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Raptipedia

Insecta - Mecoptera

Synonymy list
YearName and author
1987Raptipedia Willmann
1996Raptipedia Ansorge p. 73
2004Raptipedia Petrulevičius and Jarzembowski

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaLatreille 1829
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
RankNameAuthor
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Panorpida
superorderAntliophora
orderMecopteraPackard 1886
suborderPistillifera
infraorderRaptipedia
infraorderRaptipedia

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Infraor. Raptipedia Willmann 1987
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Fm. Bittacidae Handlirsch 1906 [hangingfly]
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G. †Antiquanabittacus Petrulevičius and Jarzembowski 2004
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†Antiquanabittacus dispar Bashkuev 2023
†Antiquanabittacus nanus Petrulevičius and Jarzembowski 2004
†Antiquanabittacus punctatus Yu et al. 2025
†Antiquanabittacus robustus Bashkuev 2023
†Antiquanabittacus saltator Bashkuev 2023
G. †Archebittacus Riek 1955
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†Archebittacus exilis Riek 1955
G. †Baissobittacus Novokshonov 1997
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†Baissobittacus angustipennatus Novokshonov 1997
Subfm. †Bittacinae Handlirsch 1906
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G. †Asiobittacus Novokshonov 1993
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†Asiobittacus sajsagulis Novokshonov 1993
G. †Mesobittacus Handlirsch 1939
+
+
Invalid names: Auxobittacus praeacutus Bode 1953 [synonym]
†Mesobittacus kubekovensis Novokshonov 1993
†Mesobittacus minutus Handlirsch 1939
+
Invalid names: Protobittacus pediculatus Bode 1953 [synonym]
Invalid names: Auxobittacus Bode 1953 [synonym], Microbittacus Novokshonov 1993 [synonym]
G. Orobittacus Villegas and Byers 1981
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†Orobittacus antiquus Novokshonov 1993
†Orobittacus minutus Novokshonov 1993
G. Bittacus Latreille 1802
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†Bittacus biamensis Novokshonov 1993
†Bittacus egestionis Carpenter 1955
†Bittacus lepiduscretaceus Li et al. 2018
†Bittacus succinus Carpenter 1954
†Bittacus veternus Cockerell 1921
G. †Burmobittacus Zhao et al. 2016
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†Burmobittacus jarzembowskii Zhao et al. 2016
G. †Composibittacus Liu et al. 2016
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†Composibittacus bipunctatus Liu et al. 2016
†Composibittacus reticulatus Liu et al. 2016
†Composibittacus retroflexus Yu et al. 2025
G. †Cretobittacus Novokshonov 1993
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†Cretobittacus baisensis Novokshonov 1993
G. †Decoribittacus Li and Ren 2009
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†Decoribittacus euneurus Li and Ren 2009
†Decoribittacus stictus Li and Ren 2009
G. Eremobittacus Byers 1997
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†Eremobittacus opeth Poschmann and Nel 2022
G. †Exilibittacus Yang et al. 2012
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†Exilibittacus foliaceus Liu et al. 2014
†Exilibittacus lii Yang et al. 2012
†Exilibittacus plagioneurus Liu et al. 2014
G. †Formosibittacus Li et al. 2008
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†Formosibittacus macularis Li et al. 2008
†Formosibittacus multifarius Yu et al. 2025
G. Hylobittacus Byers 1979
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†Hylobittacus antiquus Pictet and Hagen 1856
†Hylobittacus fossilis Carpenter 1954
†Hylobittacus minimus Carpenter 1954
†Hylobittacus picteti Krzemiński 2007
Invalid names: Electrobittacus Carpenter 1931 [synonym]
G. †Jurahylobittacus Li et al. 2008
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†Jurahylobittacus astictus Li et al. 2008
G. †Karattacus Novokshonov 1997
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†Karattacus longialatus Li and Ren 2009
