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Tenebrionoidea
Taxonomy
Tenebrionoidea was named by Latreille (1802). It is extant. Its type is Tenebrio.
It was assigned to Cucujiformia by Lawrence and Newton (1995), Bouchard et al. (2011), Robertson et al. (2015), Cai et al. (2022).
It was assigned to Cucujiformia by Lawrence and Newton (1995), Bouchard et al. (2011), Robertson et al. (2015), Cai et al. (2022).
Subtaxa
Aderidae (syn. Circaeidae, Xylophilidae), Afreminae, Anthicidae, Archaeoripiphorus, Archeocrypticidae, Asiamordella, Boridae, Chalcodryidae, Ciidae, Cossyphodinae, Elodophthalmidae, Eurygenius wickhami, Fushunicucujus, Glypta, Ischaliidae, Kulindrobor, Lagrioididae, Liaoximordellidae, Melandryidae, Meloidae, Mordellidae (syn. Praemordellidae), Mycetophagidae, Mycteridae, Neopolypria, Oedemeridae, Perimylopidae, Prostomidae, Pterogeniidae, Pyrochroidae, Pythidae, Rhipiphoridae, Ripiphoridae, Salpingidae, Samlandotoma, Scraptiidae, Stenotrachelidae, Synchroidae, Tenebrionidae, Tetratomidae, Trachelostenidae, Trictenotomidae, Ulodidae, Zopheridae (syn. Colydiidae)
Synonyms
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Synonymy list
Year | Name and author |
---|---|
1802 | Tenebrionoidea Latreille p. 165 |
1917 | Eurygenius wickhami Cockerell p. 324 fig. 2 |
1995 | Tenebrionoidea Lawrence and Newton, Jr. p. 786 |
2000 | Eurygenius wickhami Ross and York p. 12 |
2002 | Asiamordella Hong p. 126 |
2002 | Asiamordella furvis Hong p. 128 figs. Pl 10, fig 1; text-fig 2.6.73-81 |
2002 | Fushunicucujus Hong p. 137 |
2002 | Fushunicucujus maculatus Hong p. 139 figs. 2.6.106-114 |
2011 | Tenebrionoidea Bouchard et al. p. 64 |
2015 | Tenebrionoidea Robertson et al. |
2022 | Tenebrionoidea Cai et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Superfm. Tenebrionoidea Latreille 1802
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Fm. Aderidae Csiki 1909
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Subfm. Aderinae Csiki 1909
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G. †Palaeocnopus Alekseev and Grzymala 2015
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†Palaeocnopus densipunctatus Alekseev and Grzymala 2015
†Palaeocnopus glabricornis Alekseev and Grzymala 2015
†Palaeocnopus mara Alekseev and Grzymala 2015
†Palaeocnopus saeticornis Alekseev and Grzymala 2015
G. Vanonus Casey 1895
+
†Vanonus aestiorum Alekseev and Grzymala 2015
†Vanonus ulmerigicus Alekseev and Grzymala 2015
Invalid names: Circaeidae Iablokoff-Khnzorian 1961 [synonym], Xylophilidae [replaced]
Fm. Anthicidae Latreille 1819 [ant like flower beetle]
+
Subfm. Anthicinae Latreille 1819
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Tr. Anthicini Latreille 1819
+
G. Anthicus Paykull 1798
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†Anthicus neli Arillo and Ortuño 1997
†Anthicus sepultulus Cockerell 1926
Subfm. Eurygeniinae LeConte 1862
+
Subfm. Lemodinae Lawrence and Britton 1991
+
Subfm. Macratriinae LeConte 1862
+
Tr. †Camelomorphini Kirejtshuk et al. 2008
+
Tr. Macratriini Le Conte 1862
+
G. Macratria Newman 1838
+
†Macratria alleni Telnov 2012
†Macratria appendiculata Abdullah 1964
†Macratria gigantea Wickham 1910
†Macratria succinia Abdullah 1965
†Macratria tripunctata Abdullah 1964
Invalid names: Protomacratria Abdullah 1964 [synonym]
Subfm. Steropinae Jacquelin du Val 1863
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Subfm. Tomoderinae Bonadona 1961
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G. Tomoderus Laferte-Senectere 1849
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†Tomoderus balticus Telnov 2012
