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Archostemata

Insecta - Coleoptera

Synonyms
Synonymy list
YearName and author
1908Archostemata Kolbe
1966Notocupedini Ponomarenko
1967Notocupedini Ponomarenko
1968Moravocoleus perditus Kukalová p. 146 figs. 5; Pl IV fig 3
1969Avocoleus perditus Ponomarenko p. 52
1981Archostemata Crowson p. 694
1981Cupediformia Crowson p. 694
1986Priacma corrupta Ponomarenko p. 85 figs. 51; Pl VI fig 2
1992Archostemata Carpenter
1997Priacma longicapitis Ponomarenko p. 397 figs. Pl 6, fig 15; text-fig 12
1997Priacma oculata Ponomarenko p. 397 figs. Pl 6, fig 14; text-fig 11
2000Priacma striata Ponomarenko p. S321 figs. Pl 8, figs 8,9; text-fig 5
2006Notocupedini Soriano and Delclos
2009Notocupedini Tan and Ren p. 74
2011Archostemata Bouchard et al. p. 95
2011Notocupedini Bouchard et al. p. 96
2012Notocupedini Ponomarenko et al. p. 476
2012Notocupedini Tan et al.
2013Moravocoleus perditus Kirejtshuk and Nel fig. 6
2014Notocupedini Ponomarenko p. 1658
2015Notocupedini Jarzembowski et al. p. 432
2019Notocupedini Strelnikova p. 295
2019Notocupedini Tihelka et al. p. 571
2020Notocupedini Jiang et al.
2020Archostemata Kirejtshuk p. 9
2020Cupediformia Kirejtshuk p. 9
2021Notocupedini Strelnikova and Yan p. 501
2022Archostemata Boudinot et al.
2022Archostemata Cai et al.
2023Notocupedini Li et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
RankNameAuthor
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderArchostemataKolbe 1908
suborderArchostemataKolbe 1908

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subor. Archostemata Kolbe 1908
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G. †Anthocoleus Hong 1983
+
G. †Argentinocupes Martins-Neto and Gallego 2006
+
†Argentinocupes abdalai Martins-Neto and Gallego 2006
†Argentinocupes pulcher Martins-Neto and Gallego 2006
†Argentinocupes sara Martins-Neto and Gallego 2009
Superfm. †Asiocoleoidea Rohdendorf 1961
+
Fm. †Tricoleidae Ponomarenko 1969
+
G. †Sogdelytron Ponomarenko 1969
+
†Sogdelytron latum Ponomarenko 1969
Invalid names: Longxianocupes Hong 1985 [synonym]
G. †Triassocupes Hong 1985
+
G. †Tricoleodes Ponomarenko 1969
+
†Tricoleodes acutus Ponomarenko 1969
†Tricoleodes longus Ponomarenko 1969
G. †Tricoleus Ponomarenko 1969
+
†Tricoleus incertus Ponomarenko 1969
†Tricoleus punctatus Ponomarenko 1969
G. †Willcoxia Dunstan 1923
+
†Willcoxia magnopunctata Dunstan 1923
G. †Bothynophora Heer 1864
+
†Bothynophora elegans Heer 1864
G. †Brachilatus Strelnikova and Yan 2023
+
†Brachilatus caducus Ponomarenko 1969
†Brachilatus dundulaensis Ponomarenko 1994
†Brachilatus foersteri Ponomarenko 1971
+
Invalid names: Notocupes reticulatus Ponomarenko 1968 [replaced]
†Brachilatus longicoxa Soriano and Delclos 2006
†Brachilatus nigrimonticola Ponomarenko 1968
†Brachilatus oculatus Soriano and Delclos 2006
†Brachilatus premeris Lee et al. 2022
†Brachilatus spinosus Li and Cai 2023
†Brachilatus viridis Soriano and Delclos 2006
G. †Celocoleus Hong 1983
+
†Celocoleus densus Hong 1983
G. †Clathrocupes Hong 1980
+
G. †Conexicoxa Lin 1986
+
†Conexicoxa brachycephala Ponomarenko 1994
†Conexicoxa crassa Ponomarenko 1969
†Conexicoxa denticollis Jiang et al. 2020
†Conexicoxa epicharis Tan et al. 2005
†Conexicoxa eumeura Tan et al. 2006
†Conexicoxa exigua Ponomarenko 1994
†Conexicoxa homora Lin 1986
†Conexicoxa jurassica Hong 1983
†Conexicoxa khetanensis Ponomarenko 1993
†Conexicoxa kirghizica Ponomarenko 1969
†Conexicoxa longicollis Ponomarenko 1994
†Conexicoxa martinclosas Soriano and Delclos 2006
†Conexicoxa porrecta Tan et al. 2006