†Karattacus persibus Novokshonov 1997
G. †Liaobittacus Ren 1993
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G. †Megabittacus Ren 1997
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†Megabittacus spatiosus Yang et al. 2012
G. †Mongolobittacus Petrulevičius et al. 2007
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†Mongolobittacus daohugouensis Petrulevičius et al. 2007
†Mongolobittacus oligophlebius Liu et al. 2014
†Mongolobittacus speciosus Liu et al. 2014
Subfm. †Neorthophlebiinae Handlirsch 1920
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G. †Ctenobittacus Villet 2025
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Invalid names: Ctenophlebia Hong 2009 [replaced]
G. †Haplobittacus Bode 1953
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†Haplobittacus parvus Bode 1953
G. †Neorthophlebia Handlirsch 1906
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†Neorthophlebia acutipennis Handlirsch 1939
†Neorthophlebia debilis Handlirsch 1906
†Neorthophlebia maculipennis Handlirsch 1906
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Invalid names: Metaxybittacus vittatus Bode 1953 [synonym], Neorthophlebia minor Handlirsch 1906 [synonym]
†Neorthophlebia medialis Handlirsch 1939
†Neorthophlebia nana Martynova 1948
†Neorthophlebia pallida Handlirsch 1939
†Neorthophlebia robusta Martynov 1937
†Neorthophlebia simillima Handlirsch 1939
†Neorthophlebia stigmatica Handlirsch 1939
†Neorthophlebia unica Martynova 1948
†Neorthophlebia yunnanensis Zhang and Hong 2003
G. †Neorthophlebopsis Hong et al. 2005
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†Neorthophlebopsis qishuiheensis Hong et al. 2005
G. †Parabittacus Handlirsch 1939
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†Parabittacus analis Handlirsch 1939
†Parabittacus lingula Bode 1953
Invalid names: Polydicrobittacus Bode 1953 [synonym]
G. †Pleobittacus Bode 1953
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G. †Protobittacus Tillyard 1933
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†Protobittacus handlirschi Tillyard 1933
†Protobittacus liassicus Tillyard 1933
†Protobittacus maculatus Tillyard 1933
†Protobittacus moestus Sukatsheva 1985
G. †Orthobittacus Willmann 1989
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†Orthobittacus abshiricus Martynova 1951
†Orthobittacus maculosus Liu et al. 2016
†Orthobittacus polymitus Novokshonov 1996
†Orthobittacus suni Kopeć et al. 2018
G. †Paekthobittacus Ri et al. 2023
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†Paekthobittacus sinuijuensis Ri et al. 2023
G. †Palaeobittacus Carpenter 1928
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†Palaeobittacus eocenicus Carpenter 1928
Subfm. †Plesiobittacinae Novokshonov 1997
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G. †Plesiobittacus Novokshonov 1997
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†Plesiobittacus martynovi Novokshonov 1997
†Plesiobittacus primigenius Novokshonov 1997
G. †Preanabittacus Novokshonov 1993
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†Preanabittacus karatavensis Novokshonov 1993
†Preanabittacus validus Yang et al. 2012
G. †Probittacus Martynov 1927
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†Probittacus avitus Martynov 1927
G. †Prohylobittacus Novokshonov 1993
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†Prohylobittacus sorosi Novokshonov 1993
G. †Scharabittacus Novokshonov 1993
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†Scharabittacus karatavicus Novokshonov 1993
†Scharabittacus mongolicus Novokshonov 1993
†Scharabittacus pictus Martynova 1949
G. †Sibirobittacus Sukacheva 1990
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†Sibirobittacus kazakhstanensis Novokshonov 1993
†Sibirobittacus undus Sukacheva 1990
G. Thyridates Navas 1908
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†Thyridates novokschonovi Petrulevičius 2003
G. Tytthobittacus Smithers 1973
