| Basic info | Taxonomic history | Classification | Included Taxa |
| Morphology | Ecology and taphonomy | External Literature Search | Age range and collections |
Taxonomy
Bibionoidea was named by Fleming (1821). It is extant. Its type is Bibio.
It was assigned to Eudiptera by Hong and Wang (1985); and to Bibionomorpha by Yeates et al. (2007), Poinar (2010).
It was assigned to Eudiptera by Hong and Wang (1985); and to Bibionomorpha by Yeates et al. (2007), Poinar (2010).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1821 | Bibionoidea Fleming |
| 1985 | Bibionoidea Hong and Wang p. 17 |
| 2007 | Bibionoidea Yeates et al. |
| 2010 | Bibionoidea Poinar |
Is something missing? Join the Paleobiology Database and enter the data
|
|
If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Superfm. Bibionoidea Fleming 1821
show all | hide all
Fm. Bibionidae Fleming 1821 [march fly]
hide
Subfm. Bibioninae Fleming 1821
+
Tr. Dilophini Pinto and Amorim 2000
+
G. Dilophus Meigen 1803
+
†Dilophus anasiformis Dürrenfeldt 1968
†Dilophus andrewrossi Nel et al. 2019
†Dilophus campbelli Harris 1983
†Dilophus cerestensis Nel et al. 2022
†Dilophus crassicornis Skartveit 2009
†Dilophus duerrenfeldti Skartveit et al. 2025
+
Invalid names: Bibio tenuis Dürrenfeldt 1968 [replaced]
†Dilophus flavipes Skartveit et al. 2025
†Dilophus idanos Fitzgerald and Greenwalt 2022
†Dilophus krantzii von Heyden 1870
†Dilophus luteipennis Theobald 1937
†Dilophus magnus Dürrenfeldt 1968
†Dilophus matilei Waller et al. 2000
†Dilophus palaeofebrilis Skartveit 2009
†Dilophus pinguis Heer 1849
+
Invalid names: Bibio pinguis oeningensis Heer 1849 [synonym], Bibio pinguis radobojanus Heer 1849 [objective synonym]
†Dilophus plionanus Skartveit et al. 2025
†Dilophus priscus Loew 1850
†Dilophus pumilio Skartveit and Pika 2014
†Dilophus succineus Skartveit 2009
Invalid names: Dilophus deletus Von Heyden 1859 [nomen dubium]
Subfm. †Burmahesperininae Ševčík et al. 2021
+
G. †Burmahesperinus Ševčík et al. 2021
+
†Burmahesperinus antennatus Ševčík et al. 2021
†Burmahesperinus conicus Ševčík et al. 2021
†Burmahesperinus pedicellatus Ševčík et al. 2021
Subfm. †Cretobibioninae Skartveit and Ansorge 2020
+
G. †Cretobibio Skartveit and Ansorge 2020
+
†Cretobibio burmiticus Skartveit 2023
†Cretobibio montsecensis Skartveit and Ansorge 2020
†Cretobibio rasnitsyni Lukashevich et al. 2021
Subfm. Hesperininae Walker 1848
+
G. Hesperinus Walker 1848
+
†Hesperinus electrus Skartveit 2009
†Hesperinus heeri von Heyden and von Heyden 1865
+
Invalid names: Penthetria rottensis Statz 1943 [synonym], Penthetria scita Statz 1943 [synonym], Plecia elegantula Meunier 1915 [synonym]
†Hesperinus hyalopterus Skartveit 2009
†Hesperinus macroculatus Skartveit 2009
G. Penthetria Meigen 1803
+
†Penthetria abacula Lewis 1971
†Penthetria aestimata Statz 1943
†Penthetria alderensis Lewis 1971
†Penthetria anthracina Heer 1849
†Penthetria beskonakensis Skartveit and Nel 2012
†Penthetria blanchardi Oustalet 1870
†Penthetria bohemica Novák 1878
†Penthetria brevicollis Heer 1849
†Penthetria carbonum von Heyden and von Heyden 1865