†Conexicoxa psilata Tan et al. 2005
†Conexicoxa siniestri Soriano and Delclos 2006
†Conexicoxa stabilis Tan et al. 2005
†Conexicoxa tripartita Oppenheim 1888
G. †Cupedites Ponomarenko 1985
+
†Cupedites minutissimus Ponomarenko 1985
Superfm. Cupedoidea Laporte 1836
+
Fm. Cupedidae Laporte 1836 [reticulated beetle]
+
G. †Cainocups Kirejtshuk et al. 2016
+
†Cainocups aixensis Kirejtshuk et al. 2016
Subfm. Cupedinae Lacordaire 1857
+
G. †Anaglyphites Ponomarenko 1964
+
†Anaglyphites admotus Ponomarenko 1964
†Anaglyphites capitatus Ponomarenko 1966
†Anaglyphites clavatus Ponomarenko 1964
†Anaglyphites crassipygus Ponomarenko 2014
†Anaglyphites minimus Ponomarenko 1997
†Anaglyphites minutissimus Ponomarenko 2014
†Anaglyphites mongolicus Ponomarenko 1997
G. †Apriacma Kirejtshuk et al. 2016
+
†Apriacma clavata Tan et al. 2006
†Apriacma latidentata Tan et al. 2006
†Apriacma renaria Tan et al. 2006
†Apriacma tuberculosa Tan et al. 2006
G. †Asimma Ponomarenko 1966
+
†Asimma rara Ponomarenko 1966
G. †Barbaticupes Jarzembowski et al. 2017
+
†Barbaticupes combertiae Jarzembowski et al. 2017
G. †Cainomerga Kirejtshuk et al. 2016
+
†Cainomerga brevicornis Kirejtshuk et al. 2016
†Cainomerga fraterna Kirejtshuk et al. 2016
†Cainomerga immaculata Piton 1940
†Cainomerga palaeocenica Kirejtshuk et al. 2016
†Cainomerga ponti Kirejtshuk et al. 2016
G. †Cretomerga Kirejtshuk et al. 2016
+
†Cretomerga subtilis Tan and Ren 2006
G. Cupes Fabricius 1801
+
†Cupes balticus Yamamoto 2024
†Cupes distinctissimus Kirejtshuk et al. 2016
†Cupes eckfeldensis Tröster 1993
†Cupes golovatchi Kirejtshuk 2020
†Cupes groehni Kirejtshuk 2005
†Cupes hoffeinsorum Kirejtshuk 2005
†Cupes kerneggeri Kirejtshuk 2005
†Cupes komissari Kirejtshuk 2005
†Cupes legalovi Kirejtshuk 2020
†Cupes longus Hong and Wang 1987
†Cupes lutzi Kirejtshuk 2020
†Cupes manifestus Kirejtshuk et al. 2010
†Cupes messelensis Tröster 1993
†Cupes nabozhenkoi Kirejtshuk 2020
†Cupes ponomarenkoi Kirejtshuk et al. 2010
†Cupes praeglacialis Gersdorf 1976
†Cupes rohdendorfi Yablokov-Khnzorian 1960
†Cupes simillimus Kirejtshuk et al. 2016
†Cupes tesselatus Motschulsky 1856
†Cupes tessellatus Motschulsky 1856
†Cupes wedmannae Kirejtshuk 2020
†Cupes weitschati Kirejtshuk 2005
Invalid names: Cupoides Motschulsky 1856 [synonym], Distocupes Neboiss 1984 [synonym], Tenomerga Neboiss 1984 [synonym]
G. †Cupidium Ponomarenko 1968
+
†Cupidium abavum Ponomarenko 1968
G. †Cupopsis Kirejtshuk et al. 2016
+
†Cupopsis svitkoi Lubkin 2003
G. †Ensicupes Hong 1976
+
†Ensicupes guyangensis Hong 1976
†Ensicupes obtusus Zhang 1997
G. †Furcicupes Tan and Ren 2006
+
†Furcicupes raucus Tan and Ren 2006
G. †Gracilicupes Tan et al. 2006
+
†Gracilicupes crassicruralis Tan et al. 2006
†Gracilicupes minimus Strelnikova 2019
†Gracilicupes tenuicruralis Tan et al. 2006
G. †Kirghizocupes Ponomarenko 1966
+
†Kirghizocupes cellulosus Ponomarenko 1966
†Kirghizocupes indistinctus Ponomarenko 1969
†Kirghizocupes proporeius Ponomarenko 1969
Invalid names: Mesocupoides Ponomarenko 1969 [synonym]
G. †Latocupes Ren and Tan 2006
+
†Latocupes angustilabialis Tan et al. 2007
†Latocupes bellus Ren and Tan 2006
†Latocupes collaris Tan et al. 2007
†Latocupes fortis Ren and Tan 2006
†Latocupes jiensis Ren 1995
†Latocupes latilabialis Tan et al. 2007
Invalid names: Pulchicupes Ren 1995 [synonym]
G. †Mallecupes Jarzembowski et al. 2017
+
Subg. †Mallecupes (Mallecupes) Jarzembowski et al. 2017
Subg. †Mallecupes (Mallecupoides) Jarzembowski et al. 2017
†Mallecupes cheni Wang et al. 2023
†Mallecupes prokini Kirejtshuk 2020
