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Chrysidoidea

Insecta - Hymenoptera

Synonyms
Synonymy list
YearName and author
1802Chrysidoidea Latreille p. 316
1975Bethyloidea Rasnitsyn p. 104
1999Chrysidoidea Brothers p. 233
2003Chrysidoidea Wappler p. 118
2011Chrysidoidea Heraty et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
RankNameAuthor
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Hymenopterida
orderHymenoptera
—Unicalcarida
suborderApocritaGerstaecker 1867
—Aculeata()
superfamilyChrysidoidea
superfamilyChrysidoidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Chrysidoidea Latreille 1802
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Fm. Bethylidae Forster 1856 [flat wasp]
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Subfm. Bethylinae Halliday 1840
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G. Bethylus Latreille 1802
G. †Crassibethylus Barbosa and Melo 2023
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†Crassibethylus dominicanus Barbosa and Melo 2023
G. †Cretobethylellus Rasnitsyn 1990
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†Cretobethylellus lucidus Rasnitsyn 1990
G. Eupsenella Westwood 1874
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†Eupsenella aulax Ramos and Azevedo 2014
†Eupsenella eocenica De Ploëg and Nel 2004
†Eupsenella klesoviana Ramos and Azevedo 2014
†Eupsenella rossica Ramos and Azevedo 2014
†Eupsenella yantarnica Ramos and Azevedo 2014
Invalid names: Eupsenella neoeocenica Ramos and Azevedo 2018 [nomen nudum], Eupsenella neohongi Ramos and Azevedo 2018 [nomen nudum], Fushunochrysites Hong 2002 [nomen nudum], Protobethylus De Ploëg and Nel 2004 [synonym], Sinibethylus Hong 2002 [nomen nudum]
G. Goniozus Förster 1856
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†Goniozus antiquus Olmi 2005
†Goniozus contractus Brues 1933
†Goniozus copalinus Meunier 1916
†Goniozus cotyi Brazidec and Perrichot 2022
†Goniozus definitus Ramos and Azevedo 2014
†Goniozus respectus Sorg 1988
Invalid names: Messoria Meunier 1916 [synonym], Perisierola Kieffer 1914 [synonym], Pterosclerogibba Olmi 2005 [synonym]
G. Lytopsenella Kieffer 1911
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†Lytopsenella baltica Ramos and Azevedo 2014
†Lytopsenella crastina Brues 1923
†Lytopsenella maritima Ramos and Azevedo 2014
†Lytopsenella setigera Brues 1923
†Lytopsenella simplex Brues 1923
G. †Nucifrangibulum Cockx et al. 2016
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†Nucifrangibulum carentonensis Cockx et al. 2016
G. Prosierola Kieffer 1905
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†Prosierola submersa Brues 1933
G. Sierola Cameron 1881
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†Sierola colomboi Brazidec and Perrichot 2022
†Sierola hastata Sorg 1988
†Sierola rovniana Ramos and Azevedo 2014
Subfm. †Cretabythinae Brazidec et al. 2023
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G. †Cretabythus Evans 1973
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†Cretabythus sibiricus Evans 1973
G. †Holopsenelliscus Engel 2019
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G. †Megalopsenella Jouault et al. 2020
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†Megalopsenella pouilloni Jouault et al. 2020
Subfm. †Elektroepyrinae Colombo and Azevedo 2020
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G. †Elektroepyris Perrichot and Nel 2008
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†Elektroepyris magnificus Perrichot and Nel 2008
Subfm. Epyrinae Kieffer 1914
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G. Anisepyris Kieffer 1905
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†Anisepyris gradates Sorg 1988
†Anisepyris gradatus Sorg 1988
G. Bakeriella Kieffer 1910
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†Bakeriella nanyhelae Brazidec and Perrichot 2022
G. Calyoza Westwood 1837
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Invalid names: Calyoza ramosa Meunier 1906 [synonym]
Invalid names: Artiepyris Kieffer 1913 [synonym]
G. Chlorepyris Kieffer 1913
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†Chlorepyris concaptus Brues 1933
†Chlorepyris deploegi Colombo and Azevedo 2021
†Chlorepyris engeli Colombo and Azevedo 2021
†Chlorepyris gallicus Perrichot and Nel 2008
†Chlorepyris hopei Colombo and Azevedo 2021
†Chlorepyris invelatus Brues 1933
†Chlorepyris jouaulti Colombo and Azevedo 2021
†Chlorepyris mckellari Colombo and Azevedo 2021
†Chlorepyris meunieri Colombo and Azevedo 2021
†Chlorepyris neli Colombo and Azevedo 2021
†Chlorepyris perrichoti Colombo and Azevedo 2021
†Chlorepyris setosus Brues 1933
G. Dolus Motschulsky 1863
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Invalid names: Homoglenus Kieffer 1904 [synonym]
G. Epyris Westwood 1832
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†Epyris deletus Brues 1910
†Epyris kiefferi Brues 1933
†Epyris moulyi Falieres and Nel 2018
†Epyris rectinervis Cockerell 1921
†Epyris tenellus Statz 1938
