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Bittacidae (hangingfly)

Insecta - Mecoptera - Bittacidae

Synonymy list
YearName and author
1906Bittacidae Handlirsch p. 43
1978Bittacidae Willmann p. 97
1987Bittacidae Willmann
1992Bittacidae Carpenter
1996Bittacidae Ansorge p. 73
2004Bittacidae Petrulevičius and Jarzembowski p. 1198
2018Bittacidae Kopeć et al. p. 158

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
RankNameAuthor
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
superorderAntliophora
orderMecopteraPackard 1886
suborderPistillifera
infraorderRaptipedia
familyBittacidaeHandlirsch 1906
familyBittacidaeHandlirsch 1906

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Bittacidae Handlirsch 1906 [hangingfly]
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G. †Antiquanabittacus Petrulevičius and Jarzembowski 2004
+
†Antiquanabittacus dispar Bashkuev 2023
†Antiquanabittacus nanus Petrulevičius and Jarzembowski 2004
†Antiquanabittacus robustus Bashkuev 2023
†Antiquanabittacus saltator Bashkuev 2023
G. †Archebittacus Riek 1955
+
†Archebittacus exilis Riek 1955
G. †Baissobittacus Novokshonov 1997
+
†Baissobittacus angustipennatus Novokshonov 1997
Subfm. †Bittacinae Handlirsch 1906
+
G. †Asiobittacus Novokshonov 1993
+
†Asiobittacus sajsagulis Novokshonov 1993
G. †Mesobittacus Handlirsch 1939
+
+
Invalid names: Auxobittacus praeacutus Bode 1953 [synonym]
†Mesobittacus kubekovensis Novokshonov 1993
†Mesobittacus minutus Handlirsch 1939
+
Invalid names: Protobittacus pediculatus Bode 1953 [synonym]
Invalid names: Auxobittacus Bode 1953 [synonym], Microbittacus Novokshonov 1993 [synonym]
G. Orobittacus Villegas and Byers 1981
+
†Orobittacus antiquus Novokshonov 1993
†Orobittacus minutus Novokshonov 1993
G. Bittacus Latreille 1802
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†Bittacus biamensis Novokshonov 1993
†Bittacus egestionis Carpenter 1955
†Bittacus lepiduscretaceus Li et al. 2018
†Bittacus succinus Carpenter 1954
†Bittacus veternus Cockerell 1921
G. †Burmobittacus Zhao et al. 2016
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†Burmobittacus jarzembowskii Zhao et al. 2016
G. †Composibittacus Liu et al. 2016
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†Composibittacus bipunctatus Liu et al. 2016
†Composibittacus reticulatus Liu et al. 2016
G. †Cretobittacus Novokshonov 1993
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†Cretobittacus baisensis Novokshonov 1993
G. †Decoribittacus Li and Ren 2009
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†Decoribittacus euneurus Li and Ren 2009
†Decoribittacus stictus Li and Ren 2009
G. Eremobittacus Byers 1997
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†Eremobittacus opeth Poschmann and Nel 2022
G. †Exilibittacus Yang et al. 2012
+
†Exilibittacus foliaceus Liu et al. 2014
†Exilibittacus lii Yang et al. 2012
†Exilibittacus plagioneurus Liu et al. 2014
G. †Formosibittacus Li et al. 2008
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†Formosibittacus macularis Li et al. 2008
G. Hylobittacus Byers 1979
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†Hylobittacus antiquus Pictet and Hagen 1856
†Hylobittacus fossilis Carpenter 1954
†Hylobittacus minimus Carpenter 1954
†Hylobittacus picteti Krzemiński 2007
Invalid names: Electrobittacus Carpenter 1931 [synonym]
G. †Jurahylobittacus Li et al. 2008
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†Jurahylobittacus astictus Li et al. 2008
G. †Karattacus Novokshonov 1997
+
†Karattacus longialatus Li and Ren 2009
†Karattacus persibus Novokshonov 1997
G. †Liaobittacus Ren 1993
+
G. †Megabittacus Ren 1997
+
†Megabittacus spatiosus Yang et al. 2012