†Mallecupes zhangi Song et al. 2022
G. †Menatops Kirejtshuk et al. 2016
+
†Menatops bartenevi Kirejtshuk et al. 2016
†Menatops orbiculatus Kirejtshuk et al. 2010
G. †Mesocupes Martynov 1926
+
†Mesocupes bidens Ponomarenko 1964
†Mesocupes minor Ponomarenko 1968
†Mesocupes paulus Ponomarenko 1964
†Mesocupes primitivus Martynov 1926
†Mesocupes spinosus Ponomarenko 1964
†Mesocupes zherikhini Soriano and Delclos 2006
Invalid names: Mesocupes (Mesocupes) Martynov 1926 [invalid subgroup]
G. †Miocupes Ponomarenko 1973
+
†Miocupes rihai Ponomarenko 1973
G. †Priacmopsis Ponomarenko 1966
+
†Priacmopsis adumbrata Ponomarenko 1966
G. †Pterocupes Ponomarenko 1966
+
†Pterocupes antennatus Ponomarenko 1966
†Pterocupes leptocerus Ponomarenko 1966
G. †Taxopsis Kirejtshuk et al. 2016
+
†Taxopsis motschulskyi Kirejtshuk 2005
Invalid names: Anaglyphites pluricavus Soriano and Delclos 2006 [nomen dubium], Cupedini Crowson 1962 [empty], Platycupes sogdianus Ponomarenko 1966 [nomen dubium], Priacma sanzii Soriano and Delclos 2006 [nomen dubium]
Subfm. Priacminae Crowson 1962
+
G. Paracupes Kolbe 1898
G. Priacma Leconte 1861
+
†Priacma megapuncta Li and Cai 2019
Invalid names: Mesocupedinae Ponomarenko 1969 [empty]
G. †Magnocoleus Hong 1998
+
Fm. Micromalthidae Barber 1913 [telephone pole beetle]
+
G. †Archaeomalthus Yan et al. 2020
+
†Archaeomalthus synoriakos Yan et al. 2020
G. †Cretomalthus Kirejtshuk and Azar 2008
+
†Cretomalthus acracrowsonorum Kirejtshuk and Azar 2008
G. Micromalthus LeConte 1878
+
Micromalthus debilis LeConte 1878
+
Invalid names: Micromalthus anansi Perkovsky 2007 [synonym]
†Micromalthus eocenicus Kirejtshuk et al. 2010
†Micromalthus priabonicus Perkovsky 2016
G. †Protomalthus Tihelka et al. 2020
+
†Protomalthus burmaticus Tihelka et al. 2020
Fm. Ommatidae Sharp and Muir 1912
+
G. †Asiania Lee et al. 2023
+
†Asiania pax Lee et al. 2023
G. †Clessidromma Jarzembowski et al. 2017
+
†Clessidromma palmeri Jarzembowski et al. 2017
G. †Kirejtomma Li and Cai 2021
+
†Kirejtomma zengi Kirejtshuk 2020
G. †Limnomma Li and Cai 2021
+
†Limnomma daohugouense Li and Cai 2021
G. †Miniomma Li et al. 2020
+
†Miniomma chenkuni Li et al. 2020
Subfm. Ommatinae Sharp and Muir 1912
+
G. †Allophalerus Kirejtshuk 2020
+
†Allophalerus antiquus Ponomarenko 1964
†Allophalerus aphaleratus Ponomarenko 1969
†Allophalerus bontsaganensis Ponomarenko 1997
†Allophalerus incertus Ponomarenko 1969
†Allophalerus latus Tan et al. 2007
†Allophalerus maximus Ponomarenko 1968
†Allophalerus okhotensis Ponomarenko 1993
†Allophalerus tenuipes Ponomarenko 1964
†Allophalerus verrucosus Ponomarenko 1966
G. †Blapsium Westwood 1854
+
†Blapsium egertoni Westwood 1854
Invalid names: Blapsidium Phillips 1871 [synonym]
G. †Brochocoleus Hong 1982
+
†Brochocoleus alatus Ponomarenko 1994
†Brochocoleus cossyphus Ponomarenko 1994
†Brochocoleus keenani Jarzembowski et al. 2013
†Brochocoleus maculatus Whalley 1985
†Brochocoleus maximus Jarzembowski et al. 2013
†Brochocoleus planus Ponomarenko 1994
†Brochocoleus rostratus Ponomarenko 1999
†Brochocoleus sacheonensis Lee et al. 2023
†Brochocoleus tobini Jarzembowski et al. 2013
G. †Burmocoleus Kirejtshuk 2020
+
†Burmocoleus prisnyi Kirejtshuk 2020
†Burmocoleus zhiyuani Liu et al. 2017
G. †Cionocoleus Ren 1995
+
†Cionocoleus cervicalis Tan et al. 2007
†Cionocoleus elizabethae Jarzembowski et al. 2013
†Cionocoleus jepsoni Jarzembowski et al. 2013
†Cionocoleus longicapitis Soriano and Delclos 2006
†Cionocoleus magicus Ren 1995
†Cionocoleus minimus Jarzembowski et al. 2013
†Cionocoleus olympicus Jarzembowski et al. 2013
