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Pelecaniformes

Reptilia - Theropoda

Synonymy list
YearName and author
1891Pelecaniformes Sharpe
1930Pelecaniformes Hay p. 289
1940Pelecaniformes Wetmore p. 10
1956Pelecaniformes Wetmore p. 12
1960Pelecaniformes Wetmore p. 24
1972Pelecaniformes Howard p. 5
1972Pelicaniformes Simpson p. 30
1988Pelecaniformes Carroll
1992Pelecaniformes Baird p. 24
1992Pelecaniformes Guthrie p. 321
2000Pelecaniformes Kurochkin p. 551
2002Pelecaniformes Hope p. 367
2002Pelecaniformes Sepkoski
2005Pelecaniformes Mayr p. 318
2007Pelecaniformes Livezey and Zusi p. 91
2008Pelecaniformes Louchart et al. p. 151
2014Pelecaniformes Stidham et al. p. 2
2014Pelecaniformes del Hoyo et al.
2021Pelecaniformes Sangster and Mayr p. 50
2022Pelecaniformes Mayr p. 119 figs. Fig. 7.1

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
RankNameAuthor
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
orderTheropoda
Neotheropoda
AverostraPaul 2002
Tetanurae
Coelurosauria()
Maniraptora
Paraves
classAves
subclassNeornithesGadow 1893
Aequornithes
Feraequornithes
superorderPelecanimorphaeHuxley 1867
orderPelecaniformesSharpe 1891
orderPelecaniformesSharpe 1891

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Pelecaniformes Sharpe 1891
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Fm. Ardeidae Leach 1820 [heron]
+
G. Agamia Reichenbach 1853 [Agami heron]
+
Agamia agami Gmelin 1789 [Agami heron]
G. Ardea Linnaeus 1758 [heron]
+
Ardea alba Linnaeus 1758 [great egret]
Ardea alba egretta Gmelin 1789
Ardea aurelianensis Milne-Edwards 1871
Ardea brachyrhyncha Brehm 1854 [Yellow-billed egret]
Ardea brunhuberi von Ammon 1911
Ardea cinerea Linnaeus 1758 [gray heron]
Ardea cocoi Linnaeus 1766 [cocoi heron]
Ardea effosa von Meyer 1843
Ardea formosa Milne-Edwards 1871
Ardea goliath Cretzschmar 1827 [Goliath heron]
Ardea herodias Linnaeus 1758 [great blue heron]
Ardea humbloti Milne-Edwards and Grandidier 1885 [Humblot's heron]
Ardea insignis Hume 1878 [White-bellied heron]
Ardea intermedia Wagler 1829 [intermediate egret]
Ardea latipes von Meyer 1843
Ardea lignitum Gibel 1860
Ardea melanocephala Vigors and Children 1826 [Black-headed heron]
Ardea modesta Gray 1831
Ardea pacifica Latham 1801 [White-necked heron]
Ardea paloccidentalis Shufeldt 1891
Ardea piveteaui Brunet 1970
Ardea plumifera Gould 1848 [Plumed egret]
Ardea polkensis Brodkorb 1955
Ardea purpurea Linnaeus 1766 [purple heron]
Ardea sumatrana Raffles 1822 [Great-billed heron]
Invalid names: Casmerodius Gloger 1842 [synonym], Herodias Boie 1822 [synonym]
G. †Ardeacites Haushalter 1855
+
Ardeacites molassicus Haushalter 1855
G. †Ardeagrandis Kurochkin and Ganea 1972
+
Ardeagrandis arborea Kurochkin and Ganea 1972
Subfm. Ardeinae
+
G. Ardeola Boie 1822 [pond heron]
+
Ardeola ralloides Scopoli 1769 [Squacco heron]
G. †Proardea Lambrecht 1933
+
Proardea amissa Milne-Edwards 1892