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†Tytthobittacus jarzembowski Kopeć et al. 2016
Fm. †Cimbrophlebiidae Willmann 1977 [hangingfly]
+
G. †Bellicimbrophlebia Yang et al. 2013
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†Bellicimbrophlebia angusta Yang et al. 2013
†Bellicimbrophlebia cruciata Yang et al. 2013
†Bellicimbrophlebia disvena Yang et al. 2013
†Bellicimbrophlebia eumorpha Yang et al. 2013
†Bellicimbrophlebia heteroneura Zhang et al. 2015
G. †Cimbrophlebia Willmann 1977
+
†Cimbrophlebia amoena Zhang et al. 2015
†Cimbrophlebia bittaciformis Willmann 1977
†Cimbrophlebia brooksi Archibald 2009
†Cimbrophlebia flabelliformis Archibald 2009
†Cimbrophlebia gracilenta Zhang et al. 2015
†Cimbrophlebia leahyi Archibald 2009
†Cimbrophlebia rara Wang et al. 2014
†Cimbrophlebia westae Archibald 2009
G. †Juracimbrophlebia Wang et al. 2012
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†Juracimbrophlebia ginkgofolia Wang et al. 2012
G. †Malmocimbrophlebia Bechly and Schweigart 2000
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†Malmocimbrophlebia buergeri Bechly and Schweigart 2000
G. †Mirorcimbrophlebia Yang et al. 2013
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G. †Perfecticimbrophlebia Yang et al. 2012
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†Perfecticimbrophlebia laetus Yang et al. 2012
G. †Telobittacus Zhang 1993
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†Telobittacus bellus Yang et al. 2013
†Telobittacus decorus Zhang et al. 2015
†Telobittacus fragosus Zhang 1993
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Longobardian to the top of the MP 28 or 239.48000 to 23.04000 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma

Collections (73 total)


Time interval Ma Country or state Original ID and collection number
Longobardian239.48 - 237.0China (Shaanxi) Ctenophlebia tongchuanensis, Neorthophlebopsis qishuiheensis (117138)
Norian227.3 - 205.7Australia (Queensland) Archebittacus exilis (106484 191411)
Rhaetian205.7 - 201.4United Kingdom (England) Protobittacus liassicus, Protobittacus maculatus (113961)
Hettangian - Sinemurian201.4 - 192.9Kyrgyzstan (Tonskiy) Neorthophlebia robusta, Neorthophlebia unica, Neorthophlebia nana (105087)
Pliensbachian192.9 - 184.2Kyrgyzstan Neorthophlebia abshirica (160469)
Early/Lower Toarcian184.2 - 182.9Germany (Mecklenburg-Vorpommern) Bittacidae indet., Cimbrophlebiidae indet., Neorthophlebia maculipennis, Mesobittacus minutus (114993)
Early/Lower Toarcian184.2 - 182.9Germany Neorthophlebia minor, Mesobittacus minutus, Orthophlebia megapolitana, Neorthophlebia debilis, Parabittacus analis, Neorthophlebia pallida, Neorthophlebia simillima, Neorthophlebia medialis, Neorthophlebia stigmatica, Neorthophlebia acutipennis, Orthophlebia megapolitana (123987)
Early/Lower Toarcian184.2 - 182.9Germany (Niedersachsen) Auxobittacus marginelaesus (107322) Auxobittacus praeacutus (107343) Polydicrobittacus lingula (107553) Polydicrobittacus lingula, Haplobittacus parvus (107321) Protobittacus arculatus, Protobittacus pediculatus, Auxobittacus praeacutus, Metaxybittacus vittatus, Pleobittacus retroflexus, Polydicrobittacus lingula (105002) Protobittacus clavaeformis (107323) Protobittacus desacuminatus (107555)
Falciferum184.2 - 182.9Germany Bittacidae indet. (192037)
Falciferum184.2 - 182.9United Kingdom (England) Bittacidae indet. (234543) Protobittacus handlirschi (154307)
Toarcian184.2 - 174.7Kyrgyzstan Neorthophlebia robusta (114702)
Toarcian184.2 - 174.7Russian Federation (Kemerovo) Protobittacus moestus (125484)
Toarcian184.2 - 174.7Kyrgyzstan (Batken) Asiobittacus sajsagulis, Plesiobittacus martynovi, Plesiobittacus primigenius (105003)