†Cionocoleus ommamimus Ponomarenko 1997
†Cionocoleus planiusculus Tan et al. 2007
†Cionocoleus punctatus Martynov 1926
†Cionocoleus sibiricus Ponomarenko 2000
†Cionocoleus tanae Jarzembowski et al. 2013
†Cionocoleus watsoni Jarzembowski et al. 2013
Tr. †Clessidrommatini Jarzembowski et al. 2017
+
G. †Lepidomma Jarzembowski et al. 2019
G. †Diluticupes Ren 1995
+
†Diluticupes applanatus Tan and Ren 2009
†Diluticupes crowsonae Jarzembowski et al. 2013
†Diluticupes magnus Tan and Ren 2009
†Diluticupes minor Ponomarenko 2000
†Diluticupes validus Tan and Ren 2009
†Diluticupes yangshuwanziensis Jarzembowski et al. 2013
G. †Echinocups Kirejtshuk and Jarzembowski 2020
G. †Eurydictyon Ponomarenko 1969
+
†Eurydictyon conspicuum Ponomarenko 1969
G. †Jarzembowskiops Kirejtshuk 2020
+
†Jarzembowskiops caseyi Jarzembowski et al. 2016
G. †Liassocupes Zeuner 1962
+
†Liassocupes parvus Zeuner 1962
G. †Lithocupes Ponomarenko 1966
+
†Lithocupes gigas Ponomarenko 1969
†Lithocupes incertus Ponomarenko 1966
†Lithocupes punctatus Ponomarenko 1969
G. †Monticupes Ren 1995
+
†Monticupes curtinervis Tan et al. 2007
†Monticupes decorosus Tan et al. 2012
G. †Notocupoides Ponomarenko 1966
+
†Notocupoides capitatus Ponomarenko 1966
†Notocupoides fasciatus Ponomarenko 1966
†Notocupoides triassicus Ponomarenko 1966
G. †Odontomma Ren et al. 2006
+
†Odontomma patula Ponomarenko 1985
†Odontomma sulcatum Tan et al. 2007
†Odontomma trachylaenum Ren et al. 2006
Tr. Ommatini Sharp and Muir 1912
+
G. Omma Newman 1839
+
Invalid names: Chalepocarabus Handlirsch 1906 [synonym], Cionocups Kirejtshuk 2020 [synonym], Ommomima Ponomarenko 1964 [synonym], Procarabus Oppenheim 1888 [synonym], Pyrochroophana Handlirsch 1906 [synonym]
G. †Paraodontomma Yamamoto 2017
+
†Paraodontomma burmitica Yamamoto 2017
†Paraodontomma leptocristatum Li et al. 2021
†Paraodontomma szwedoi Jarzembowski et al. 2018
G. †Pareuryomma Tan et al. 2012
+
†Pareuryomma angustum Tan et al. 2007
†Pareuryomma cardiobasis Tan et al. 2012
†Pareuryomma tylodes Tan et al. 2006
Invalid names: Euryomma Tan et al. 2006 [replaced]
G. †Polyakius Kirejtshuk 2020
+
†Polyakius alberti Kirejtshuk 2020
†Polyakius pubescens Kirejtshuk 2020
G. †Rhopalomma Ashman et al. 2015
+
†Rhopalomma stefaniae Ashman et al. 2015
G. †Stegocoleus Jarzembowski and Wang 2016
+
†Stegocoleus arkonus Tihelka et al. 2019
†Stegocoleus caii Jarzembowski and Wang 2016
†Stegocoleus lawrencei Tihelka et al. 2019
G. †Tetraphalerites Crowson 1962
+
G. Tetraphalerus Waterhouse 1901
+
†Tetraphalerus brevis Ponomarenko 1964
†Tetraphalerus collaris Ponomarenko 1997
†Tetraphalerus glabratus Ponomarenko 1997
†Tetraphalerus mongolicus Ponomarenko 1986
†Tetraphalerus notatus Ponomarenko 1997
†Tetraphalerus penalveri Soriano and Delclos 2006
†Tetraphalerus ponomarenkoi Soriano and Delclos 2006
Invalid names: Lithocupedini Ponomarenko 1969 [invalid subgroup], Notocupedinae Ponomarenko 1966 [synonym], Pronotocupedini Tan et al. 2012 [invalid subgroup], Tetraphalerini Crowson 1962 [invalid subgroup], Tetraphalerus brevicapitis Ponomarenko and Martínez-Delclòs 2000 [nomen dubium]
Subfm. Tetraphalerinae Crowson 1962
+
G. †Bukhkalius Kirejtshuk and Jarzembowski 2020
+
†Bukhkalius lindae Jarzembowski et al. 2017
Invalid names: Brochocoleinae Hong 1982 [empty]
Invalid names: Crowsoniellidae Iablokoff-Khnzorian 1983 [empty], Magnocoleidae Hong 1998 [invalid subgroup]
G. †Dictyocoleus Hong 1982
+
G. †Doggeriopsis Handlirsch 1906
+
†Doggeriopsis stonesfieldiana Handlirsch 1906
G. †Eumecoleus Haupt 1950
+
†Eumecoleus tenuis Haupt 1950