Proardea deschutteri Mayr et al. 2019
G. †Proardeola Harrison 1979
+
Proardeola walkeri Harrison 1979
Invalid names: Ardeini Bock 1956 [empty], Nycticoracini Bonaparte 1854 [empty]
Subfm. Botaurinae Reichenbach 1850
+
G. †Botauroides Shufeldt 1915
+
Botauroides parvus Shufeldt 1915
G. †Eoceornis Shufeldt 1915
+
Eoceornis ardetta Shufeldt 1915
G. †Pikaihao Worthy et al. 2013
+
Pikaihao bartlei Worthy et al. 2013
Tr. Tigriornithini Bock 1956
+
G. Tigrisoma Swainson 1827
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Tigrisoma lineatum Boddaert 1783 [Rufescent tiger-heron]
Tigrisoma mexicanum Swainson 1834 [bare-throated tiger-heron]
Invalid names: Botaurini [empty]
G. †Botaurites von Ammon 1918
+
Botaurites avitus von Ammon 1918
G. Botaurus Stephens 1819
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Botaurus lentiginosus Rackett 1813 [American bittern]
Botaurus pinnatus Wagler 1829 [Pinnated bittern]
Botaurus poiciloptilus Wagler 1827 [Australasian bittern]
Botaurus stellaris Linnaeus 1758 [great bittern]
G. Bubulcus Bonaparte 1855 [Cattle egret]
+
Bubulcus ibis Linnaeus 1758 [cattle egret]
G. Butorides Blyth 1852
+
Butorides striata Linnaeus 1758 [striated heron]
Butorides validipes Campbell 1976
Butorides virescens Linnaeus 1758 [green heron]
G. Cochlearius Brisson 1760
+
Cochlearius cochlearius Linnaeus 1766 [Boat-billed heron]
G. Egretta Forster 1817 [egret]
+
Egretta caerulea Linnaeus 1758 [little blue heron]
Egretta garzetta Linnaeus 1766 [little egret]
Egretta garzetta dimorpha Hartert 1914 [dimorphic egret]
Egretta novaehollandiae Latham 1790 [White-faced heron]
Egretta picata Gould 1845 [pied heron]
Egretta sacra Gmelin 1789 [Pacific reef-heron]
Egretta subfluvia Becker 1985
Egretta thula Molina 1782 [snowy egret]
Egretta tricolor Statius Muller 1776 [tricolor heron]
Invalid names: Florida Baird 1858 [synonym], Hydranassa Baird 1858 [synonym], Leucophoyx Sharpe 1894 [synonym]
G. Ixobrychus Billberg 1828 [Least bittern]
+
Ixobrychus cinnamomeus Gmelin 1789 [Cinnamon bittern]
Ixobrychus dubius Matthews 1912 [Australian little bittern]
Ixobrychus eurhythmus Swinhoe 1873 [Schrenck's bittern]
Ixobrychus exilis Gmelin 1789 [least bittern]
Ixobrychus flavicollis Latham 1790 [Black bittern]
Ixobrychus involucris Vieillot 1823 [Stripe-backed bittern]
Ixobrychus minutus Linnaeus 1766 [Common little bittern]
Ixobrychus novaezelandiae Potts 1871 [New Zealand little bittern]
Ixobrychus sinensis Gmelin 1789 [Yellow bittern]
Ixobrychus sturmii Wagler 1827 [Dwarf bittern]
G. †Matuku Scofield et al. 2010
+
Matuku otagoense Scofield et al. 2010
G. Nyctanassa Stejneger 1887
+
Nyctanassa carcinocatactes Olson and Wingate 2006 [Bermuda night heron]
Nyctanassa violacea Linnaeus 1758 [yellow-crowned night heron]
+
Invalid names: Larus vero Shufeldt 1917 [synonym]
G. Nycticorax Forster 1817 [night heron]