†Tomoderus longelytratus Telnov 2013
†Tomoderus saecularis Telnov and Bukejs 2019
Invalid names: Copobaeninae Abdullah 1969 [empty]
Fm. Boridae Thomson 1859
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Invalid names: Borinae Thomson 1859 [empty], Synercticinae Lawrence and Pollock 1994 [empty]
Fm. Ciidae Leach 1819 [minute tree-fungus beetle]
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Subfm. Ciinae Leach 1819
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Tr. Ciini Leach 1819
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Tr. Orophiini Thomson 1863
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G. Orthocis Casey 1898
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†Orthocis kompantsevi Legalov and Perkovsky 2024
†Orthocis larssoni Legalov and Perkovsky 2024
†Orthocis vilhelmseni Legalov and Perkovsky 2024
Invalid names: Sphindociinae Lawrence 1974 [empty]
Fm. †Elodophthalmidae Kirejtshuk and Azar 2008
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G. †Elodophthalmus Kirejtshuk and Azar 2008
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†Elodophthalmus gracilis Kirejtshuk and Azar 2008
†Elodophthalmus harmonicus Kirejtshuk and Azar 2008
†Elodophthalmus maksoudae Cai et al. 2024
Subfm. Ischaliidae Blair 1920 [false fire-colored beetle]
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G. Ischalia Pascoe 1860
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Subg. Ischalia (Eupleurida) LeConte 1862
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†Ischalia (Eupleurida) dohnaturris Alekseev and Bukejs 2017
Subg. †Ischalia (Telnovia) Alekseev and Bukejs 2017
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†Ischalia (Telnovia) danieli Alekseev and Bukejs 2017
G. †Prototrichalus Molina-Olmedo et al. 2020
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†Prototrichalus jingpo Telnov and Kundrata 2023
†Prototrichalus meiyingae Molina-Olmedo et al. 2020
†Prototrichalus milleri Molina-Olmedo et al. 2020
†Prototrichalus sepronai Molina-Olmedo et al. 2020
G. †Kulindrobor Tihelka et al. 2021
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†Kulindrobor enigmaticus Tihelka et al. 2021
†Kulindrobor magnus Tihelka et al. 2021
Fm. Melandryidae Leach 1815 [false darkling beetle]
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Subfm. Melandryinae Leach 1815
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Tr. Dircaeini Mulsant 1856
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Tr. Hypulini Gistel 1848
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G. †Madelinia Alekseev and Pankowski 2020
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†Madelinia capillata Alekseev and Bukejs 2021
†Madelinia gedanoposita Alekseev and Pankowski 2020
Tr. Orchesiini Mulsant 1856
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G. Microscapha Leconte 1866
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†Microscapha andrzeji Bukejs and Alekseev 2015
†Microscapha kugelanni Alekseev and Bukejs 2021
Tr. Serropalpini Latreille 1829
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G. Serropalpus Hellenius 1786
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†Serropalpus ingemmescus Alekseev 2014
†Serropalpus ryzhkovianus Alekseev 2014
†Serropalpus vivax Alekseev 2014
G. Symphora LeConte 1866
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†Symphora glaesonauta Alekseev and Bukejs 2021
†Symphora pollocki Alekseev and Bukejs 2021
Tr. Xylitini Thomson 1864
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Invalid names: Osphyinae Mulsant 1856 [empty]
Fm. Meloidae Gyllenhal 1810 [blister beetle]