Invalid names: Isobrachium Forster 1856 [synonym], Rhabdepyris Kieffer 1904 [synonym]
G. †Gloxinius Colombo and Azevedo 2021
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†Gloxinius bifossatus Brues 1939
G. Holepyris Kieffer 1904
G. Laelius Ashmead 1893
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†Laelius pallidus Brues 1933
†Laelius preteritus Barbosa and Azevedo 2013
†Laelius rovnensis Barbosa and Azevedo 2013
G. Rysepyris Kieffer 1906
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†Rysepyris minor Brues 1939
†Rysepyris neodubius Azevedo 2018
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Invalid names: Misepyris dubius Brues 1933 [replaced]
†Rysepyris planiceps Brues 1933
†Rysepyris precursor Brues 1933
†Rysepyris rasnitsyni Colombo and Azevedo 2021
†Rysepyris robustus Brues 1933
†Rysepyris terayamai Colombo and Azevedo 2021
Invalid names: Misepyris Kieffer 1913 [synonym]
Subfm. †Lancepyrinae Azevedo and Azar 2012
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G. †Archaepyris Evans 1973
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†Archaepyris minutus Evans 1973
G. †Azepyris Brazidec et al. 2023
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†Azepyris delamarrei Brazidec et al. 2023
G. †Burmapyris Jouault et al. 2021
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†Burmapyris azevedoi Jouault et al. 2021
†Burmapyris ohmkuhnlei Brazidec et al. 2023
G. †Cretepyris Ortega-Blanco and Engel 2013
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†Cretepyris martini Ortega-Blanco and Engel 2013
G. †Gwesped Brazidec et al. 2023
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†Gwesped groehni Brazidec et al. 2023
G. †Lancepyris Azevedo and Azar 2012
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†Lancepyris opertus Azevedo and Azar 2012
G. †Liztor Ortega-Blanco and Engel 2013
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†Liztor pilosus Ortega-Blanco and Engel 2013
G. †Paralanceis Brazidec et al. 2023
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†Paralanceis chotardi Brazidec et al. 2023
G. †Protopyris Jouault and Nel 2021
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†Protopyris myanmarensis Jouault and Nel 2021
G. †Yunbayin Brazidec et al. 2023
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†Yunbayin rossei Brazidec et al. 2023
G. †Zophepyris Engel et al. 2016
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†Zophepyris alavaensis Ortega-Blanco and Engel 2013
Subfm. Mesitiinae Kieffer 1914
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G. Mesitius Spinola 1851
G. †Nadezhdabythus Zhang et al. 2020
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†Nadezhdabythus burmensis Zhang et al. 2020
Subfm. Pristocerinae Mocsary 1881
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G. †Ajkanesia Szabó and Brazidec 2022
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†Ajkanesia harmincipsziloni Szabó and Brazidec 2022
G. Apenesia Westwood 1874
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†Apenesia miki Terayama 2004
Invalid names: Archaeopristocera Terayama 2004 [synonym]
G. Cleistepyris Kieffer 1910
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†Cleistepyris allosaurus Colombo et al. 2020
†Cleistepyris baryonyx Colombo et al. 2020
G. Dissomphalus Ashmead 1893
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†Dissomphalus schubnelli Falières and Nel 2020
G. †Ekaterina Colombo et al. 2020
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†Ekaterina volgatitan Colombo et al. 2020
G. Eleganesia Alencar and Azevedo 2018
G. †Eopristocera Falières and Nel 2019
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†Eopristocera bilobata Falières and Nel 2019
G. Foenobethylus Kieffer 1913
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†Foenobethylus electriphilus Cockerell 1917
G. †Merascylla Colombo and Azevedo 2021
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†Merascylla atavella Cockerell 1920
G. Parascleroderma Kieffer 1904
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†Parascleroderma palaeosinica Brazidec and Perrichot 2022
G. Pristocera Klug 1808
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†Pristocera alaini Falières and Nel 2019
†Pristocera skwarrae Brues 1933
Invalid names: Parapristocera Brues 1933 [synonym]
G. Pseudisobrachium Kieffer 1904
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†Pseudisobrachium megalosaurus Colombo et al. 2020
†Pseudisobrachium stegosaurus Colombo et al. 2020
G. †Uniceratops Colombo et al. 2020
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†Uniceratops trex Colombo et al. 2020
Subfm. †Protopristocerinae Nagy 1974
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G. †Bethylitella Cockerell 1917
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†Bethylitella cylindrella Cockerell 1917
G. †Bethylopteron Brues 1933
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†Bethylopteron ambiguum Brues 1933
G. †Cretapristocera Jouault et al. 2020