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Nycticorax caledonicus Gmelin 1789 [rufous night-heron]
Nycticorax duboisi Rothschild 1907 [Réunion night heron]
+
Invalid names: Nycticorax borbonensis Cowles 1994 [synonym]
Nycticorax fidens Brodkorb 1963
Nycticorax kalavikai Steadman et al. 2000 [Niue night heron]
Nycticorax mauritianus Newton and Gadow 1893 [Mauritius night heron]
Nycticorax megacephalus Milne-Edwards 1874 [Rodrigues night heron]
Nycticorax nycticorax Linnaeus 1758 [black-crowned night heron]
Nycticorax olsoni Bourne et al. 2003 [Ascension night heron]
G. †Palaeophoyx McCoy 1963
+
Palaeophoyx columbiana McCoy 1963
G. Pilherodius Reichenbach 1853 [Capped heron]
+
Pilherodius pileatus Boddaert 1783 [Capped heron]
G. Syrigma Ridgway 1824 [whistling heron]
+
Syrigma sanctimartini Campbell 1979
Syrigma sibilatrix Temminck 1824 [Whistling heron]
G. †Taphophoyx Steadman and Takano 2019
+
Taphophoyx hodgei Steadman and Takano 2019
G. †Zeltornis Balouet 1981
Subor. †Cladornithes Wetmore 1960
+
Fm. †Cladornithidae Ameghino
+
G. †Cladornis Ameghino 1895
+
Cladornis pachypus Ameghino 1894
G. †Cruschedula Ameghino 1899
+
Cruschedula revola Ameghino 1899
Invalid names: Cruschedulidae Ameghino 1899 [synonym]
G. Gorsachius Bonaparte 1855
+
Fm. Pelecanidae Vigors 1825 [pelican]
+
G. †Eopelecanus El Adli et al. 2021
+
Eopelecanus aegyptiacus El Adli et al. 2021
G. †Miopelecanus Cheneval 1984
+
Miopelecanus gracilis Milne-Edwards 1867
+
Invalid names: Pelecanus fraasi Lyddekker 1891 [synonym]
G. Pelecanus Linnaeus 1758 [pelican]
+
Subg. †Pelecanus (Cyrtopelicanus) Reichenbach 1852
+
Subg. Pelecanus (Leptopelicanus) Reichenback 1852
Pelecanus cadimurka Rich and Van Tets 1981
Pelecanus cautleyi Davies 1880
Pelecanus conspicillatus Temminck 1824 [Australian pelican]
+
Invalid names: Pelecanus grandiceps De Vis 1905 [synonym]
Pelecanus crispus Bruch 1832 [Dalmatian pelican]
Pelecanus erythrorhynchos Gmelin 1789 [American white pelican]
Pelecanus novaezealandiae Scarlett 1966
Pelecanus occidentalis Linnaeus 1766 [brown pelican]
Pelecanus odessanus Widhalm 1866
Pelecanus onocrotalus Linnaaeus 1758
Pelecanus proavus De Vis 1892
Pelecanus rufescens Gmelin 1789 [pink-backed pelican]
Pelecanus schreiberi Olson 1999
Pelecanus sivalensis Davies 1880
Pelecanus thagus Molina 1782 [Peruvian pelican]
Pelecanus tirarensis Miller 1966
G. †Protopelicanus Reichenbach 1852
+
Protopelicanus cuvierii Reichenbach 1852
Fm. †Protoplotidae van Tets et al. 1989
+
G. †Protoplotus Lambrecht 1931
+
Protoplotus beauforti Lambrecht 1931
Subor. †Steganopodes
+
Infraor. Fregatides
+
Invalid names: Fregatoidea [empty]
Unr. †Pelecani
+
Superfm. Pelecanoidea Rafinesque 1815
+
Fm. †Cyphornithidae Wemore 1928
+
G. †Palaeochenoides Shufeldt 1916
Infraor. Pelecanides
+
Unr. Sulida Reichenbach 1849
+
Superfm. Suloidea Hay 1930
+
Fm. †Elopterygidae Lambrecht 1933
Invalid names: Phalacrocoracoidea [empty]
Invalid names: Pelecanida [empty]