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Subfm. Meloinae Gyllenhal 1810
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Tr. Epicautini Parker and Boving 1924
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G. Epicauta Dejean 1834
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†Epicauta rufithoraca Zhang 1989
†Epicauta sanctoruensis Zaragoza-Caballero and Velasco-de León 2003
†Epicauta subneglecta Wickham 1914
Tr. Lyttini Solier 1851
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Tr. Meloini Gyllenhal 1810
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Tr. Mylabrini Rafinesque 1815
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G. Mylabris Fabricius 1775
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†Mylabris deflorata von Heyden and von Heyden 1866
†Mylabris punctatissima Théobald 1935
G. Zonabris Harold 1879
Tr. Tetraonycini Böving and Craighead 1931
+
Invalid names: Eleticinae Wellman 1910 [empty]
Fm. Mordellidae Latreille 1802 [tumbling flower beetle]
+
Subfm. †Apotomourinae Bao et al. 2018
+
G. †Multispinus Bao et al. 2018
+
†Multispinus multispinosus Bao et al. 2018
†Multispinus parvus Bao 2020
G. Falsomordellistena Ermisch 1941
G. Glipostena Ermisch 1941
G. Mordellaria Ermisch 1950
Subfm. Mordellinae Latreille 1802
+
Tr. †Mediumiugini Peris and Ruzzier 2013
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Tr. Mordellini Latreille 1802
+
G. †Baltimorda Batelka et al. 2023
+
†Baltimorda friedrichi Perkovsky and Odnosum 2013
†Baltimorda scheelei Ermisch 1941
†Baltimorda succinea Bao et al. 2018
G. Tomoxia Costa 1854
Tr. Mordellistenini Ermisch 1941
+
G. †Baltistena Batelka et al. 2023
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†Baltistena amplicollis Ermisch 1941
†Baltistena antiqua Ermisch 1941
†Baltistena atronigra Batelka et al. 2023
†Baltistena aurata Batelka et al. 2023
†Baltistena brevispina Batelka et al. 2023
†Baltistena concava Batelka et al. 2023
†Baltistena goeckei Ermisch 1941
†Baltistena hoffeinsorum Batelka et al. 2023
†Baltistena korschefskyi Ermisch 1941
†Baltistena longistrigata Batelka et al. 2023
†Baltistena nigrispinata Batelka et al. 2024
†Baltistena sergeli Ermisch 1943
†Baltistena soror Ermisch 1941
†Baltistena ultima Batelka et al. 2023
Tr. Raynoldsiellini Franciscolo 1957
+
Tr. Reynoldsiellini Franciscolo 1957
+
G. †Petrimordella Batelka et al. 2023
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†Petrimordella indata Statz 1952
†Petrimordella inundata Wickham 1914
†Petrimordella lapidicola Wickham 1909
†Petrimordella nearctica Wickham 1914
†Petrimordella nigrapilosa Statz 1952
†Petrimordella oligocenica Nel 1985
†Petrimordella priscula Cockerell 1925
†Petrimordella protogaea Wickham 1914
†Petrimordella rasnitsyni Odnosum and Perkovsky 2016
†Petrimordella scudderiana Wickham 1914
†Petrimordella smithiana Wickham 1913
†Petrimordella stygia Wickham 1914
Subfm. †Praemordellinae Shchegoleva-Barovskaya 1929
+
G. †Bellimordella Liu et al. 2008
+
†Bellimordella capitulifera Liu et al. 2008
†Bellimordella longispina Liu et al. 2008
†Bellimordella robusta Liu et al. 2008
G. †Wuhua Wang and Zhang 2011
+
†Wuhua jurassica Wang and Zhang 2011
†Wuhua peregrina Bao et al. 2019
Invalid names: Ctenidiinae Franciscolo 1951 [empty], Praemordellidae Shchegoleva-Barovskaya 1929 [synonym]
Fm. Mycetophagidae Leach 1815 [hairy fungus beetle]
+
Subfm. Mycetophaginae Leach 1815
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Tr. Mycetophagini Leach 1815
+
G. Litargus Erichson 1846
+
Subg. Litargus (Litargosomus) Motschulsky 1858
Subg. Litargus (Litargus) Erichson 1846