G. †Euteticoleus Hong 1983
+
†Euteticoleus radiatus Hong 1983
G. †Gansucupes Hong 1982
+
†Gansucupes attenuatus Hong 1982
G. †Hebeicoleus Hong 1992
+
G. †Hebeicupes Zhang 1986
+
G. †Helopides Roemer 1876
+
†Helopides hildesiensis Roemer 1876
G. †Ironicus Handlirsch 1906
+
†Ironicus nothrus Westwood 1854
G. †Kargalarva Prokin et al. 2019
+
†Kargalarva permosialis Prokin et al. 2019
G. †Katapontisus Handlirsch 1906
+
†Katapontisus brodiei Giebel 1856
G. †Kelidus Handlirsch 1906
+
†Kelidus bolbus Westwood 1854
G. †Lecanopteryx Haupt 1950
+
†Lecanopteryx miranda Haupt 1950
G. †Longaevicupes Ren 1995
+
G. †Metacupes Gardiner 1961
+
†Metacupes harrisi Gardiner 1961
G. †Nannocurculionites Handlirsch 1906
+
G. †Notocupes Ponomarenko 1964
+
†Notocupes alienus Tan and Ren 2006
†Notocupes caudatus Ponomarenko 1966
†Notocupes cyclodontus Tan et al. 2006
†Notocupes daohugouensis Li and Cai 2023
†Notocupes diazromerali Soriano and Delclos 2006
†Notocupes elegans Ponomarenko 1994
†Notocupes excellens Ponomarenko 1966
†Notocupes kezuoensis Hong 1987
†Notocupes khasurtyiensis Strelnikova 2019
†Notocupes laetus Lin 1976
†Notocupes laiyangensis Hong and Wang 1990
†Notocupes lapidarius Ponomarenko 1968
†Notocupes latus Ponomarenko 1969
†Notocupes lini Ponomarenko et al. 2012
†Notocupes ludongensis Wang and Liu 1996
†Notocupes mongolicus Ponomarenko 1994
†Notocupes neli Tihelka et al. 2019
†Notocupes ohmkuhnlei Jarzembowski et al. 2019
†Notocupes picturatus Ponomarenko 1964
†Notocupes pingi Ponomarenko and Ren 2010
†Notocupes pulcher Ponomarenko 1968
†Notocupes reticulatus Oppenheim 1888
†Notocupes robustus Li and Cai 2023
†Notocupes rudis Tan et al. 2005
†Notocupes shiluoensis Hong 1984
†Notocupes tuanwangensis Hong and Wang 1990
†Notocupes validus Lin 1976
Invalid names: Amblomma Tan et al. 2005 [synonym], Chengdecupes Hong 1983 [synonym], Forticupes Hong and Wang 1990 [synonym], Lupicupes Ren 1995 [synonym], Ovatocupes Tan and Ren 2006 [synonym], Picticupes Hong and Wang 1990 [synonym], Sinocupes Lin 1976 [synonym]
G. †Parabuprestites Handlirsch 1906
+
G. †Paracurculionites Handlirsch 1906
+
G. †Paradoggeria Handlirsch 1906
+
†Paradoggeria acuminata Handlirsch 1906
G. †Pseudosilphites Zeuner 1930
+
G. †Rhabdocupes Ponomarenko 1966
+
†Rhabdocupes baculatus Ponomarenko 1969
†Rhabdocupes cellulosus Ponomarenko 1969
†Rhabdocupes dischides Zhang 1986
†Rhabdocupes issykkulensis Ponomarenko 1969
†Rhabdocupes laticella Ponomarenko 1969
†Rhabdocupes lentus Ren 1995
†Rhabdocupes longus Ponomarenko 1966
†Rhabdocupes minisculus Tan et al. 2006
†Rhabdocupes minor Ponomarenko 1966
†Rhabdocupes oxypygus Ponomarenko 1969
†Rhabdocupes protensus Tan et al. 2006
†Rhabdocupes rostratus Ponomarenko 1969
†Rhabdocupes sogutensis Ponomarenko 1969
†Rhabdocupes tenuis Ponomarenko 1969
†Rhabdocupes vitimensis Ponomarenko 1966
Superfm. †Rhombocoleoidea Rohdendorf 1961
+
Fm. †Rhombocoleidae Rohdendorf 1961
+
G. †Rhombocoleites Ponomarenko 1969
+
†Rhombocoleites adumbratus Ponomarenko 1969
†Rhombocoleites danutae Ponomarenko and Mostovski 2005
†Rhombocoleites uralensis Ponomarenko 2013
G. †Rhombocoleus Rohdendorf 1961
+
†Rhombocoleus andreae Rohdendorf 1961
†Rhombocoleus gomankovi Ponomarenko 2013
G. †Rossocoleus Rohdendorf 1961
+
†Rossocoleus altus Ponomarenko 2004
†Rossocoleus angustus Ponomarenko 2013
†Rossocoleus aristovi Ponomarenko 2013
†Rossocoleus avus Rohdendorf 1961
†Rossocoleus ellipticus Martynov 1937
†Rossocoleus grandis Rohdendorf 1961
†Rossocoleus latior Rohdendorf 1961
†Rossocoleus latus Rohdendorf 1961
†Rossocoleus novojilovi Ponomarenko 2013
