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Culicoidini

Insecta - Diptera - Ceratopogonidae

Synonymy list
YearName and author
1911Culicoidinae Kieffer p. 1
1911Culicoidini Kieffer p. 1
1944Culicoidinae Statz p. 142
1988Culicoidini Szadziewski p. 27
1996Culicoidini Szadziewski p. 58
1997Culicoidini Borkent and Wirth p. 59
2011Culicoidini Sontag and Szadziewski p. 783
2013Culicoidini Borkent p. 72
2014Culicoidini Choufani et al. p. 38
2020Culicoidini Borkent and Dominiak p. 102
2021Culicoidini Peñalver et al. p. 427

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
RankNameAuthor
classInsecta
orderDiptera
infraorderCulicomorpha
superfamilyChironomoideaMacquart 1838
familyCeratopogonidae
subfamilyCeratopogoninae
tribeCulicoidini
tribeCulicoidini

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Tr. Culicoidini Kieffer 1911
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G. Culicoides Latreille 1809
+
Subg. Culicoides (Groganomyia) Szadziewski and Dominiak 2019
+
†Culicoides (Groganomyia) bojarskii Szadziewski and Dominiak 2019
†Culicoides (Groganomyia) burmiticus Szadziewski and Dominiak 2019
†Culicoides (Groganomyia) ellenbergeri Szadziewski and Dominiak 2019
†Culicoides (Groganomyia) myanmaricus Szadziewski and Dominiak 2019
Subg. Culicoides (Oecacta) Poey 1853
+
†Culicoides agamus Borkent 1995
†Culicoides ambericus Szadziewski and Grogan 1998
†Culicoides annosus Borkent 1995
†Culicoides antilleanus Szadziewski and Grogan 1998
†Culicoides atratus Statz 1944
†Culicoides austerus Statz 1944
†Culicoides balticus Szadziewski 1988
†Culicoides bicolor Statz 1944
†Culicoides bifidus Borkent 1996
†Culicoides brisaci Choufani and Nel 2013
†Culicoides brodzinskyi Szadziewski and Grogan 1998
†Culicoides bullus Borkent 1995
†Culicoides canadensis Boesel 1937
†Culicoides casei Grogan and Szadziewski 1988 [biting midge]
†Culicoides ceranowiczi Szadziewski 1988
†Culicoides dasyheleiformis Szadziewski 1988
†Culicoides doyeni Choufani et al. 2014
†Culicoides elongatulus Statz 1944
†Culicoides eoselficus Szadziewski 1988
†Culicoides filipalpis Remm 1976
†Culicoides gedanensis Szadziewski 1988
†Culicoides gracilior Statz 1944
†Culicoides grandibocus Borkent 1996
†Culicoides hispanicola Szadziewski and Grogan 1998
†Culicoides jucundus Statz 1944
†Culicoides kaluginae Remm 1976
†Culicoides liliputanus Statz 1944
†Culicoides mammalicola Szadziewski and Grogan 1998
†Culicoides megacanthus Palmer 1957
†Culicoides obesus Statz 1944
†Culicoides obscuratus Statz 1944
†Culicoides obuncus Borkent 1995
†Culicoides paleopestis Peñalver et al. 2021
†Culicoides prussicus Szadziewski 1988
†Culicoides speciosus Meunier 1904
†Culicoides subgedanensis Szadziewski 1993
†Culicoides succineus Remm 1976
†Culicoides succivarius Szadziewski 1988
†Culicoides tenuipennis Statz 1944
†Culicoides truncatus Borkent 2000
†Culicoides tyrrelli Boesel 1937
†Culicoides ventralis Borkent and Wirth 1997
+
Invalid names: Culicoides abdominalis Statz 1944 [replaced]
†Culicoides yoosti Borkent 2000
Invalid names: Devalquia Choufani and Nel 2013 [synonym]
G. Washingtonhelea Wirth and Grogan 1988
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Hemingfordian or 93.90000 to 15.97000 Ma
Minimum age of oldest fossil (stem group age): 93.5 Ma