Invalid names: Limniattagenus Poinar and Háva 2015 [synonym]
G. Mycetophagus Hellwig 1792
+
†Mycetophagus exterminatus Wickham 1913
†Mycetophagus willistoni Wickham 1913
Invalid names: Bergininae Leng 1920 [empty], Esarcinae Reitter 1882 [empty]
Fm. Mycteridae Oken 1843 [palm and flower beetle]
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Subfm. Eurypinae Thomson 1860
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G. †Glesoconomorphus Alekseev et al. 2019
+
†Glesoconomorphus ekaterinae Telnov et al. 2021
†Glesoconomorphus nachzehrer Alekseev et al. 2019
Invalid names: Lacconotinae Le Conte 1862 [synonym]
Invalid names: Hemipeplinae Lacordaire 1854 [empty]
Fm. Oedemeridae Latreille 1810 [false blister beetle]
+
Subfm. Calopodinae Costa 1852
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G. †Ditysparedrus Vitali and Legalov 2020
+
†Ditysparedrus gigas Vitali and Legalov 2020
†Ditysparedrus obscurus Vitali and Legalov 2020
†Ditysparedrus serixioides Vitali and Legalov 2020
†Ditysparedrus uniformis Vitali and Legalov 2020
Subfm. Oedemerinae Latreille 1810
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Tr. Ditylini Mulsant 1858
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Invalid names: Saperda absyrti Heer 1847 [nomen dubium]
Fm. Prostomidae Thomson 1859 [jugular-horned beetle]
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G. Prostomis Latreille 1825
G. †Vetuprostomis Engel and Grimaldi 2008
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†Vetuprostomis angularis Li and Cai 2022
†Vetuprostomis consimilis Engel and Grimaldi 2008
†Vetuprostomis gaoi Li and Cai 2022
Fm. Pyrochroidae Latreille 1807
+
Subfm. Pedilinae Lacordaire 1859
+
Subfm. †Waidelotinae Bukejs et al. 2019
+
Invalid names: Cretaceimelittomoides Vulcano and Pereira 1987 [nomen nudum], Cretaceimelittomoides cearensis Vulcano and Pereira 1987 [nomen nudum], Pilipalpinae Abdullah 1964 [empty], Tydessinae Nikitsky 1986 [empty]
Fm. Pythidae Solier 1834
+
Fm. Rhipiphoridae Gemminger 1870 [wedge shaped beetle]
+
G. Rhipiphorus Bosc 1792
Invalid names: Rhipidiinae Gerstaecker 1855 [empty], Rhipiphorinae Gemminger 1870 [empty]
Fm. Ripiphoridae Gemminger 1870 [wedge shaped beetle]
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Subfm. Ripidiinae Gerstaecker 1855
+
Tr. Eorhipidiini Iablokoff-Khnzorian 1986
+
G. †Olemehliella Batelka 2017
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†Olemehliella krali Batelka 2011
Invalid names: Olemehlia Batelka 2011 [replaced]
G. †Pauroripidius Kaupp and Nagel 2001
+
†Pauroripidius groehni Kaupp and Nagel 2001
†Pauroripidius pyrrholophus Dalman 1826
Tr. Ripidiini Gerstaecker 1855
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G. †Amberocula Batelka et al. 2018
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†Amberocula costata Batelka et al. 2018
†Amberocula fallax Batelka et al. 2018
†Amberocula muelleri Batelka et al. 2018
G. †Ripidinelia Batelka and Prokop 2019
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†Ripidinelia burmiticola Batelka and Prokop 2019
†Ripidinelia daiboyui Li and Cai 2024
G. †Ripilarva Batelka and Beutel 2024
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†Ripilarva kachinensis Batelka and Beutel 2024
†Ripilarva parabolica Batelka and Beutel 2024
Invalid names: Ripidius megalophus Dalman 1826 [nomen dubium]
Subfm. Ripiphorinae Gemminger 1870
+
Tr. Macrosiagonini von Heyden 1908
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Tr. Ripiphorini Gemminger 1870
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G. Ripiphorus Bosc 1792
+
†Ripiphorus meyeri von Heyden and von Heyden 1866