†Penthetria cimicoides Heer 1849
†Penthetria claripennis Theobald 1937
+
Invalid names: Plecia difficilis Theobald 1937 [synonym], Plecia graciosa Theobald 1937 [synonym], Plecia varia Theobald 1937 [synonym], Plecia variegata Theobald 1937 [synonym]
†Penthetria creedensis James 1938
†Penthetria egerana Novák 1878
†Penthetria fryi Rice 1959
†Penthetria fuchsi Handlirsch 1907
+
Invalid names: Penthetria marinosi Bachmayer et al. 1971 [objective synonym]
†Penthetria gigantea Theobald 1937
+
Invalid names: Plecia crassiventris Theobald 1937 [synonym], Plecia luteipennis Theobald 1937 [synonym], Plecia miegi Theobald 1937 [synonym]
†Penthetria globularis Oustalet 1870
†Penthetria gracilis Unger 1841
†Penthetria graciliventris Theobald 1937
†Penthetria immutabilis Melander 1949
†Penthetria imperialis Novák 1878
†Penthetria incerta Oustalet 1870
†Penthetria integroneura Skartveit 2009
†Penthetria intermedia Scudder 1892
†Penthetria longa Heer 1849
†Penthetria longifurca Melander 1949
†Penthetria luberonica Skartveit and Nel 2017
†Penthetria lugens Oustalet 1870
†Penthetria montanaregis Skartveit 2009
†Penthetria murchisonii Unger 1841
†Penthetria nervisinuata Theobald 1937
†Penthetria nigra Statz 1943
†Penthetria parschlugiana Heer 1849
†Penthetria rubescens Oustalet 1870
†Penthetria rubiensis Lewis 1971
†Penthetria subterranea Theobald 1937
†Penthetria togoensis Fujiyama and Iwao 1974
†Penthetria tropica Statz 1943
Penthetria velutina Loew 1858
†Penthetria whipsawensis Rice 1959
Invalid names: Bibio alacris Oustalet 1870 [nomen dubium], Bibio gigas Oustalet 1870 [nomen dubium], Bibio macer Oustalet 1870 [nomen dubium], Bibio minor Oustalet 1870 [nomen dubium], Bibiopsis Heer 1849 [synonym], Mycetophaetus Scudder 1892 [synonym], Protomyia Heer 1849 [synonym]
Invalid names: Longicornia Hong 2002 [nomen nudum], Penthetriinae Rohdendorf 1946 [synonym]
Subfm. Pleciinae Duda 1930
+
G. Plecia Wiedemann 1823 [love bug]
+
†Plecia acourti Cockerell 1921
†Plecia aculeolata Zhang 1989
†Plecia akerionana Fitzgerald 1999
†Plecia amagua Cockerell 1925
†Plecia angustipennis Handlirsch 1910
†Plecia angustiventris Theobald 1937
†Plecia assonensis Gentilini 1993
†Plecia avus Handlirsch 1910
†Plecia axeliana Cockerell 1915
†Plecia baglii Gentilini 1991
†Plecia bivalvula Zhang 1993
†Plecia borussica Meunier 1904
†Plecia brunneipennis Piton 1939
†Plecia brunniptera Skartveit 2009
†Plecia bucklandi Heer 1849
+
Invalid names: Penthetria edwardsi Oustalet 1870 [synonym], Plecia dilatativentris Theobald 1937 [synonym], Protomyia elegans Heer 1856 [synonym], Protomyia gracilis Heer 1856 [synonym]
†Plecia cairnesi Rice 1959
†Plecia canadensis Handlirsch 1910
†Plecia capitata Zhang et al. 1994
†Plecia castellaroi Gentilini 1991
†Plecia chapuisii Oustalet 1872
†Plecia clavifemur Skartveit 2009
†Plecia collossea von Heyden and von Heyden 1865
+
Invalid names: Plecia imperialis Statz 1943 [synonym], Plecia philippi Statz 1943 [synonym], Plecia superba Meunier 1914 [synonym]
†Plecia decapitata Cockerell 1917
†Plecia dejecta Scudder 1890
†Plecia diatoma Zhang 1989