G. †Mongolobittacus Petrulevičius et al. 2007
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†Mongolobittacus daohugouensis Petrulevičius et al. 2007
†Mongolobittacus oligophlebius Liu et al. 2014
†Mongolobittacus speciosus Liu et al. 2014
Subfm. †Neorthophlebiinae Handlirsch 1920
+
G. †Ctenophlebia Hong 2009
+
G. †Haplobittacus Bode 1953
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†Haplobittacus parvus Bode 1953
G. †Neorthophlebia Handlirsch 1906
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†Neorthophlebia acutipennis Handlirsch 1939
†Neorthophlebia debilis Handlirsch 1906
†Neorthophlebia maculipennis Handlirsch 1906
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Invalid names: Metaxybittacus vittatus Bode 1953 [synonym], Neorthophlebia minor Handlirsch 1906 [synonym]
†Neorthophlebia medialis Handlirsch 1939
†Neorthophlebia nana Martynova 1948
†Neorthophlebia pallida Handlirsch 1939
†Neorthophlebia robusta Martynov 1937
†Neorthophlebia simillima Handlirsch 1939
†Neorthophlebia stigmatica Handlirsch 1939
†Neorthophlebia unica Martynova 1948
†Neorthophlebia yunnanensis Zhang and Hong 2003
G. †Neorthophlebopsis Hong et al. 2005
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†Neorthophlebopsis qishuiheensis Hong et al. 2005
G. †Parabittacus Handlirsch 1939
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†Parabittacus analis Handlirsch 1939
†Parabittacus lingula Bode 1953
Invalid names: Polydicrobittacus Bode 1953 [synonym]
G. †Pleobittacus Bode 1953
+
G. †Protobittacus Tillyard 1933
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†Protobittacus handlirschi Tillyard 1933
†Protobittacus liassicus Tillyard 1933
†Protobittacus maculatus Tillyard 1933
†Protobittacus moestus Sukatsheva 1985
G. †Orthobittacus Willmann 1989
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†Orthobittacus abshiricus Martynova 1951
†Orthobittacus maculosus Liu et al. 2016
†Orthobittacus polymitus Novokshonov 1996
†Orthobittacus suni Kopeć et al. 2018
G. †Paekthobittacus Ri et al. 2023
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†Paekthobittacus sinuijuensis Ri et al. 2023
G. †Palaeobittacus Carpenter 1928
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†Palaeobittacus eocenicus Carpenter 1928
Subfm. †Plesiobittacinae Novokshonov 1997
+
G. †Plesiobittacus Novokshonov 1997
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†Plesiobittacus martynovi Novokshonov 1997
†Plesiobittacus primigenius Novokshonov 1997
G. †Preanabittacus Novokshonov 1993
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†Preanabittacus karatavensis Novokshonov 1993
†Preanabittacus validus Yang et al. 2012
G. †Probittacus Martynov 1927
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†Probittacus avitus Martynov 1927
G. †Prohylobittacus Novokshonov 1993
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†Prohylobittacus sorosi Novokshonov 1993
G. †Scharabittacus Novokshonov 1993
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†Scharabittacus karatavicus Novokshonov 1993
†Scharabittacus mongolicus Novokshonov 1993
†Scharabittacus pictus Martynova 1949
G. †Sibirobittacus Sukacheva 1990
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†Sibirobittacus kazakhstanensis Novokshonov 1993
†Sibirobittacus undus Sukacheva 1990
G. Thyridates Navas 1908
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†Thyridates novokschonovi Petrulevičius 2003
G. Tytthobittacus Smithers 1973
+
†Tytthobittacus jarzembowski Kopeć et al. 2016
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Longobardian to the top of the MP 28 or 242.00000 to 23.03000 Ma
Minimum age of oldest fossil (stem group age): 235.0 Ma