†Rossocoleus parvus Rohdendorf 1961
†Rossocoleus sakmara Ponomarenko 2013
†Rossocoleus surijokovensis Rohdendorf 1961
†Rossocoleus verrucosus Rohdendorf 1961
Invalid names: Curculiopsis Martynov 1937 [replaced]
G. †Sinorhombocoleus Tan and Ren 2009
+
†Sinorhombocoleus papposus Tan and Ren 2009
Superfm. †Schizocoleoidea Rohdendorf 1961
+
Fm. †Phoroschizidae Bouchard and Bousquet 2020
+
G. †Phoroschizus Bouchard and Bousquet 2020
+
†Phoroschizus crassus Ponomarenko 1968
Invalid names: Schizophorus Ponomarenko 1968 [replaced]
Invalid names: Schizophoridae Ponomarenko 1968 [replaced]
Fm. †Schizocoleidae Rohdendorf 1961
+
G. †Aenigmocoleus Rohdendorf 1961
+
†Aenigmocoleus borissiaki Rohdendorf 1961
†Aenigmocoleus uralensis Ponomarenko 2013
G. †Argentinosyne Martins-Neto and Gallego 2006
+
†Argentinosyne bonapartei Martins-Neto and Gallego 2006
†Argentinosyne duraznoensis Martins-Neto and Gallego 2008
†Argentinosyne frenguellii Martins-Neto and Gallego 2006
†Argentinosyne gonaldiae Martins-Neto and Gallego 2006
†Argentinosyne gualoensis Martins-Neto and Gallego 2006
†Argentinosyne ischichucaensis Martins-Neto and Gallego 2009
†Argentinosyne losrastrosensis Martins-Neto and Gallego 2006
†Argentinosyne rugosa Martins-Neto and Gallego 2006
G. †Erunakicupes Rohdendorf 1961
+
†Erunakicupes angustus Ponomarenko 2011
†Erunakicupes brevis Ponomarenko 2004
†Erunakicupes fomitshevi Rohdendorf 1961
†Erunakicupes grossus Ponomarenko 2013
†Erunakicupes kryshtofovichi Rohdendorf 1961
†Erunakicupes obrutshevi Rohdendorf 1961
†Erunakicupes schmalhauseni Rohdendorf 1961
Invalid names: Schizocupes Rohdendorf 1961 [synonym]
G. †Karakanocoleus Rohdendorf 1961
+
†Karakanocoleus europeus Ponomarenko 2004
†Karakanocoleus janischewskyi Rohdendorf 1961
†Karakanocoleus magnus Rohdendorf 1961
†Karakanocoleus major Rohdendorf 1961
+
Invalid names: Karakanocoleodes latissimus Rohdendorf 1961 [synonym]
†Karakanocoleus mutovinensis Ponomarenko 2013
†Karakanocoleus pincombeae Tillyard 1924
†Karakanocoleus striatus Rohdendorf 1961
†Karakanocoleus venjukovi Rohdendorf 1961
+
Invalid names: Karakanocoleus lebedevi Rohdendorf 1961 [synonym]
Invalid names: Karakanocoleodes Rohdendorf 1961 [synonym]
G. †Mesolobites Carpenter 1986
+
†Mesolobites granulatus Dunstan 1923
†Mesolobites margacrispus Lin and Mou 1989
†Mesolobites quadripartita Dunstan 1923
†Mesolobites trivittatus Dunstan 1923
†Mesolobites tuberculatus Dunstan 1923
Invalid names: Lobites Dunstan 1923 [replaced], Shijingocoleus Lin and Mou 1989 [synonym]
G. †Mesosagrites Martynov 1935
+
†Mesosagrites multipunctatus Martynov 1935
G. †Metrorhynchites Tillyard 1916
+
†Metrorhynchites angustior Rohdendorf 1961
†Metrorhynchites angustum Tillyard 1918
†Metrorhynchites angustus Rohdendorf 1961
†Metrorhynchites baculum Ponomarenko 2011
†Metrorhynchites dunstani Tillyard 1916
†Metrorhynchites elongatus Ponomarenko 2011
†Metrorhynchites grandis Tillyard 1918
†Metrorhynchites longus Rohdendorf 1961
†Metrorhynchites sydneiensis Tillyard 1916
Invalid names: Mancicupes Lin 1986 [synonym]
G. †Mimemala Strand 1936
+
†Mimemala giganteum Whalley 1985
†Mimemala punctatum Handlirsch 1906
Invalid names: Mimema Handlirsch 1906 [replaced]
G. †Pseudochrysomelites Handlirsch 1906
+
†Pseudochrysomelites anser Ponomarenko 2008
†Pseudochrysomelites bashkuevi Ponomarenko 2013
†Pseudochrysomelites convexus Ponomarenko 2015
†Pseudochrysomelites dilatatus Ponomarenko 2008
†Pseudochrysomelites dvinensis Ponomarenko 2018
†Pseudochrysomelites elongatus Ponomarenko 2004
†Pseudochrysomelites humeralis Ponomarenko 2016