Bajocian - Bathonian170.9 - 165.3Russian Federation (Krasnoyarsk) Microbittacus kubekovensis (118010)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Bellicimbrophlebia cruciata, Bellicimbrophlebia disvena (117610) Exilibittacus lii (137887) Formosibittacus macularis, Jurahylobittacus astictus, Decoribittacus euneurus, Decoribittacus stictus, Karattacus longialatus (112664) Formosibittacus multifarius, Antiquanabittacus punctatus, Composibittacus retroflexus (112663) Mongolobittacus daohugouensis (113371) Mongolobittacus speciosus, Mongolobittacus oligophlebius, Exilibittacus foliaceus, Exilibittacus plagioneurus (155751) Orthobittacus suni (183155) Perfecticimbrophlebia laetus, Preanabittacus validus, Juracimbrophlebia ginkgofolia, Telobittacus bellus, Mirorcimbrophlebia daohugouensis, Bellicimbrophlebia cruciata, Bellicimbrophlebia angusta, Bellicimbrophlebia eumorpha, Bellicimbrophlebia disvena (113366) Telobittacus decorus, Bellicimbrophlebia heteroneura, Cimbrophlebia amoena, Cimbrophlebia gracilenta, Composibittacus bipunctatus, Composibittacus reticulatus, Orthobittacus maculosus (123761)
Callovian - Oxfordian165.3 - 154.8China (Liaoning) Liaobittacus longantennatus (107004)
Callovian - Oxfordian165.3 - 154.8Kazakhstan (Chimkent) Karattacus persibus, Orobittacus antiquus (105971) Orthobittacus polymitus, Probittacus karatavicus (105426) Preanabittacus karatavensis, Probittacus karatavicus (106101) Probittacus avitus (114692) Prohylobittacus sorosi (113439) Prohylobittacus sorosi, Orobittacus antiquus (105970) Sibirobittacus kazakhstanensis (106102)
Oxfordian - Kimmeridgian161.5 - 149.2China (Yunnan) Neorthophlebia yunnanensis (164859)
Late/Upper Jurassic161.5 - 143.1Russian Federation (Buryatia) Neorthophlebia indet. (125483)
Early/Lower Tithonian149.2 - 145.06Germany Malmocimbrophlebia buergeri (122880)
Tithonian149.2 - 143.1Mongolia Orobittacus minutus (104748) Probittacus mongolicus (121393)
Tithonian149.2 - 143.1Russian Federation (Chita) Sibirobittacus undus (123197)
Early/Lower Berriasian143.1 - 141.0United Kingdom (England) Tytthobittacus jarzembowski (104007)
Late/Upper Hauterivian132.6 - 125.77United Kingdom (England) Antiquanabittacus nanus (106417)
Late/Upper Hauterivian - Early/Lower Barremian132.6 - 121.4China (Shaanxi) Telobittacus fragosus (140940)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Megabittacus colosseus, Megabittacus beipiaoensis, Sibirobittacus atalus (138436) Megabittacus spatiosus (140491)
Late/Upper Barremian125.77 - 121.4China (Nei Mongol) Cimbrophlebia rara (125656)
Barremian - Aptian125.77 - 113.2North Korea Paekthobittacus sinuijuensis (200272)
Aptian121.4 - 113.2Russian Federation (Buryatia) Antiquanabittacus saltator, Antiquanabittacus dispar (113349) Baissobittacus angustipennatus, Cretobittacus baisensis (105969) Raptipedia indet., Antiquanabittacus sp., Antiquanabittacus saltator, Antiquanabittacus robustus, Antiquanabittacus dispar (113350)
Early/Lower Cenomanian100.5 - 93.9Myanmar Bittacus lepiduscretaceus (178121) Burmobittacus jarzembowskii (161726)
Late/Upper Paleocene59.24 - 56.0Argentina (Jujuy) Thyridates novokschonovi (113004)
Ypresian56.0 - 48.07USA (Washington) Cimbrophlebia brooksi, Cimbrophlebia westae (103206)
Ypresian56.0 - 48.07Canada (British Columbia) Cimbrophlebia sp. A, Cimbrophlebia flabelliformis, Cimbrophlebia leahyi (103205)
Ypresian56.0 - 48.07Denmark Cimbrophlebia bittaciformis (113025)
Bridgerian50.5 - 46.2USA (Colorado) Palaeobittacus eocenicus (163519)
Bridgerian50.5 - 46.2USA (Utah) Bittacus egestionis (122882)
Priabonian37.71 - 33.9Russian Federation Hylobittacus fossilis, Hylobittacus minimus, Hylobittacus picteti (123419)
Priabonian37.71 - 33.9Poland Bittacus antiquus (109625) Hylobittacus antiquus, Hylobittacus fossilis, Hylobittacus minimus, Hylobittacus picteti (123215) Hylobittacus fossilis (120813)
Priabonian37.71 - 33.9Germany (Halle) Hylobittacus picteti (124273)
Priabonian37.71 - 33.9Russian Federation (Primorskiy) Bittacus biamensis (123794)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Bittacus fossilis, Bittacus minimus, Bittacus succinus, Hylobittacus antiquus (124205) Bittacus minimus (123766)
Priabonian37.71 - 33.9United Kingdom (England) Bittacus sp.A, Panorpa veterna, Bittacus veternus (123960)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Eremobittacus opeth (160097)