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†Cretapristocera longiscapa Jouault et al. 2020
G. †Gynopteron Falières and Nel 2019
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†Gynopteron inexpectatus Falières and Nel 2019
G. †Protopristocera Brues 1923
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†Protopristocera sucini Brues 1923
G. †Ramageoptera Jouault and Brazidec 2021
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†Ramageoptera platycephala Jouault and Brazidec 2021
Subfm. Scleroderminae Kieffer 1914
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G. Allobethylus Kieffer 1905
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†Allobethylus bei Colombo and Azevedo 2019
G. Alongatepyris Azevedo 1992
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†Alongatepyris pedrocai Colombo and Azevedo 2022
G. †Amissidigitus Szabó and Brazidec 2022
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†Amissidigitus belae Szabó and Brazidec 2022
G. †Archaeonoxus Colombo and Azevedo 2019
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†Archaeonoxus scintillatus Colombo and Azevedo 2019
G. †Celonophamia Evans 1973
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†Celonophamia granama McKellar and Engel 2014
†Celonophamia taimyria Evans 1973
G. Glenosema Kieffer 1905
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†Glenosema electrum Colombo and Azevedo 2019
G. †Mael Colombo and Azevedo 2021
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†Mael longiceps Brues 1923
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Invalid names: Epyris neolongiceps Azevedo 2018 [objective synonym]
G. Nothepyris Evans 1973
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†Nothepyris pretiosus Colombo and Azevedo 2019
G. †Paleoscleroderma Falières and Nel 2019
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†Paleoscleroderma lamarrei Falières and Nel 2019
G. Plastanoxus Kieffer 1905
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†Plastanoxus atrescens Sorg 1988
G. Sclerodermus Latreille 1809
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†Sclerodermus quadridentatum Cockerell 1917
†Sclerodermus quadridentatus Cockerell 1917
G. Solepyris Azevedo 2006
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†Solepyris electromexicanus Brazidec and Perrichot 2022
G. †Voronknoxus Colombo and Perkovsky 2022
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†Voronknoxus zerovae Colombo and Perkovsky 2022
Invalid names: Epyrinae Kieffer 1914 [invalid subgroup], Fushunochrysidae Hong 2002 [synonym]
Fm. Chrysididae Latreille 1802 [cuckoo wasp]
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Subfm. Amiseginae Mocsary 1889
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G. Atoposega Krombein 1960
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†Atoposega rosai Brazidec and Perrichot 2022
G. †Eosega Martynova 2017
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†Eosega heterotarsata Martynova 2017
G. †Foveorisus Martynova 2017
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†Foveorisus kilimniki Martynova 2017
G. †Palaeobethyloides Brues 1933
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G. †Palaeobethylus Brues 1923
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†Palaeobethylus politus Brues 1933
G. †Protadelphe Krombein 1986
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†Protadelphe aenea Krombein 1986
G. †Protamisega Evans 1973
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†Protamisega khatanga Evans 1973
G. †Uromesitius Brues 1933
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†Uromesitius caudatus Brues 1933
G. †Auricleptes Lucena and Melo 2018
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†Auricleptes nebulosus Lucena and Melo 2018
G. †Burmasega Lucena and Melo 2018
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†Burmasega ammirabilis Lucena and Melo 2018
Subfm. Chrysidinae Latreille 1802
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G. †Azanichrum Lucena and Melo 2018
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†Azanichrum pilosum Lucena and Melo 2018
G. †Bohartiura Lucena and Melo 2018
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†Bohartiura glabrata Lucena and Melo 2018
Tr. Chrysidini Latreille 1802
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G. Ceratochrysis Cooper 1952
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G. Chrysis Linnaeus 1761
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†Chrysis amoena Förster 1891
†Chrysis miocenica Rohwer 1909
†Chrysis rohweri Cockerell 1907
†Chrysis viridicyanea Giebel 1862
G. Euchroeus Latreille 1809
G. Primeuchroeus Linsenmaier 1968
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†Primeuchroeus groehni Brazidec et al. 2024
Tr. Elampini Dahlbom 1854
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G. Hedychridium Abeille de Perrin 1878
G. Omalus Panzer 1801
G. †Palaeochrum Krombein 1986