Invalid names: Myodites Latreille 1829 [synonym]
Invalid names: Hemirhipidiinae Heller 1921 [empty], Ptilophorinae Gerstaecker 1855 [empty]
Fm. Salpingidae Leach 1815 [narrow-waisted bark beetle]
+
Subfm. Inopeplinae Grouvelle 1908
+
Subfm. Prostominiinae Grouvelle 1914
+
Subfm. Salpinginae Leach 1815
+
Invalid names: Aegialitinae LeConte 1862 [empty], Dacoderinae LeConte 1862 [empty], Othniinae LeConte 1861 [empty]
Fm. Scraptiidae Gistel 1848 [false flower beetle]
+
Subfm. Anaspidinae Mulsant 1856
+
Tr. Anaspidini Mulsant 1856
+
G. Anaspis Geoffroy 1762
+
Subg. Anaspis (Silaria) Mulsant 1856
Subg. Anaspis (Spanisa) Emery 1876
†Anaspis longispina Ermisch 1941
Anaspis maculata Geoffroy 1785
†Anaspis parallela Ermisch 1941
Invalid names: Anaspis antica Guérin-Méneville 1838 [nomen nudum]
Subfm. Scraptiinae Gistel 1848
+
Tr. Scraptiini Gistel 1848
+
G. Scraptia Latreille 1807
+
†Scraptia inclusa Ermisch 1941
+
Invalid names: Pentaroides fischeri Yablokov-Khnzorian 1960 [synonym]
†Scraptia longelytrata Ermisch 1943
†Scraptia ovata Guérin-Méneville 1838
†Scraptia pseudofuscula Ermisch 1941
Invalid names: Pentaroides Yablokov-Khnzorian 1960 [synonym]
G. †Scraptiomima Medvedev 1969
+
†Scraptiomima brachycornis Medvedev 1969
†Scraptiomima minima Ponomarenko 1990
Fm. Stenotrachelidae Thomson 1859
+
Invalid names: Cephaloinae LeConte 1862 [empty], Nematoplinae LeConte 1862 [empty], Stenotrachelinae Thomson 1859 [empty], Stoliinae Nikitsky 1985 [empty]
Fm. Tenebrionidae Latreille 1802 [darkling beetle]
+
Subfm. Alleculinae Laporte 1840 [comb-clawed beetle]
+
Tr. Alleculini Laporte 1840
+
Subtr. Alleculina Laporte 1840
+
G. Allecula Fabricius 1801
G. Hymenalia Mulsant 1856
G. Hymenorus Mulsant 1852
G. †Jurallecula Medvedev 1969
G. Lobopoda Solier 1835
G. Oracula Novák 2019
G. Parahymenorus Campbell 1971
Subtr. Gonoderina Seidlitz 1896
+
G. Asiomira Dubrovina 1973
G. †Calcarocistela Nabozhenko 2015
G. Capnochroa Le Conte 1862
G. Gonodera Mulsant 1856
G. Isomira Mulsant 1856
G. Pseudocistela Crotch 1873
Subtr. Mycetocharina Gistel 1848
+
G. Mycetochara Guérin-Méneville 1827
G. †Mycetocharoides Schaufuss 1889
Tr. Cteniopodini Solier 1835
+
G. Cteniopinus Seidlitz 1896
Invalid names: Cisteleniae Latreille 1802 [replaced], Omophlini Mulsant 1856 [synonym]
Invalid names: Dascillus exiguus Zhang 1989 [nomen dubium]
Subfm. Blaptinae Leach 1815
+
Tr. Amphidorini LeConte 1862
+
G. Eleodes Eschscholtz 1829
+
Subg. Eleodes (Blapylis) Horn 1870
Subg. Eleodes (Cratidus) LeConte 1862
Subg. Eleodes (Eleodes) Eschscholtz 1829
Subg. Eleodes (Litheleodes) Blaisdell 1909
Subg. Eleodes (Melaneleodes) Blaisdell 1909
Subg. Eleodes (Xysta) Eschscholtz 1829
Tr. Opatrini Brullé 1832
+
Subtr. Opatrina Brullé 1832
+
G. Ephalus Le Conte 1862
G. Gonocephalum Chevrolat 1849
G. Opatrum Fabricius 1775
Subfm. Diaperinae Latreille 1802
+
Tr. Diaperini Latreille 1802
+
Subtr. Diaperina Latreille 1802
+
G. Ceropria Laporte and Brulle 1831
G. Liodema Horn 1870
G. Neomida Latreille 1829
G. Pentaphyllus Dejean 1821
G. Platydema Laporte and Brulle 1831
Invalid names: Adelina Le Conte 1862 [empty]
Tr. Gnathidiini Gieben 1921
+
Tr. Hypophlaeini Billberg 1820
+
Tr. Scaphidemini Reitter 1922
+
G. †Palaeobasanus Nabozhenko and Kirejtshuk 2020
+
†Palaeobasanus neli Nabozhenko and Kirejtshuk 2020
Subfm. Lagriinae Latreille 1825
+
Tr. Belopini Reitter 1917
+
G. Centorus Mulsant 1854
+
†Centorus inhumatus Hope 1842
Invalid names: Calcar Dejean 1821 [replaced]
Tr. †Gonialaenini Nabozhenko et al. 2019
+
Tr. Lagriini Latreille 1825
+
Tr. Prateini Aalbu et al. 2023
+
G. Lorelopsis Champion 1896
+