†Pseudochrysomelites latissimus Ponomarenko 2015
†Pseudochrysomelites latum Ponomarenko 2004
†Pseudochrysomelites longus Ponomarenko 2013
†Pseudochrysomelites major Ponomarenko 2011
†Pseudochrysomelites medialis Ponomarenko 2013
†Pseudochrysomelites ovum Ponomarenko 2004
†Pseudochrysomelites planus Ponomarenko 2008
†Pseudochrysomelites tumidus Ponomarenko 2016
G. †Reeveana Dunstan 1923
+
†Reeveana intermedia Dunstan 1923
†Reeveana major Dunstan 1923
†Reeveana minor Dunstan 1923
G. †Schizocoleus Rohdendorf 1961
+
†Schizocoleus depressus Ponomarenko 2013
†Schizocoleus glabrus Ponomarenko 2004
†Schizocoleus kuznetskiensis Rohdendorf 1961
†Schizocoleus longus Ponomarenko 2004
†Schizocoleus major Ponomarenko 2013
†Schizocoleus minimus Ponomarenko 2013
†Schizocoleus robustus Ponomarenko 2013
†Schizocoleus septentrionalis Ponomarenko 2008
†Schizocoleus triassicus Ponomarenko 2016
G. †Shepherdia Dunstan 1923
+
†Shepherdia quadrivittata Dunstan 1923
†Shepherdia trivittata Ponomarenko 1985
G. †Stegosyne Rohdendorf 1961
+
†Stegosyne borealis Ponomarenko 2008
†Stegosyne rugosa Rohdendorf 1961
G. †Uskatocoleus Rohdendorf 1961
+
†Uskatocoleus artus Ponomarenko 2015
†Uskatocoleus avitus Rohdendorf 1961
†Uskatocoleus convexus Rohdendorf 1961
†Uskatocoleus convexus Ponomarenko 2013
†Uskatocoleus eurypygus Ponomarenko 2011
†Uskatocoleus foveolatus Rohdendorf 1961
†Uskatocoleus hirsutus Ponomarenko 2015
†Uskatocoleus iljinskiensis Rohdendorf 1961
†Uskatocoleus kaltanicus Rohdendorf 1961
†Uskatocoleus kushejakovicus Rohdendorf 1961
†Uskatocoleus lutugini Rohdendorf 1961
†Uskatocoleus micron Ponomarenko 2013
†Uskatocoleus minimus Rohdendorf 1961
†Uskatocoleus minor Ponomarenko 2015
†Uskatocoleus rhynchophorus Ponomarenko 2011
†Uskatocoleus sukhonensis Ponomarenko 2013
†Uskatocoleus surijokovicus Rohdendorf 1961
†Uskatocoleus uralensis Ponomarenko 2013
†Uskatocoleus zalesskyi Rohdendorf 1961
Invalid names: Schizophoroidea Ponomarenko 1968 [synonym]
G. †Stigmenamma Handlirsch 1906
+
†Stigmenamma heeri Giebel 1856
G. †Synodus Hong 1982
+
†Synodus changmaensis Hong 1982
G. †Tentyridium Westwood 1854
+
†Tentyridium dilatatum Handlirsch 1906
†Tentyridium peleus Westwood 1854
G. †Tetrocupes Hong 1983
+
†Tetrocupes cavernasus Hong 1983
G. †Tillyardiopsis Dunstan 1923
+
†Tillyardiopsis granulata Dunstan 1923
G. †Zygadenia Handlirsch 1906
+
†Zygadenia alexrasnitsyni Strelnikova and Yan 2021
†Zygadenia angliae Giebel 1856
†Zygadenia cornigera Lee et al. 2023
†Zygadenia floodpagei Jarzembowski et al. 2015
†Zygadenia giebeli Ponomarenko 2014
†Zygadenia handlirschi Ponomarenko 2014
†Zygadenia liui Jarzembowski et al. 2015
†Zygadenia martinae Jarzembowski et al. 2015
†Zygadenia semen Ponomarenko 2000
†Zygadenia sibirica Ponomarenko 2000
†Zygadenia simpsoni Jarzembowski et al. 2015
†Zygadenia sinitzae Ponomarenko 2000
†Zygadenia tuberculata Giebel 1856
Invalid names: Camtodontus Dejean 1826 [synonym], Kakoselia Handlirsch 1906 [synonym]
Invalid names: Cupediformia [invalid subgroup], Moravocoleus perditus Kukalová 1968 [nomen dubium], Notocupedini Ponomarenko 1966 [invalid subgroup], Priacma corrupta Ponomarenko 1986 [nomen dubium], Priacma longicapitis Ponomarenko 1997 [nomen dubium], Priacma oculata Ponomarenko 1997 [nomen dubium], Priacma striata Ponomarenko 2000 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Asselian to the top of the Piacenzian or 298.90000 to 2.58800 Ma
Minimum age of oldest fossil (stem group age): 295.5 Ma

Collections (284 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Asselian298.9 - 295.5Czech Republic Moravocoleus perditus (114186)