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†Palaeochrum diversum Krombein 1986
G. †Sphaerocleptes Cockx et al. 2016
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†Sphaerocleptes neraudeaui Cockx et al. 2016
Subfm. Cleptinae Latreille 1802
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G. Cleptes Latreille 1802
G. †Hypocleptes Evans 1973
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†Hypocleptes rasnitsyni Evans 1973
G. †Procleptes Evans 1969
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†Procleptes carpenteri Evans 1969
†Procleptes eoliami McKellar and Engel 2014
†Procleptes hopejohnsonae McKellar and Engel 2014
G. †Dahurochrysis Rasnitsyn 1990
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†Dahurochrysis veta Rasnitsyn 1990
G. †Miracorium Lucena and Melo 2018
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†Miracorium tetrafoveolatum Lucena and Melo 2018
G. †Protochrysidis Carpenter 1986
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†Protochrysidis succinalis Bischoff 1916
Invalid names: Eochrysis Doweld 2015 [objective synonym], Protochrysis Bischoff 1916 [replaced]
Fm. †Chrysobythidae Melo and Lucena 2019
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G. †Aureobythus Melo and Lucena 2019
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†Aureobythus decoloratus Melo and Lucena 2019
†Aureobythus punctatus Melo and Lucena 2019
†Aureobythus villosus Melo and Lucena 2019
G. †Bethylochrysis Melo and Lucena 2019
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†Bethylochrysis clypeata Melo and Lucena 2019
G. †Chrysobythus Melo and Lucena 2019
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†Chrysobythus areolatus Melo and Lucena 2019
Fm. Dryinidae Haliday 1833
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Subfm. Anteoninae Kieffer 1907
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G. †Anteonopsis Olmi et al. 2010
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†Anteonopsis antiquus Olmi et al. 2010
G. †Burmanteon Engel 2003
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†Burmanteon olmii Engel 2003
G. Deinodryinus Perkins 1907
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†Deinodryinus aptianus Olmi et al. 2010
†Deinodryinus areolatus Ponomareko 1975
†Deinodryinus velteni Guglielmino and Olmi 2011
Invalid names: Electrodryinus Ponomareko 1975 [synonym]
G. †Janzeniola Olmi 2009
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†Janzeniola baltica Olmi 2000
Invalid names: Janzenia Olmi 2000 [replaced]
G. Lonchodryinus Kieffer 1905
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†Lonchodryinus balticus Olmi and Guglielmino 2012
†Lonchodryinus groehni Olmi et al. 2021
Subfm. Aphelopinae Perkins 1912
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G. Aphelopus Dalman 1823
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†Aphelopus poinari Olmi 1998
Subfm. Apodryininae Olmi 1984
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Tr. †Rovnodryinini Olmi et al. 2022
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G. †Rovnodryinus Olmi et al. 2022
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†Rovnodryinus khomychi Olmi et al. 2022
Subfm. †Archaeodryininae Olmi et al. 2020
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G. †Archaeodryinus Olmi et al. 2020
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Subfm. Bocchinae Richards 1939
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G. Bocchus Ashmead 1893
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†Bocchus primaevus Martins and Melo 2019
†Bocchus rex Olmi et al. 2022
†Bocchus schmalhauseni Perkovsky et al. 2020
†Bocchus vetustus Olmi 1991
Subfm. †Burmadryininae Olmi et al. 2014
+
G. †Burmadryinus Olmi et al. 2014
+
†Burmadryinus cenomanianus Olmi et al. 2014
Subfm. Dryininae Haliday 1833
+
G. †Cretodryinus Ponomarenko 1975
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†Cretodryinus zherichini Ponomarenko 1975
G. Dryinus Latreille 1804
+
†Dryinus acutifrons Martynova et al. 2019
†Dryinus alamellatus Olmi and Guglielmino 2011
†Dryinus antiquus Ponomarenko 1981
†Dryinus arakanesis Martynova et al. 2019
†Dryinus balticus Olmi 1984
†Dryinus bamar Martynova et al. 2019
†Dryinus brachytarsis Martynova et al. 2019
†Dryinus bruesi Olmi 1984
†Dryinus burmensis Martynova et al. 2019
†Dryinus canadensis Ponomarenko 1981
†Dryinus carsteni Olmi et al. 2022
†Dryinus chin Martynova et al. 2019
†Dryinus grimaldii Olmi 1995
†Dryinus janzeni Olmi 2000
†Dryinus katyae Martynova et al. 2019
†Dryinus laminatus Martynova et al. 2019
†Dryinus latitarsis Martynova et al. 2019
†Dryinus maderai Guglielmino et al. 2018
†Dryinus mortuorum Brues 1933
†Dryinus muenchi Olmi and Bechly 2001
†Dryinus nadezhdae Martynova et al. 2019
†Dryinus palaeodominicanus Currado and Olmi 1983
†Dryinus parisei Martynova et al. 2019
†Dryinus poinari Olmi 1998
†Dryinus pristinus Olmi 1998
†Dryinus rasnitsyni Olmi and Guglielmino 2011
†Dryinus reifi Olmi and Bechly 2001
†Dryinus taron Martynova et al. 2019
†Dryinus velteni Olmi 2011