Collections (65 total)


Time interval Ma Country or state Original ID and collection number
Longobardian242.0 - 235.0China (Shaanxi) Ctenophlebia tongchuanensis, Neorthophlebopsis qishuiheensis (117138)
Norian228.0 - 208.5Australia (Queensland) Archebittacus exilis (106484 191411)
Rhaetian208.5 - 201.3United Kingdom (England) Protobittacus liassicus, Protobittacus maculatus (113961)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Neorthophlebia robusta, Neorthophlebia unica, Neorthophlebia nana (105087)
Pliensbachian190.8 - 182.7Kyrgyzstan Neorthophlebia abshirica (160469)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Auxobittacus marginelaesus (107322) Auxobittacus praeacutus (107343) Polydicrobittacus lingula (107553) Polydicrobittacus lingula, Haplobittacus parvus (107321) Protobittacus arculatus, Protobittacus pediculatus, Auxobittacus praeacutus, Metaxybittacus vittatus, Pleobittacus retroflexus, Polydicrobittacus lingula (105002) Protobittacus clavaeformis (107323) Protobittacus desacuminatus (107555)
Early/Lower Toarcian183.0 - 182.0Germany Neorthophlebia minor, Mesobittacus minutus, Orthophlebia megapolitana, Neorthophlebia debilis, Parabittacus analis, Neorthophlebia pallida, Neorthophlebia simillima, Neorthophlebia medialis, Neorthophlebia stigmatica, Neorthophlebia acutipennis, Orthophlebia megapolitana (123987)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Bittacidae indet., Neorthophlebia maculipennis, Mesobittacus minutus (114993)
Falciferum183.0 - 182.0United Kingdom (England) Bittacidae indet. (234543) Protobittacus handlirschi (154307)
Falciferum183.0 - 182.0Germany Bittacidae indet. (192037)
Toarcian182.7 - 174.1Russian Federation (Kemerovo) Protobittacus moestus (125484)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Asiobittacus sajsagulis, Plesiobittacus martynovi, Plesiobittacus primigenius (105003)
Toarcian182.7 - 174.1Kyrgyzstan Neorthophlebia robusta (114702)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Microbittacus kubekovensis (118010)
Callovian - Oxfordian166.1 - 157.3China (Liaoning) Liaobittacus longantennatus (107004)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Karattacus persibus, Orobittacus antiquus (105971) Orthobittacus polymitus, Probittacus karatavicus (105426) Preanabittacus karatavensis, Probittacus karatavicus (106101) Probittacus avitus (114692) Prohylobittacus sorosi (113439) Prohylobittacus sorosi, Orobittacus antiquus (105970) Sibirobittacus kazakhstanensis (106102)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Composibittacus bipunctatus, Composibittacus reticulatus, Orthobittacus maculosus (123761) Exilibittacus lii (137887) Formosibittacus macularis, Jurahylobittacus astictus, Decoribittacus euneurus, Decoribittacus stictus, Karattacus longialatus (112664) Mongolobittacus daohugouensis (113371) Mongolobittacus speciosus, Mongolobittacus oligophlebius, Exilibittacus foliaceus, Exilibittacus plagioneurus (155751) Orthobittacus suni (183155) Preanabittacus validus (113366)
Oxfordian - Kimmeridgian163.5 - 152.1China (Yunnan) Neorthophlebia yunnanensis (164859)
Late/Upper Jurassic161.5 - 145.0Russian Federation (Buryatia) Neorthophlebia indet. (125483)
Tithonian152.1 - 145.0Russian Federation (Chita) Sibirobittacus undus (123197)
Tithonian152.1 - 145.0Mongolia Orobittacus minutus (104748) Probittacus mongolicus (121393)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Tytthobittacus jarzembowski (104007)
Late/Upper Hauterivian136.4 - 130.0United Kingdom (England) Antiquanabittacus nanus (106417)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Megabittacus colosseus, Megabittacus beipiaoensis, Sibirobittacus atalus (138436) Megabittacus spatiosus (140491)
Barremian - Aptian129.4 - 113.0North Korea Paekthobittacus sinuijuensis (200272)
Aptian125.0 - 113.0Russian Federation (Buryatia) Antiquanabittacus saltator, Antiquanabittacus dispar (113349) Antiquanabittacus sp., Antiquanabittacus saltator, Antiquanabittacus robustus, Antiquanabittacus dispar (113350) Baissobittacus angustipennatus, Cretobittacus baisensis (105969)
Early/Lower Cenomanian99.6 - 93.5Myanmar Bittacus lepiduscretaceus (178121) Burmobittacus jarzembowskii (161726)
Late/Upper Paleocene58.7 - 55.8Argentina (Jujuy) Thyridates novokschonovi (113004)
Bridgerian50.3 - 46.2USA (Colorado) Palaeobittacus eocenicus (163519)
Bridgerian50.3 - 46.2USA (Utah) Bittacus egestionis (122882)
Priabonian38.0 - 33.9United Kingdom (England) Bittacus sp.A, Panorpa veterna, Bittacus veternus (123960)
Priabonian38.0 - 33.9Russian Federation (Primorskiy) Bittacus biamensis (123794)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Bittacus fossilis, Bittacus minimus, Bittacus succinus, Hylobittacus antiquus (124205) Bittacus minimus (123766)
Priabonian38.0 - 33.9Russian Federation Hylobittacus fossilis, Hylobittacus minimus, Hylobittacus picteti (123419)
Priabonian38.0 - 33.9Poland Bittacus antiquus (109625) Hylobittacus antiquus, Hylobittacus fossilis, Hylobittacus minimus, Hylobittacus picteti (123215) Hylobittacus fossilis (120813)
Priabonian38.0 - 33.9Germany (Halle) Hylobittacus picteti (124273)
MP 2828.4 - 23.03Germany (Rheinland-Pfalz) Eremobittacus opeth (160097)