Roadian272.3 - 268.8Russian Federation (Kemerovo) Rossocoleus avus (122567) Schizocoleus kuznetskiensis (124604) Uskatocoleus kaltanicus (104704)
Roadian272.3 - 268.8Russian Federation Stegosyne rugosa (124611)
Roadian272.3 - 268.8Russian Federation (Kirov) Rossocoleus altus (105058)
Wordian266.9 - 264.28Russian Federation (Udmurtia) Rhombocoleites uralensis (104750) Schizophoridae indet., Rossocoleus novojilovi, Rossocoleus angustus (103502)
Wordian266.9 - 264.28Russian Federation (Kemerovo) Schizocupes obrutshevi (124600) Uskatocoleus avitus, Uskatocoleus iljinskiensis, Uskatocoleus surijokovicus, Karakanocoleus janischewskyi, Rossocoleus grandis, Rossocoleus latior, Rossocoleus parvus, Rossocoleus surijokovensis, Rossocoleus latus (122566) Uskatocoleus convexus, Rossocoleus verrucosus (124603)
Wordian - Capitanian266.9 - 259.51Russian Federation Uskatocoleus kushejakovicus (124610)
Capitanian264.28 - 259.51Russian Federation (Orenburg) Curculiopsis ellipticus (104994) Kargalarva permosialis, Rhombocoleites adumbratus (104057) Schizocoleus major, Aenigmocoleus uralensis, Rossocoleus sakmara (152235) Schizocoleus minimus, Schizocoleus robustus, Schizocoleus longus, Schizocoleus glabrus, Erunakicupes brevis, Karakanocoleus europeus, Palademosyne ovum, Rhombocoleus gomankovi (104717)
Capitanian264.28 - 259.51China (Anhui) Dikerocoleus divisus (103239)
Wuchiapingian259.9 - 254.17Russian Federation (Orenburg) Uskatocoleus uralensis (152236)
Wuchiapingian259.9 - 254.17Russian Federation (Vologda) Uskatocoleus sukhonensis, Uskatocoleus micron, Uskatocoleus convexus, Schizocoleus depressus, Erunakicupes grossus, Karakanocoleus mutovinensis, Pseudochrysomelites longus, Pseudochrysomelites sphenoidalis, Pseudochrysomelites medialis, Pseudochrysomelites bashkuevi, Rossocoleus aristovi (104754)
Wuchiapingian259.9 - 254.17Russian Federation Uskatocoleus zalesskyi (122565) Uskatocoleus zalesskyi, Uskatocoleus minimus, Uskatocoleus foveolatus, Uskatocoleus lutugini, Erunakicupes schmalhauseni, Erunakicupes fomitshevi, Erunakicupes kryshtofovichi, Erunakicupes venjukovi, Karakanocoleus lebedevi, Karakanocoleus major, Karakanocoleodes latissimus, Karakanocoleus magnus, Karakanocoleus striatus, Uskatocoleus angustior, Uskatocoleus angustus, Uskatocoleus longus, Aenigmocoleus borissiaki, Rhombocoleus andreae (124608)
Changhsingian254.17 - 252.17Russian Federation (Vologda) Pseudochrysomelites dvinensis (105041) Schizocoleidae indet., Uskatocoleus hirsutus, Uskatocoleus minor, Uskatocoleus artus, Pseudochrysomelites circumflexus, Pseudochrysomelites latissimus, Palademosyne latum, Pseudochrysomelites convexus (105046)
Changhsingian254.17 - 252.17South Africa (Kwazulu-Natal) Rhombocoleites danutae (14357)
Changhsingian254.17 - 252.17Russian Federation (Vladimir) Schizocoleidae indet., Uskatocoleus eurypygus, Uskatocoleus rhynchophorus, Erunakicupes angustus, Metrorhynchites elongatus, Metrorhynchites baculum, Pseudochrysomelites major (113351)
Changhsingian254.17 - 252.17Australia (New South Wales) Permosyne pincombeae (122414)
Griesbachian251.902 - 250.5Russian Federation (Vologda) Pseudochrysomelites tumidus (105056)
Griesbachian251.902 - 250.5Russian Federation (Kemerovo) Archaeomalthus synoriakos, Permocatinus tomiensis (104713)
Olenekian251.2 - 247.2Russian Federation (Yaroslavl) Schizocoleus triassicus, Pseudochrysomelites humeralis, Palademosyne elongatum (105043)
Olenekian251.2 - 247.2Russian Federation (Arkhangelsk) Schizocoleus septentrionalis (108318)