†Dryinus vetus Brues 1933
†Dryinus vetustus Olmi 1995
†Dryinus wunderlichi Olmi and Bechly 2001
Invalid names: Avodryinus Ponomarenko 1981 [synonym], Richardsidryinus Moczar 1965 [synonym]
G. †Hybristodryinus Engel 2005
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†Hybristodryinus anomalus Perkovsky et al. 2019
†Hybristodryinus castaneus Olmi et al. 2021
†Hybristodryinus concavifrons Perkovsky et al. 2019
†Hybristodryinus cretacicus Perkovsky et al. 2019
†Hybristodryinus karen Perkovsky et al. 2019
†Hybristodryinus kayin Perkovsky et al. 2019
†Hybristodryinus konbaung Perkovsky et al. 2019
†Hybristodryinus ligulatus Perkovsky et al. 2019
†Hybristodryinus magnificus Perkovsky et al. 2019
†Hybristodryinus mon Perkovsky et al. 2019
†Hybristodryinus moutesoe Tribull et al. 2020
†Hybristodryinus nalae Perkovsky et al. 2019
†Hybristodryinus pyu Perkovsky et al. 2019
†Hybristodryinus shan Perkovsky et al. 2019
†Hybristodryinus viriosus Wang et al. 2021
†Hybristodryinus zaifui Chen et al. 2021
G. †Palaeodryinus Olmi and Bechly 2001
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†Palaeodryinus groehni Olmi and Bechly 2001
G. Pseudodryinus Olmi 1991
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†Pseudodryinus burmensis Olmi et al. 2014
†Pseudodryinus parisiensis Peinado et al. 2006
G. †Rasnitsynum Olmi et al. 2021
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†Rasnitsynum burmense Olmi et al. 2021
Invalid names: Dryinini Haliday 1833 [invalid subgroup]
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G. Gonatopus Ljungh 1810
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†Gonatopus cretacicus Olmi et al. 2010
Invalid names: Dicondylus Curtis 1829 [synonym], Laberius Kieffer 1914 [synonym]
G. Neodryinus Perkins 1905
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†Neodryinus somniatus Brues 1933
Subfm. †Palaeoanteoninae Olmi and Bechly 2001
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G. †Palaeoanteon Olmi 2000
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†Palaeoanteon cenomanianum Perkovsky et al. 2020
†Palaeoanteon janzeni Olmi 2000
Subfm. †Protodryininae Olmi and Guglielmino 2012
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G. †Protodryinus Olmi and Guglielmino 2012
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†Protodryinus eocenicus Olmi and Guglielmino 2012
Subfm. †Raptodryininae Olmi et al. 2020
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G. †Raptodryinus Olmi et al. 2020
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†Raptodryinus patrickmuelleri Olmi et al. 2020
Subfm. Thaumatodryininae Perkins 1905
+
G. Thaumatodryinus Perkins 1905
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†Thaumatodryinus fuscescens Martins and Melo 2020
G. †Thaumatorrhinos Martins and Melo 2024
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†Thaumatorrhinos athrix Martins and Melo 2024
Fm. Embolemidae Forster 1856
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G. Ampulicomorpha Ashmead 1893
+
†Ampulicomorpha janzeni Olmi et al. 2014
†Ampulicomorpha perialla Ortega-Blanco et al. 2011
†Ampulicomorpha quesnoyensis Chény et al. 2020
†Ampulicomorpha testacea Olmi et al. 2020
G. †Baissobius Rasnitsyn 1975
+
†Baissobius carolianus Rasnitsyn 1996
†Baissobius minimus Rasnitsyn 1996
†Baissobius minutus Olmi et al. 2010
†Baissobius parvus Rasnitsyn 1975
G. †Cretembolemus Olmi et al. 2014
+
†Cretembolemus orapensis Olmi et al. 2014
G. †Embolemopsis Olmi et al. 2010
+
†Embolemopsis baissensis Olmi et al. 2010
†Embolemopsis maryannae Olmi et al. 2021
G. Embolemus Westwood 1833
+
†Embolemus antiquus Perkovsky et al. 2021
†Embolemus brachypterus Olmi et al. 2021
†Embolemus burmensis Perkovsky et al. 2021
†Embolemus cretacicus Perkovsky et al. 2021
†Embolemus excitus Perrichot and Engel 2011
†Embolemus micropterus Olmi et al. 2021
†Embolemus ohmkuhnlei Perkovsky et al. 2021
†Embolemus zherikhini Perkovsky et al. 2021
G. †Ponomarenkoa Olmi 2010
+
†Ponomarenkoa burmensis Perkovsky et al. 2021
†Ponomarenkoa ellenbergeri Olmi et al. 2013
†Ponomarenkoa polonica Ponomarenko 1988
Invalid names: Laberites Ponomarenko 1988 [replaced]
Invalid names: Embolemus breviscapus Brues 1933 [nomen dubium], Ponomarenkoinae Olmi 2010 [invalid subgroup]
Fm. †Plumalexiidae Brothers 2011
+
G. †Plumalexius Brothers 2011
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†Plumalexius ohmkuhnlei Rasnitsyn and Brothers 2020
†Plumalexius rasnitsyni Brothers 2011
Fm. †Radiophronidae Ortega-Blanco et al. 2010
+
G. †Microcostaphron Ortega-Blanco et al. 2010
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†Microcostaphron parvus Ortega-Blanco et al. 2010
G. †Radiophron Ortega-Blanco et al. 2010
+
†Radiophron ibericus Ortega-Blanco et al. 2010
Fm. Sclerogibbidae Ashmead 1902 [sclerogibbid wasp]
+
G. †Burmasclerogibba Perkovsky et al. 2020
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†Burmasclerogibba aptera Perkovsky et al. 2020
G. †Chosia Fujiyama 1994
+
†Chosia yamadai Fujiyama 1994