Collections (44 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Cenomanian99.6 - 93.5Myanmar Culicoides bojarskii, Culicoides myanmaricus (161726) Culicoides burmiticus (187929) Culicoides ellenbergeri (165560)
Late/Upper Cenomanian99.6 - 93.5USA (New Jersey) Culicoides casei (110959)
Turonian93.9 - 89.8USA (New Jersey) Culicoides bifidus (123016) Culicoides casei, Culicoides filipalpis, Culicoides grandibocus, Culicoides truncatus, Culicoides yoosti (110895)
Turonian93.9 - 89.8France Culicoides doyeni (164386)
Early/Lower Santonian85.8 - 83.5France Devalquia brisaci (152462)
Late/Upper Santonian85.8 - 83.5Russian Federation Culicoides sp., Culicoides filipalpis, Culicoides kaluginae, Culicoides sphenostylus, Culicoides succineus (35056)
Late/Upper Campanian83.5 - 70.6Canada (Manitoba) Culicoides filipalpis, Culicoides canadensis, Culicoides obuncus (122353) Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis, Culicoides agamus, Culicoides obuncus, Culicoides annosus (122354) Culicoides filipalpis, Dasyhelea tyrrelli, Atrichopogon canadensis, Culicoides obuncus (122356)
Middle Campanian83.5 - 70.6Canada (Alberta) Culicoides filipalpis (122385) Culicoides filipalpis, Culicoides bullus (123143) Culicoides filipalpis, Culicoides tyrrelli, Culicoides annosus (104932) Culicoides filipalpis, Culicoides tyrrelli, Culicoides bullus (122388) Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis (122383) Culicoides filipalpis, Culicoides tyrrelli, Culicoides canadensis, Culicoides obuncus (122384) Culicoides sp. (34896) Culicoides sp., Culicoides filipalpis (122389)
Bartonian41.3 - 38.0Australia (Victoria) Culicoides paleopestis (224365)
Priabonian38.0 - 33.9Denmark Culicoides sp., Culicoides speciosus, Culicoides sp. B, Culicoides eoselficus, Culicoides balticus (123762)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Ceratopogon speciosus (123927) Culicoides sp. (123962 123993) Culicoides sp., Culicoides ceranowiczi, Culicoides dasyheleiformis (123221)
Priabonian38.0 - 33.9Ukraine Culicoides sp. (122995) Culicoides sp., Culicoides speciosus (113621 122997)
Priabonian38.0 - 33.9Poland Culicoides prussicus, Culicoides dasyheleiformis (109625) Culicoides sp., Culicoides sp. A, Culicoides sp. B, Culicoides sp. C, Culicoides ceranowiczi, Culicoides eoselficus, Culicoides gedanensis, Culicoides speciosus, Culicoides dasyheleiformis, Culicoides succivarius, Culicoides balticus (120813) Culicoides sp., Culicoides speciosus (154301)
Priabonian38.0 - 33.9Germany (Halle) Culicoides sp., Culicoides speciosus (129228) Culicoides sp., Culicoides speciosus, Culicoides ceranowiczi, Culicoides subgedanensis (129227)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Culicoides abbreviatipennis, Culicoides atratus, Culicoides austerus, Culicoides bicolor, Culicoides elongatulus, Culicoides gracilior, Culicoides jucundus, Culicoides liliputanus, Culicoides obesus, Culicoides tenuipennis, Culicoides abdominalis, Culicoides obscuratus (122010)
Early/Lower Miocene23.03 - 15.97Mexico Culicoides sp. (21977)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Culicoides antilleanus, Culicoides ambericus, Culicoides mammalicolus, Culicoides sp. (122473) Culicoides brodzinskyi, Culicoides ambericus, Culicoides sp. (122982) Culicoides hispanicolus, Culicoides sp. (123857)
Hemingfordian20.43 - 15.97USA (California) Culicoides megacanthus (106431 123022 123030 123034 152222)