†Lorelopsis foraminosa Doyen and Poinar 1994
†Lorelopsis minutulis Doyen and Poinar 1994
Lorelopsis wolcotti Doyen 1993
G. Prateus LeConte 1862
+
†Prateus angulatus Doyen and Poinar 1994
Invalid names: Lorelus Sharp 1876 [synonym]
Subfm. Nilioninae Oken 1843 [false ladybird beetle]
+
Invalid names: Nilionini Thomson 1860 [invalid subgroup]
Subfm. Phrenapatinae Solier 1834
+
Tr. Penetini Lacordaire 1859
+
G. Scolytocaulus Fairmaire 1896
+
Scolytocaulus brevis Carter 1911
Invalid names: Platycilibe Carter 1911 [synonym]
Subfm. Pimeliinae Latreille 1802
+
Tr. Asidini Fleming 1821
+
G. Asida Latreille 1802
+
G. Micrasida Smith 2013
G. Pelecyphorus Solier 1836
+
Subg. Pelecyphorus (Plesiasida) Smith 2013
+
Invalid names: Pelecyphorus (Parasida) Casey 1912 [replaced]
Subg. Pelecyphorus (Stenosides) Solier 1836
+
Invalid names: Ologlyptus Lacordaire 1859 [objective synonym], Pactostoma Le Conte 1858 [synonym]
Tr. Coniontini Waterhouse 1858
+
G. Coniontis Eschscholtz 1829
+
Coniontis abdominalis LeConte 1859
+
Invalid names: Coniontis abdominalis caseyi Pierce 1954 [synonym], Coniontis abdominalis fragmans Pierce 1954 [synonym], Coniontis abdominalis labreae Pierce 1954 [synonym], Coniontis blissi Pierce 1954 [synonym], Coniontis pectoralis interrupta Pierce 1954 [synonym], Coniontis pectoralis paraelliptica Pierce 1954 [synonym], Coniontis rugosa Casey 1908 [synonym], Coniontis tristis Casey 1908 [synonym], Coniontis tristis asphalti Pierce 1954 [synonym], Coniontis tristis latigula Pierce 1954 [synonym]
Coniontis elliptica Casey 1884
Coniontis lamentabilis Blaisdell 1924
Coniontis pectoralis Casey 1908
Coniontis puncticollis LeConte 1851
†Coniontis remnans Pierce 1954
Coniontis robusta Horn 1870
Invalid names: Coniontis tristis alpha Pierce 1954 [synonym]
Tr. Nycteliini Solier 1834
+
Invalid names: Akidini Billberg 1820 [empty]
G. †Rhinohelaeites Haupt 1950
+
†Rhinohelaeites longipes Haupt 1950
†Rhinohelaeites punctatulus Haupt 1950
Subfm. Stenochiinae Kirby 1837
+
G. †Anthracohelops Haupt 1950
+
†Anthracohelops gigas Haupt 1950
†Anthracohelops minutus Haupt 1950
†Anthracohelops molassicus Heer 1883
†Anthracohelops wetteravicus von Heyden and von Heyden 1865
Tr. Cnodalonini Oken 1843
+
G. Nesocyrtosoma Marcuzzi 1976
+
†Nesocyrtosoma antiquum Kaszab and Schwaller 1984
†Nesocyrtosoma celadonum Doyen and Poinar 1994
†Nesocyrtosoma hadratum Doyen and Poinar 1994
†Nesocyrtosoma impensum Doyen and Poinar 1994
†Nesocyrtosoma minisculum Hopp and Ivie 2009
†Nesocyrtosoma phthanatum Doyen and Poinar 1994
Invalid names: Hesiodobates Kaszab and Schwaller 1984 [synonym]
Invalid names: Coelometopini Lacordaire 1859 [synonym]
G. †Eodromus Pongrácz 1935
+
†Eodromus helopoides Haupt 1950
†Eodromus parvus Haupt 1956
†Eodromus punctatostriatus Haupt 1950
†Eodromus punctatosulcatus Pongrácz 1935
Subfm. Tenebrioninae Latreille 1802
+
Tr. Alphitobiini Reitter 1917
+
Tr. Amarygmini Gistel 1848
+
Tr. Bolitophagini Kirby 1837
+
Tr. Eulabini Horn 1870
+
G. Eulabis Eschscholtz 1829
Tr. Helopini Latreille 1802
+
Subtr. Helopina Latreille 1802
+
G. †Cryptohelops Nabozhenko and Kirejtshuk 2014
G. Euboeus Boieldieu 1865
G. Helops Fabricius 1775
G. Stenohelops Reitter 1922
Tr. Melanimonini Seidlitz 1894
+
G. Melanimon Stevens 1829
+
†Melanimon veteratum von Heyden and von Heyden 1866
Invalid names: Microzoum Dejean 1834 [synonym]
Tr. Metaclisini Steiner 2016
+
Tr. Palorini Matthews 2003
+
Tr. Tenebrionini Latreille 1802
+
G. Tenebrio Linnaeus 1758
+
†Tenebrio effossus Germar 1837
†Tenebrio primigenius Scudder 1879
†Tenebrio senex Von Heyden 1859
Tr. Toxicini Oken 1843
+
Subtr. Dysantina Gebien 1922
+
G. Diceroderes Solier 1841