G. †Cretosclerogibba Perkovsky et al. 2020
+
†Cretosclerogibba antennalis Perkovsky et al. 2020
†Cretosclerogibba contractocollis Perkovsky et al. 2020
†Cretosclerogibba neli Perkovsky et al. 2020
†Cretosclerogibba rasnitsyni Perkovsky et al. 2020
G. †Edrossia Perkovsky et al. 2020
+
†Edrossia vetusta Perkovsky et al. 2020
G. †Gallosclerogibba Perrichot et al. 2020
+
†Gallosclerogibba alnensis Perrichot et al. 2020
G. Sclerogibba Riggio and De Stefani-Perez 1888
+
†Sclerogibba cretacica Martynova et al. 2019
Subfm. Sclerogibbinae Ashmead 1902
+
G. Probethylus Ashmead 1902
+
†Probethylus poinari Olmi 2005
Subfm. †Sclerogibbodinae Engel and Grimaldi 2006
+
G. †Sclerogibbodes Engel and Grimaldi 2006
+
†Sclerogibbodes embioleia Engel and Grimaldi 2006
Fm. Scolebythidae Evans 1963
+
G. †Acanthabythus Lepeco and Melo 2022
+
†Acanthabythus exilispineus Lepeco and Melo 2022
G. †Arcapenesia Lepeco and Melo 2022
+
†Arcapenesia compacta Lepeco and Melo 2022
G. †Burmapenesia Lepeco and Melo 2022
+
†Burmapenesia iridescens Lepeco and Melo 2022
†Burmapenesia longicephala Lepeco and Melo 2022
G. †Cephalobythus Lepeco and Melo 2022
+
†Cephalobythus clypeatus Lepeco and Melo 2022
†Cephalobythus deformis Lepeco and Melo 2022
†Cephalobythus gladiator Lepeco and Melo 2022
†Cephalobythus simplex Lepeco and Melo 2022
G. †Cretopsenella Lepeco and Melo 2022
+
†Cretopsenella spinifera Lepeco and Melo 2022
G. †Gnathapenesia Lepeco and Melo 2022
+
†Gnathapenesia burmensis Lepeco and Melo 2022
Subfm. Pristapenesiinae Engel et al. 2013
+
G. †Boreobythus Engel and Grimaldi 2007
+
†Boreobythus turonius Engel and Grimaldi 2007
G. †Cursoribythus Cockx and McKellar 2016
+
†Cursoribythus silvestris Cockx and McKellar 2016
G. †Ectenobythus Engel et al. 2013
+
†Ectenobythus iberiensis Engel et al. 2013
G. †Eobythus Lacau et al. 2000
+
†Eobythus patriciae Lacau et al. 2000
G. †Libanobythus Prentice and Poinar 1996
+
†Libanobythus milkii Prentice and Poinar 1996
G. †Necrobythus Engel et al. 2013
+
†Necrobythus pulcher Engel et al. 2013
G. Pristapenesia Brues 1933
+
†Pristapenesia inopinata Prentice and Poinar 1996
†Pristapenesia primaeva Brues 1933
Invalid names: Dominibythus Prentice and Poinar 1996 [synonym]
G. †Sphakelobythus Engel et al. 2013
+
†Sphakelobythus limnopous Engel et al. 2013
G. †Uliobythus Engel and Grimaldi 2007
+
†Uliobythus terpsichore Engel and Grimaldi 2007
G. †Zapenesia Engel and Grimaldi 2007
+
†Zapenesia libanica Engel and Grimaldi 2007
G. †Rhynchopsenella Lepeco and Melo 2022
+
†Rhynchopsenella argentea Lepeco and Melo 2022
†Rhynchopsenella bicarinata Lepeco and Melo 2022
†Rhynchopsenella diaphana Lepeco and Melo 2022
Subfm. Scolebythinae Evans 1963
+
G. Clystopsenella Kieffer 1911
+
G. Scolebythus Evans 1963
Invalid names: Bethyloidea [synonym], Plumariidae [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Late/Upper Barremian to the top of the Holocene or 130.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 125.45 Ma

Collections (127 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Barremian130.0 - 125.45United Kingdom Embolemopsis maryannae (198093)
Early/Lower Barremian130.0 - 125.45Spain Bethylidae indet. (104052)
Early/Lower Barremian130.0 - 125.45Lebanon Aphelopus palaeophoenicius (124363) Lancepyris opertus (138801) Libanobythus milkii (124925) Sclerogibbodes embioleia (122359) Zapenesia libanica, Uliobythus terpsichore (122471)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Scolebythidae indet. (138325)
Aptian125.0 - 113.0Mongolia Baissobius carolianus, Deinodryinus aptianus (113355)
Aptian125.0 - 113.0Russian Federation (Buryatia) Baissobius minimus (157328) Baissobius parvus (105536) Embolemopsis baissensis, Baissobius minutus (118216)
Aptian125.0 - 113.0Russian Federation Cretobethylellus lucidus (124811)
Late/Upper Aptian122.46 - 113.0Australia (Victoria) Bethyloidea indet. (77192)
Early/Lower Aptian121.4 - 122.46Japan (Chiba) Chosia yamadai (157033)
Early/Lower Aptian121.4 - 122.46Russian Federation (Chita) Dahurochrysis veta (119499)
Albian113.0 - 100.5Jordan Bethylidae indet. (124847)
Early/Lower Albian112.03 - 109.0Myanmar (Sagaing) Ampulicomorpha testacea (228880) Azepyris delamarrei (229858) Paralanceis chotardi, Yunbayin rossei (215115)
Late/Upper Albian105.3 - 99.6Spain (Álava) Scolebythidae indet. 1, Scolebythidae indet. 2, Embolemus periallus, Radiophron ibericus, Microcostaphron parvus, Lancepyris alavaensis, Liztor pilosus, Cretepyris martini, Ectenobythus iberiensis (108769)
Late/Upper Albian - Early/Lower Cenomanian105.3 - 93.5France (Charente-Maritime) Scolebythidae indet. (107076)
