyangopterus zhangi (79523 79941) Huaxiapterus benxiensis (92905) Nemicolopterus crypticus (78922) Sinopterus dongi (92898) Sinopterus dongi, Sinopterus gui, Huaxiapterus jii, Huaxiapterus corollatus, Eoazhdarcho liaoxiensis, Shenzhoupterus chaoyangensis, Jidapterus edentus (92652)
Aptian - Campanian	121.4 - 72.2	Brazil (Paraná)	Caiuajara dobruskii, Keresdrakon vilsoni (160057)
Late/Upper Aptian	119.57 - 113.2	Brazil (Ceará)	Caupedactylus ybaka (114920) Tupandactylus imperator (102227) Tupuxuara longicristatus, Tapejara wellnhoferi, Azhdarchoidea indet., Thalassodrominae indet., Kariridraco dianae, Thalassodromeus sethi (92290) Tupuxuara sp., Tupuxuara longicristatus, Thalassodromeus sethi, Tupuxuara leonardii, Tupuxuara deliradamus (67712)
Late/Upper Aptian	119.57 - 113.2	Brazil (Ceara)	Tapejara imperator, Tapejaridae indet., Tapejara navigans, Thalassodrominae indet., Lacusovagus magnificens, Aymberedactylus cearensis, Azhdarchidae indet. (77891)
Early/Lower Albian	113.2 - 110.1	USA (Texas)	Azhdarchidae indet. (84273)
Albian	113.2 - 100.5	France (Champagne-Ardenne)	Azhdarchidae indet. (141003)
Albian	113.2 - 100.5	USA (Oregon)	Bennettazhia oregonensis (229463) Pteranodon oregonensis (92236)
Late/Upper Albian	106.3 - 100.5	United Kingdom (England)	Ornithostoma sedgwicki (94210)
Late/Upper Albian - Early/Lower Cenomanian	106.3 - 93.9	Morocco (Drâa-Tafilalet)	Afrotapejara zouhri (208784) Azhdarchidae indet., Azhdarchidae indet., Dsungaripteroidea indet. (52887)
Early/Lower Cenomanian	100.5 - 93.9	Argentina (Río Negro)	Azhdarchidae indet. (132347)
Early/Lower Cenomanian	100.5 - 93.9	Morocco (Drâa-Tafilalet)	Alanqa saharica (96222) Alanqa saharica, Leptostomia begaaensis, Apatorhamphus gyrostega (209058) Azhdarchidae indet. (90079) Azhdarchidae indet., Alanqa saharica, Xericeps curvirostris (96223) Azhdarchidae indet., Tapejaridae indet., Pteranodontidae indet. (90076)
Early/Lower Cenomanian	100.5 - 93.9	Uzbekistan (Karakalpakstan)	Azhdarchidae indet. (58466 92327)
Late/Upper Cenomanian	100.5 - 93.9	Lebanon	Microtuban altivolans (138218)
Late/Upper Cenomanian - Early/Lower Turonian	100.5 - 89.8	Japan (Kumamoto)	Azhdarchidae indet. (93995)
Cenomanian - Santonian	100.5 - 83.6	Mongolia (Dornogov)	Azhdarchidae indet. (55758)
Late/Upper Turonian	93.9 - 89.8	Armenia (Vayots Dzor)	Azhdarchidae indet. (92185)
Middle Turonian - Late/Upper Turonian	93.9 - 89.8	Uzbekistan (Navoi)	Azhdarcho imparidens (75943)
Middle Turonian - Late/Upper Turonian	93.9 - 89.8	Uzbekistan (Xorazm)	Azhdarchidae indet. (92328)
Middle Turonian - Late/Upper Turonian	93.9 - 89.8	Czechia (Pardubice)	Cretornis hlavaci (93874)
Late/Upper Turonian - Coniacian	93.9 - 85.7	Kazakhstan (Qyzylorda)	Azhdarchidae indet. (58594)
Late/Upper Turonian - Early/Lower Coniacian	93.9 - 85.7	Argentina (Neuquén)	Azhdarchidae indet. (49415)
Late/Upper Turonian - Late/Upper Santonian	93.9 - 83.6	Mongolia (Omnogov)	Azhdarchidae indet. (55254)
Turonian - Early/Lower Campanian	93.9 - 72.2	Kenya (Rift Valley)	Azhdarchidae indet. (106492)
Late/Upper Coniacian - Early/Lower Santonian	89.8 - 83.6	Argentina (Mendoza)	Thanatosdrakon amaru (219506)
Early/Lower Santonian	85.7 - 83.6	Tajikistan (Sogd)	Azhdarcho sp., Azhdarchidae indet. (58701)
Santonian	85.7 - 83.6	Hungary (Veszprém)	Bakonydraco galaczi, Pterosauria indet. (48722)
Santonian - Early/Lower Campanian	85.7 - 72.2	Kazakhstan (Qyzylorda)	Pterosauria indet. (53134)
Campanian	83.6 - 72.2	Mongolia (Omnogov)	Azhdarchidae indet. (168986)
Early/Lower Campanian	83.6 - 72.2	Russian Federation (Saratov)	Azhdarchidae indet. (92330)