Invalid names: Eudysantina Bouchard et al. 2005 [synonym]
Invalid names: Titaenini Fauvel 1905 [empty], Ulus minutus Wickham 1914 [nomen dubium]
G. †Tenebrionites Cockerell 1920
+
†Tenebrionites alatus Cockerell 1927
†Tenebrionites anglicus Cockerell 1920
†Tenebrionites inclinans Cockerell 1925
Invalid names: Ancylochira oligostriata Piton and Théobald 1937 [nomen dubium], Tenebrio calculensis Scudder 1895 [nomen dubium]
Fm. Tetratomidae Billberg 1820 [polypore fungus beetle]
+
Subfm. Eustrophinae Gistel 1848
+
Tr. Eustrophini Gistel 1848
+
G. †Allostrophus Hsiao et al. 2018
+
†Allostrophus cretaceus Hsiao et al. 2018
†Allostrophus yangi Hsiao 2020
Tr. Holostrophini Nikitsky 1998
+
Subfm. Hallomeninae Gistel 1848
+
Subfm. Tetratominae Billberg 1820
+
Tr. †Archaeolupropini Nabozhenko et al. 2023
+
G. Tetratoma Fabricius 1790
+
Invalid names: Penthinae Lacordaire 1859 [empty], Piseninae Miyatake 1960 [empty]
Fm. Zopheridae Solier 1834 [ironclad beetle]
+
Subfm. Colydiinae Billberg 1820
+
G. †Bifurderum Cheng et al. 2020
+
†Bifurderum donoghuei Cheng et al. 2020
†Bifurderum minutum Cheng et al. 2020
Tr. Synchitini Erichson 1845
+
G. Namunaria Reitter 1882
G. †Paleoendeitoma Deng et al. 2017
+
†Paleoendeitoma antennata Deng et al. 2017
†Paleoendeitoma buryi Háva 2019
†Paleoendeitoma minuta Deng et al. 2017
†Paleoendeitoma tuberculata Bullis 2020
G. Synchita Hellwig 1792
+
†Synchita andrisbukejsi Alekseev 2022
Invalid names: Cicones Curtis 1827 [synonym]
G. Xylolaemus Redtenbacher 1857
+
†Xylolaemus legalovi Alekseev and Bukejs 2016
†Xylolaemus richardklebsi Alekseev and Bukejs 2016
†Xylolaemus sakhnovi Alekseev and Lord 2014
Subfm. Zopherinae Solier 1834
+
Tr. Pycnomerini Erichson 1845
+
G. Pycnomerus Erichson 1842
+
†Pycnomerus agtsteinicus Bukejs et al. 2019
†Pycnomerus mahanatoa Porch and Greig 2020
†Pycnomerus prebblei Porch and Smith 2017
†Pycnomerus rairua Porch and Greig 2020
†Pycnomerus raivavae Porch and Greig 2020
†Pycnomerus rimatara Porch and Smith 2017
†Pycnomerus simukovi Alekseev 2015
†Pycnomerus taralewisae Porch and Greig 2020
Pycnomerus terebrans Olivier 1790
†Pycnomerus vavitu Porch and Greig 2020
Tr. Usechini Horn 1867
+
Tr. Zopherini Solier 1834
+
G. Nosoderma Guerin-Meneville 1838
Invalid names: Nosodermini Casey 1907 [synonym]
Tr. †Zopheromimini Alekseev and Nabozhenko 2023
+
G. †Zopheromimus Alekseev and Nabozhenko 2023
+
†Zopheromimus auriborussiensis Alekseev and Nabozhenko 2023
Invalid names: Colydiidae Billberg 1820 [replaced]
Invalid names: Afreminae Levey 1985 [empty], Archeocrypticidae Kaszab 1964 [empty], Asiamordella Hong 2002 [nomen nudum], Chalcodryidae Watt 1974 [empty], Cossyphodinae Wasmann 1899 [empty], Eurygenius wickhami Cockerell 1917 [nomen dubium], Fushunicucujus Hong 2002 [nomen nudum], Lagrioididae Abdullah and Abdullah 1968 [empty], Perimylopidae St. George 1939 [empty], Pterogeniidae Crowson 1953 [empty], Trachelostenidae Lacordaire 1859 [empty], Trictenotomidae Blanchard 1845 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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Source: c = class, p = phylum | |||||
References: Kiessling 2004, Bush and Bambach 2015 |
Age range
Maximum range based only on fossils: base of the Callovian to the top of the Holocene or 165.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 154.8 Ma
Minimum age of oldest fossil (stem group age): 154.8 Ma
Collections (228 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
---|---|---|---|
Callovian - Oxfordian | China (Nei Mongol) | Archaeoripiphorus nuwa (113360 223890) Tenebrionoidea indet. (112664) Wuhua peregrina, Wuhua jurassica (113371) | |