Early/Lower Cenomanian99.6 - 93.5Myanmar Auricleptes nebulosus, Burmasega ammirabilis, Miracorium tetrafoveolatum, Azanichrum pilosum, Bohartiura glabrata, Aureobythus decoloratus, Aureobythus punctatus, Aureobythus villosus, Bethylochrysis clypeata, Chrysobythus areolatus (192864) Burmanteon olmii, Hybristodryinus moutesoe (111688) Burmapyris azevedoi, Protopyris myanmarensis, Cretapristocera longiscapa, Ramageoptera platycephala, Burmapyris ohmkuhnlei, Yunbayin rossei (179227) Cretosclerogibba neli, Cretosclerogibba rasnitsyni (198285) Cursoribythus silvestris (166234) Dryinus taron (206272) Embolemus sp., Apenesia electriphila, Bethylitella cylindrella, Epyris atavellus, Scleroderma quadridentatum (123020) Gwesped groehni, Dryinus carsteni (172900) Hukawngepyris setosus (184841) Hybristodryinus cretacicus, Dryinus taron, Hybristodryinus zaifui, Hybristodryinus viriosus (165561) Hybristodryinus resinicolus, Dryinus maderai, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus nalae, Hybristodryinus pyu, Dryinus bamar, Dryinus katyae, Dryinus parisei (184692) Hybristodryinus resinicolus, Holopsenelliscus pankowskiorum (123116) Megalopsenella pouilloni (212258) Plumalexius ohmkuhnlei, Embolemus ohmkuhnlei, Embolemus zherikhini (194182) Ponomarenkoa ellenbergeri, Ampulicomorpha janzeni, Dryinus maderai, Hybristodryinus anomalus, Hybristodryinus concavifrons, Hybristodryinus cretacicus, Hybristodryinus karen, Hybristodryinus kayin, Hybristodryinus konbaung, Hybristodryinus ligulatus, Hybristodryinus magnificus, Hybristodryinus mon, Hybristodryinus nalae, Hybristodryinus pyu, Hybristodryinus shan, Sclerogibba cretacica, Dryinus acutifrons, Dryinus arakanesis, Dryinus bamar, Dryinus brachytarsis, Dryinus chin, Dryinus laminatus, Dryinus latitarsis, Dryinus nadezhdae, Dryinus parisei, Dryinus taron, Raptodryinus patrickmuelleri, Palaeoanteon cenomanianum, Hybristodryinus castaneus, Embolemus antiquus, Embolemus burmensis, Embolemus cretacicus, Ponomarenkoa burmensis, Embolemus brachypterus, Embolemus micropterus, Rasnitsynum burmense (194884) Ponomarenkoa ellenbergeri, Ampulicomorpha janzeni, Pseudodryinus burmensis, Burmadryinus cenomanianus (123053) Radiophronidae indet., Sclerogibbidae indet., Dryinus burmensis, Dryinus katyae, Burmasclerogibba aptera, Cretosclerogibba antennalis, Cretosclerogibba contractocollis, Edrossia vetusta, Cretosclerogibba neli, Cretosclerogibba rasnitsyni, Nadezhdabythus burmensis (161726) Thaumatorrhinos athrix, Plumalexius ohmkuhnlei, Acanthabythus exilispineus, Arcapenesia compacta, Burmapenesia iridescens, Burmapenesia longicephala, Cephalobythus clypeatus, Cephalobythus deformis, Cephalobythus gladiator, Cephalobythus simplex, Cretopsenella spinifera, Gnathapenesia burmensis, Rhynchopsenella argentea, Rhynchopsenella bicarinata, Rhynchopsenella diaphana (216576)
Early/Lower Cenomanian99.6 - 93.5Myanmar (Kachin) Uniceratops trex (221971)
Early/Lower Cenomanian99.6 - 93.5France (Poitou-Charentes) Bethylidae indet., Nucifrangibulum carentonensis, Gallosclerogibba alnensis (161514) Sphaerocleptes neraudeaui (180993)
Late/Upper Cenomanian99.6 - 93.5Russian Federation Laberius antiquus (35051)
Late/Upper Cenomanian99.6 - 93.5Morocco (Er Rachida) Chrysididae indet. (120457)
Turonian93.9 - 89.8USA (New Jersey) Plumalexius rasnitsyni, Boreobythus turonius (110895)
Turonian93.9 - 89.8Botswana Cretembolemus orapensis (123430)
Santonian86.3 - 83.6Hungary Ajkanesia harmincipsziloni, Amissidigitus belae (224054)
Late/Upper Santonian85.8 - 83.5Russian Federation Archaepyris minutus, Celonophamia taimyria, Hypocleptes rasnitsyni, Ekaterina volgatitan (35056) Cretodryinus zherichini, Cretabythus sibiricus, Archaepyris minutus, Hypocleptes rasnitsyni, Protamisega khatanga (122590)
Early/Lower Campanian83.5 - 70.6Russian Federation (Tenkinskii) Gonatopus cretacicus, Anteonopsis antiquus (105375)
Late/Upper Campanian83.5 - 70.6Canada (Manitoba) Procleptes carpenteri (122354)
Middle Campanian83.5 - 70.6Canada (Alberta) Avodryinus canadensis (122384) Bethylidae indet., Celonophamia granama (122383) Chrysididae indet. (104932) Cleptinae indet., Necrobythus pulcher, Sphakelobythus limnopous, Procleptes hopejohnsonae (122357) Procleptes eoliami (122389)
Ypresian56.0 - 47.8France (Oise) Eobythus patriciae, Protobethylus eocenicus, Epyrini indet., Rhabdepyris gallicus, Elektroepyris magnificus, Pseudodryinus parisiensis, Paleoscleroderma lamarrei, Pristocera alaini, Ampulicomorpha quesnoyensis, Dissomphalus schubnelli, Epyris moulyi, Eopristocera sp., Eopristocera bilobata, Gynopteron inexpectatus (123850) Scolebythidae indet. (68174)
Ypresian56.0 - 47.8Canada (British Columbia) Chrysidinae indet. (192140)
Ypresian56.0 - 47.8China (Liaoning) Fushunochrysites eocenicus (124530) Sinibethylus eocenicus (124820)
Bridgerian50.3 - 46.2USA (Colorado) Eupsenella sp. (229578)