G. Plotus Linnaeus 1766
+
Plotus pannonicus Lambrect 1916
Fm. Threskiornithidae Richmond 1917 [ibis]
+
G. Bostrychia Gray 1847
+
G. Cercibis Wagler 1825
+
G. Eudocimus Wagler 1832
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Eudocimus albus Linnaeus 1758 [White ibis]
Eudocimus leiseyi Emslie 1995
Eudocimus peruvianus Campbell 1979 [ibis]
Eudocimus ruber Linnaeus 1758 [Scarlet ibis]
Invalid names: Guara Reichenbach 1852 [synonym]
G. Geronticus Wagler 1832 [bald ibis]
+
Geronticus eremita Linnaeus 1758 [northern bald ibis]
Geronticus perplexus Milne-Edwards 1871
G. Lophotibis Reichebach 1853
+
Lophotibis cristata Boddaert 1783 [Madagascar ibis]
G. Mesembrinibis Peters 1930
+
G. †Milnea Lydekker 1891
+
Milnea gracilis Lydekker 1891
G. Nipponia Reichenbach 1850 [Asian crested ibis]
+
Nipponia nippon Temminck 1835 [Asian crested ibis]
G. Phimosus Wagler 1832
+
Phimosus infuscatus Lichtenstein 1823
G. Platalea Linnaeus 1758 [spoonbills]
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Platalea ajaja Linnaeus 1758 [roseate spoonbill]
+
Invalid names: Platea rosea Brisson 1760 [synonym]
Platalea alba Scopoli 1786 [African spoonbill]
+
Invalid names: Platalea tenuirostris Temminck 1820 [synonym]
Platalea flavipes Gould 1838 [yellow-billed spoonbill]
Platalea leucorodia Linnaeus 1758 [Eurasian spoonbill]
Platalea minor Temminck and Schlegel 1849 [Black-faced spoonbill]
Platalea regia Gould 1838 [Royal spoonbill]
Platalea subtenuis De Vis 1892
Invalid names: Ajaja Reichenbach 1853 [synonym], Platibis Bonaparte 1856 [synonym]
G. Plegadis Kaup 1829
+
Plegadis autumnalis Hasselqvist 1762
Plegadis chihi Vieillot 1817 [white-faced ibis]
Plegadis falcinellus Linnaeus 1766 [glossy ibis]
Plegadis paganus Milne-Edwards 1868
Plegadis pharangites Olson 1981
+
Invalid names: Plegadis gracilis Miller and Bowman 1956 [replaced]
G. †Protibis Ameghino 1891
+
Protibis cnemialis Ameghino 1891
G. Pseudibis Hodgson 1844
+
Pseudibis ibis Temminck 1824
Subfm. †Rhynchaeitinae Mayr 2002
+
G. †Rhynchaeites Wittich 1898
+
Rhynchaeites litoralis Mayr and Kitchener 2023
Rhynchaeites messelensis Wittich 1898
Invalid names: Plumumida Hoch 1980 [synonym]
G. Thaumatibis Elliot 1877
+
Thaumatibis gigantea Oustalet 1877
G. Threskiornis Gray 1842
+
Threskiornis aethiopicus Latham 1790 [sacred ibis]
Threskiornis bernieri Bonaparte 1855 [Madagascar sacred ibis]
Threskiornis melanocephalus Latham 1790 [Black-headed ibis]
Threskiornis molucca Cuvier 1829 [Australian ibis]
Threskiornis solitarius Selys-Longchamps 1848 [Réunion ibis]
+
Invalid names: Borbonibis latipes Mourer-Chauviré and Moutou 1987 [synonym]
Threskiornis spinicollis Jameson 1835 [Straw-necked ibis]
Invalid names: Apterornis Selys-Longchamps 1848 [synonym], Borbonibis Mourer-Chauviré and Moutou 1987 [synonym]
Subfm. Threskiornithinae Richmond 1917 [ibis]
+
G. Theristicus Wagler 1832 [ibis]
+
Theristicus wetmorei Campbell 1979 [ibis]