Callovian - Oxfordian | Kazakhstan (Chimkent) | Jurallecula grossa (106101) Praemordella martynovi (114692) | |
Hauterivian | China (Hebei) | Glypta qingshilaensis (107218) | |
Barremian | China (Hebei) | Glypta qingshilaensis, Glypta longa (155164) | |
Early/Lower Barremian | Lebanon | Elodophthalmus maksoudae, Elodophthalmus harmonicus, Elodophthalmus gracilis, Camelomorpha longicervix (122471) | |
Late/Upper Barremian | China (Liaoning) | Alphitopsis initialis (110825) Liaoximordella hongi (156322) Mirimordella gracilicruralis, Bellimordella longispina, Bellimordella capitulifera, Bellimordella robusta (111710) Platycteniopus diversoculatus, Calcarocistela kirejtshuki (173379) | |
Early/Lower Aptian | Russian Federation (Chita) | Scraptiomima minima (119499) | |
Aptian | Russian Federation (Buryatia) | Scraptiomima brachycornis (105536) | |
Late/Upper Aptian | Brazil (Ceará) | Cretaceimelittomoides cearensis (125511) Tenebrionidae indet. (122557) | |
Late/Upper Aptian | China (Beijing Shi) | Cretanaspis lushangfenensis (140479) | |
Late/Upper Aptian | Australia (Victoria) | Mordellidae indet. (77192) | |
Early/Lower Albian | Spain | Arra legalovi (109438) | |
Late/Upper Albian | Spain (Aragón) | Arra legalovi (91626) | |
Late/Upper Albian | France (Poitou-Charentes) | Paleoripiphorus deploegi, Synchrotronia idinineteena (99114) | |
Late/Upper Albian | Spain (Álava) | Mediumiuga sinespinis, Darwinylus marcosi (108769) | |
Cenomanian | Russian Federation | Yakutia sukachevae (226224) | |
Cenomanian | France (Alpes-de-Haute- Provence) | Macrosiagon ebboi (123441) | |
Early/Lower Cenomanian | Myanmar | Allostrophus yangi (210680) Amberocula muelleri, Amberocula fallax, Paleoripiphorus deploegi (184692) Burmitoma nalae, Plesiotoma alissae, Amberocula muelleri, Amberocula costata, Amberocula fallax (184841) Cretoprostominia myanmarensis (227023) Ditysparedrus obscurus, Ditysparedrus uniformis, Ditysparedrus gigas (182624) Ditysparedrus serixioides, Sparedrus archaicus (204334) Eurygenius wickhami, Myodites burmiticus, Cryphalites rugosissmus (123020) Flabellotoma heidiae, Paleoripiphorus deploegi, Ripiphoridae tringulin B, Vetuprostomis consimilis (111688) Glyphonotum hsiaoi, Primaevomordellida burmitina, Angimordella burmitina, Multispinus multispinosus, Multispinus parvus, Apotomoura fortiscrura, Kulindrobor enigmaticus, Gryzmalia wukong, Praezolodinus pilosus, Longicrusa jaracimrmani, Homalenchodes jarzembowskii, Protoripidius burmiticus, Ripidinelia daiboyui, Cretosynstrophus archaicus, Vetuprostomis gaoi, Vetuprostomis angularis, Bifurderum minutum, Bifurderum donoghuei, Bulasconotus carinisternus, Coslonatus rasnitsyni (161726) Kulindrobor magnus (184691) Lemodicarmenia olmedoae (187030) Macratriinae indet. (201136) Melanosiagon serraticornis, Eodrias mandibularis, Ripidinelia burmiticola, Ripidinelia sp. (223712) Paleoendeitoma buryi (205556) Paleoendeitoma tuberculata (218269) Prototrichalus jingpo, Prototrichalus sp. A, Prototrichalus sp. B (189783) Prototrichalus milleri, Prototrichalus meiyingae (216411) Prototrichalus milleri, Prototrichalus sepronai, Prototrichalus meiyingae (208365) Ripidiinae indet. (123053) Ripidiinae indet., Ripiphoridae indet. (178121) Ripilarva kachinensis (234239) Spinotoma ruicheni (178159) Thescelostrophus cretaceus, Allostrophus cretaceus, Cretomysteria burmanica, Paleoendeitoma minuta, Paleoendeitoma antennata (165561) | |
Early/Lower Cenomanian | Myanmar (Kachin) | Stegastochlidus saraemcheana (104053) | |
Turonian | USA (New Jersey) | Archaeserropalpus cretaceus, Pseudocuphosis tristis (110895) |