MP 1348.6 - 40.4Germany (Eifel) Bethylidae indet. (135641)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Allobethylus bei (219707) Ampulicomorpha succinalis (218978) Ampulicomorpha succinalis, Dryinus muenchi, Dryinus reifi, Dryinus wunderlichi, Palaeoanteon janzeni, Harpactosphecion filicorne, Deinodryinus velteni, Dryinus bruesi, Dryinus janzeni, Lonchodryinus balticus, Protodryinus eocenicus (120814) Ampulicomorpha succinalis, Lytopsenella kerneggeri, Electrodryinus areolatus, Eupsenella rossica, Eupsenella yantarnica, Eosega heterotarsata, Chlorepyris engeli, Chlorepyris hopei, Chlorepyris meunieri, Chlorepyris perrichoti, Holepyris rasnitsyni, Archaeonoxus scintillatus, Glenosema electrum, Nothepyris pretiosus, Cleistepyris baryonyx (122591) Ampulicomorpha succinalis, Thaumatodryinus gracilis, Thaumatodryinus filicornis, Palaeobethylus longicollis, Calyoza longiceps, Protopristocera sucini, Epyris inhabilis, Sierola simplex, Sierola setigera, Sierola crastina, Dryinus filicornis, Omalus primordialis, Embolemus breviscapus, Uromesitius caudatus, Palaeobethylus politus, Palaeobethylus brevicollis, Palaeobethyloides longiceps, Bethylopteron ambiguum, Parapristocera skwarrae, Eupsenella setigera, Eupsenella simplex, Eupsenella crastina, Prosierola submersa, Perisierola contracta, Artiepyris kiefferi, Pristapenesia primaeva, Misepyris robustus, Misepyris dubius, Holepyris planiceps, Holepyris precursor, Isobrachium concaptum, Isobrachium invelatum, Rhabdepyris setosus, Rhabdepyris elatus, Laelius pallidus, Thaumatodryinus deletus, Lestodryinus mortuorum, Lestodryinus vetus, Neodryinus somniatus, Protochrysis succinalis (123927) Bocchus primaevus (217266) Dryinus mortuorum, Harpactosphecion filicornis (156497) Dryinus sp., Palaeoanteon janzeni, Janzenia baltica, Dryinus janzeni, Harpactosphecion filicornis (154633) Embolemus excitus (123928) Eupsenella aulax, Lytopsenella baltica, Lytopsenella maritima (160471) Goniozus definitus (120610) Harpactosphecion filicorne (123773) Harpactosphecion filicorne, Lytopsenella crastina (124318) Harpactosphecion filicornis (123757) Lytopsenella kerneggeri (138800) Misepyris minor, Homoglenus bifossatus (124205) Palaeobethyloides longiceps, Eupsenella crastina (127170) Palaeodryinus groehni, Harpactosphecion gracile, Pristapenesia primaeva, Dryinus janzeni, Bocchus primaevus, Lonchodryinus groehni, Primeuchroeus groehni (110735) Pristapenesia primaeva (123760 124275)
Priabonian38.0 - 33.9Denmark Holepyris sp., Sclerodermini indet., Pristocerinae indet., Alphadryinus balticus, Alphadryinus bruesi, Sierola hastata, Plastanoxus atrescens (123762)
Priabonian38.0 - 33.9Russian Federation Harpactosphecion gracile, Dryinus janzeni (123419) Palaeobethylus longicollis, Lytopsenella crastina, Epyrinae indet. (173264) Pristapenesia primaeva (137410)
Priabonian38.0 - 33.9United Kingdom (England) Mesitius rectinervis (123959)
Priabonian38.0 - 33.9Ukraine Ampulicomorpha succinalis, Dryinus reifi, Dryinus wunderlichi, Palaeobethylus longicollis, Palaeobethylus politus (207953) Bocchus rex (223355 227091) Bocchus schmalhauseni, Voronknoxus zerovae, Rovnodryinus khomychi (216854) Dryinus janzeni (196704) Palaeoanteon janzeni (185684) Palaeobethyloides sp., Palaeobethylus sp., Ampulicomorpha succinalis, Palaeobethylus politus, Laelius rovnensis, Laelius preteritus, Lytopsenella kerneggeri, Eupsenella aulax, Eupsenella klesoviana, Sierola rovniana, Foveorisus kilimniki, Chlorepyris deploegi, Chlorepyris jouaulti, Chlorepyris mckellari, Holepyris terayamai, Pseudisobrachium megalosaurus, Pseudisobrachium stegosaurus (122997) Pristapenesia primaeva, Chlorepyris neli, Cleistepyris allosaurus (113621)
Priabonian38.0 - 33.9Poland Protadelphe aenea, Palaeochrum diversum, Laberites polonicus (120813)
Chadronian37.2 - 33.9USA (Colorado) Bethylidae indet. (122976) Chrysis miocenica, Chrysis rohweri (106381) Epyris deletus (123119)
Early/Lower Oligocene33.9 - 28.4France Chrysis amoena (121473)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Epyris tenellus (122010)
Chattian28.1 - 23.03Russian Federation Dryinidae indet. (125971)
Early/Lower Miocene23.03 - 15.97Mexico (Chiapas) Dryinus palaeomexicanus (124928) Goniozus cotyi, Bakeriella nanyhelae, Solepyris electromexicanus (225741)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Alongatepyris pedrocai (229461) Archaeopristocera miki (151097) Bocchus vetustus (159268) Ceratochrysis dominicana, Dryinus grimaldii, Dryinus hymenaeaphilus, Dryinus priscus, Dryinus pristinus, Dryinus vetustus, Pseudodryinus sucinus, Clystopsenella mirabilis, Dryinidae indet. (122982) Dominibythus inopinatus (122473) Dryinus palaeodominicanus (154719 166828) Dryinus palaeodominicanus, Alphadryinus sucinus (157305) Dryinus palaeodominicanus, Harpactosphecion scheveni (124846) Dryinus rasnitsyni, Harpactosphecion sucinum, Thaumatodryinus miocenicus, Dryinus alamellatus (122474) Dryinus sp., Dryinus grimaldii, Aphelopus poinari, Dryinus palaeodominicanus, Dryinus poinari, Harpactosphecion sucinum, Thaumatodryinus miocenicus, Probethylus poinari, Pterosclerogibba antiqua (122472) Goniozus respectus, Epyrinae indet., Anisepyris gradatus (146557) Thaumatodryinus fuscescens, Crassibethylus dominicanus (203120) Thaumatodryinus miocenicus (155866 155867)
Langhian15.97 - 13.82China (Fujian) Sierola colomboi, Hedychridium rosai, Parascleroderma palaeosinica (220273)
Holocene0.0117 - 0.0Tanzania Bethylidae indet. (234540) Calyoza ramosa (140788) Calyoza staphylinoides (165051) Chrysis viridicyanea (129861)
Holocene0.0117 - 0.0Madagascar Messoria copalina (151372)