G. †Vadaravis Smith et al. 2013
+
Vadaravis brownae Smith et al. 2013
G. †Xenicibis Olson and Steadman 1977
+
Xenicibis xympithecus Olson and Steadman 1977
Invalid names: Plataleinae Bonaparte 1838 [empty]
G. Tigriornis Sharpe 1895
+
Tigriornis leucolopha Jardine 1846
G. †Torotix Brodkorb 1963
+
Torotix clemensi Brodkorb 1963
G. Zebrilus Bonaparte 1855
+
Zebrilus undulatus Gmelin 1789
G. Zonerodius Salvadori 1882
+
Invalid names: Fregatae Sharpe 1891 [empty], Odontopteryges Lambrecht 1933 [empty], Odontopterygia Spulski 1910 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: hydroxyapatiteo
Entire body: yeso
Adult length: 10 to < 100o
Adult width: 1.0 to < 10o
Adult height: 1.0 to < 10o
Architecture: compact or denseo
Ontogeny: accretion, modification of partso
Grouping: solitaryo
Environment: marineo
Locomotion: actively mobileo
Life habit: ground dwellingo
Diet: carnivoresubo
Reproduction: oviparouso
Dispersal: direct/internalo
Dispersal 2: mobileo
Created: 2017-04-17 10:40:27
Modified: 2017-04-17 10:40:27
Source: subo = suborder, o = order
References: Bush and Bambach 2015, Marsh 1875, Benton 1983

Age range

Maximum range based only on fossils: base of the Late/Upper Maastrichtian to the top of the Holocene or 70.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 66.0 Ma

Collections (263 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Maastrichtian70.6 - 66.0USA (Wyoming) Torotix clemensi (14585)
Paleocene - Eocene66.0 - 33.9Indonesia (Sumatra) Protoplotus beauforti (142350)
Ypresian56.0 - 47.8United Kingdom (England) Rhynchaeites sp., Rhynchaeites litoralis (227361)
Wasatchian55.8 - 50.3USA (Wyoming) Vadaravis brownae (74617)
MP 1148.6 - 40.4Germany (Hessen) Rhynchaeites messelensis (22701)
Middle Eocene47.8 - 37.71USA (Wyoming) Botauroides parvus (81457) Eoceornis ardetta (81455)
Middle Eocene - Late/Upper Eocene47.8 - 33.9Chile Ardeidae indet. (99479)
Bartonian41.3 - 38.0Myanmar Threskiornithidae indet. (41393)
Priabonian38.0 - 33.9France (Ile-de-France Region) Protopelicanus cuvierii (31739)
Priabonian38.0 - 33.9Egypt (Fayum) Eopelecanus aegyptiacus (74738)
Priabonian38.0 - 33.9France (Ile-de-France) Ardea piveteaui (177531)
Late/Upper Eocene37.2 - 33.9United Kingdom (England) Actiornis anglicus (181272)
Early/Lower Oligocene33.9 - 28.4Belgium Proardea deschutteri (31748)
Early/Lower Oligocene33.9 - 28.4France Pelecaniformes indet. (69805)
Rupelian33.9 - 28.1USA (Washington) Phalacrocoracoidea indet. (62246)
Rupelian33.9 - 28.1Egypt (Fayum) Nycticorax sp. (12960)
Oligocene33.9 - 23.03Argentina Cladornis pachypus, Cruschedula revola (74644)
Late/Upper Oligocene28.4 - 23.03Australia (South Australia) Pelecanus tirarensis (51412 216700)
Aquitanian23.03 - 20.44France (Auvergne) Proardeola walkeri, Plegadis paganus, Miopelecanus gracillis (125333)
Miocene23.03 - 5.333USA (South Carolina) Palaeochenoides mioceanus (18506)