Early/Lower Campanian	83.6 - 72.2	Russian Federation (Orenburg)	Azhdarchidae indet. (224094)
Early/Lower Campanian	83.6 - 72.2	China (Zhejiang)	Zhejiangopterus linhaiensis (92188)
Early/Lower Campanian	83.6 - 72.2	Russian Federation (Penza)	Azhdarchidae indet., Ornithostoma orientalis (92329)
Early/Lower Campanian	83.6 - 72.2	Austria (Niederosterreich)	Ornithocheirus sp. (48720)
Late/Upper Campanian	83.6 - 72.2	Russian Federation (Saratov)	Azhdarchidae indet. (92359)
Late/Upper Campanian	83.6 - 72.2	Senegal (Thiès)	Quetzalcoatlus sp. (92227)
Late/Upper Campanian	83.6 - 72.2	USA (New Mexico)	Navajodactylus boerei (138830 138831)
Late/Upper Campanian	83.6 - 72.2	Canada (Alberta)	Azhdarchidae indet. (52656 94765 189085) Quetzalcoatlus sp. (36264)
Late/Upper Campanian	83.6 - 72.2	Canada (British Columbia)	Azhdarchidae indet. (132556)
Late/Upper Campanian	83.6 - 72.2	Israel (Southern District)	Titanopteryx sp. (92192)
Late/Upper Campanian	83.6 - 72.2	France (Provence-Alpes-Côte d'Azur)	Azhdarchidae indet. (28454) Mistralazhdarcho maggii (220134)
Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Azhdarchidae indet. (22689) Montanazhdarcho minor (90189) Pterodactyloidea indet. (36934)
Late/Upper Campanian	83.6 - 72.2	USA (Utah)	Azhdarchidae indet. (217077)
Late/Upper Campanian	83.6 - 72.2	Romania (Sebeş)	Azhdarchidae indet. B (219187)
Late/Upper Campanian	83.6 - 72.2	Spain (Castilla y Leon)	Azhdarchidae indet. (28406) Azhdarcho sp. (28765)
Middle Campanian - Late/Upper Campanian	83.6 - 72.2	USA (Montana)	Azhdarchidae indet. (14462)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	France (Occitanie)	Azhdarchidae indet. (35290)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Argentina (Río Negro)	Aerotitan sudamericanus (135703)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	Spain (Valencia)	Azhdarchidae indet. (96461)
Late/Upper Campanian - Early/Lower Maastrichtian	83.6 - 66.0	France (Languedoc-Roussillon)	Azhdarchidae indet. (48870 59887)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Alba)	Albadraco tharmisensis (115074) Azhdarchidae indet. A (93729) Azhdarchidae indet. C (138892) Azhdarchoidea indet. (153475) Hatzegopteryx sp. (97813)
Early/Lower Maastrichtian	72.2 - 66.0	Romania (Hunedoara)	Azhdarchidae indet. (193773) Hatzegopteryx thambema (89991) Pterosauria indet. (13334)
Late/Upper Maastrichtian	72.2 - 66.0	Australia (Western Australia)	Azhdarchidae indet. (92270)
Late/Upper Maastrichtian	72.2 - 66.0	France (Provence-Alpes-Côte d'Azur)	Pterosauria indet. (48866)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Montana)	Infernodrakon hastacollis (239512)
Late/Upper Maastrichtian	72.2 - 66.0	Morocco (Khouribga)	Azhdarchidae indet., Phosphatodraco mauritanicus (193774)
Late/Upper Maastrichtian	72.2 - 66.0	USA (Texas)	Azhdarchidae indet. (230207) Pterosauria indet. (92249 92251 230208) Quetzalcoatlus northropi (85191 230205 230206) Quetzalcoatlus sp. (52090 85190 92247) Wellnhopterus brevirostris (230204)
Late/Upper Maastrichtian	72.2 - 66.0	Romania (Hunedoara)	Azhdarchidae indet. (167488) Hatzegopteryx sp. (26369)
Late/Upper Maastrichtian	72.2 - 66.0	Spain (Valencia)	Azhdarchidae indet. (47035)
Late/Upper Maastrichtian	72.2 - 66.0	Morocco (Chaouia-Ouardigha)	Phosphatodraco mauritanicus (92203)
Late/Upper Maastrichtian	72.2 - 66.0	Morocco (Béni Mellal-Khénifra)	Azhdarchidae indet. (193775)
Maastrichtian	72.2 - 66.0	Jordan	Titanopteryx philadelphiae, Arambourgiania philadelphiae (13366)
Maastrichtian	72.2 - 66.0	USA (New Jersey)	Titanopteryx sp. (78124)
Maastrichtian	72.2 - 66.0	Mongolia (Omnogov)	Azhdarchidae indet. (168417)