†Plecia dilatata Handlirsch 1910
†Plecia dubia Germar 1837
+
Invalid names: Helophilus primarius Germar 1837 [synonym], Plecia satyrus Statz 1943 [synonym], Protomyia abava von Heyden and von Heyden 1865 [synonym]
†Plecia dumasi Theobald 1937
†Plecia elatior Handlirsch 1910
†Plecia exigua Statz 1943
†Plecia explanata Cockerell 1917
†Plecia exposititia von Heyden 1865
†Plecia foersteri Theobald 1937
+
Invalid names: Plecia oviformis Theobald 1937 [synonym], Plecia splendida Theobald 1937 [synonym]
†Plecia fumosa Zhang et al. 1994
†Plecia gracilenta von Heyden and von Heyden 1865
†Plecia gracillima Förster 1891
†Plecia gradata Melander 1949
†Plecia grossa von Heyden and von Heyden 1865
†Plecia hilaris Heer 1849
†Plecia hypogaea von Heyden and von Heyden 1865
+
Invalid names: Plecia pulchella Meunier 1915 [synonym], Protomyia antenata von Heyden and von Heyden 1865 [synonym], Protomyia elongata von Heyden and von Heyden 1865 [synonym], Protomyia expositia von Heyden and von Heyden 1865 [synonym], Protomyia grandaeva von Heyden and von Heyden 1865 [synonym]
†Plecia inflata Oustalet 1870
†Plecia inflata Hull 1960
†Plecia intima Fujiyama and Iwao 1974
†Plecia jucunda Heer 1849
+
†Protomyia jucunda parschlugiana Heer 1849
Invalid names: Protomyia affinis Heer 1849 [synonym]
†Plecia kanetakii Fujiyama 1970
†Plecia kelownaensis Rice 1959
†Plecia kudiella Cockerell 1925
†Plecia kuznetzovi Cockerell 1925
†Plecia larguensis Gentilini 1993
†Plecia livida Heer 1849
+
Invalid names: Plecia brevithoracis Theobald 1937 [synonym], Plecia cerestensis Theobald 1937 [synonym], Plecia constricta Theobald 1937 [synonym], Plecia flichei Theobald 1937 [synonym], Plecia fusciventris Theobald 1937 [synonym], Plecia goreti Theobald 1937 [synonym], Plecia maculitarsis Theobald 1937 [synonym], Plecia maculiventris Theobald 1937 [synonym], Protomyia brevipennis Heer 1856 [synonym]
†Plecia luctuosa von Heyden and von Heyden 1865
†Plecia ludongensis Hong and Wang 1985
†Plecia lugubris Heer 1849
†Plecia luteola von Heyden and von Heyden 1865
†Plecia lygaeoides Heer 1849
+
Invalid names: Bibio detractus Theobald 1937 [synonym], Plecia acutiventris Theobald 1937 [synonym], Plecia guinieri Theobald 1937 [synonym], Protomyia latipennis Heer 1849 [synonym]
†Plecia macilenta Skartveit and Pika 2014
†Plecia macrocephala von Heyden and von Heyden 1865
+
Invalid names: Protomyia schineri von Heyden and von Heyden 1865 [synonym]
†Plecia maimensis Gentilini 1993
†Plecia melanderi Cockerell 1911
†Plecia minutula Rice 1959
†Plecia morio Heer 1849
†Plecia myersi Peterson 1975
†Plecia nana Handlirsch 1910
†Plecia obsitula Cockerell 1925
†Plecia orycta Melander 1949
†Plecia oustaleti Brongniart 1876
†Plecia parisiensis Gee et al. 2001
†Plecia pealei Scudder 1890
†Plecia pennata Statz 1943
†Plecia pictipennis Handlirsch 1910
+
Invalid names: Penthetria lambei Handlirsch 1910 [synonym], Penthetria ovalis Handlirsch 1910 [synonym], Penthetria separanda Handlirsch 1910 [synonym]
†Plecia pinguis von Heyden and von Heyden 1865
+
Invalid names: Bibio brachypteroides Meunier 1915 [synonym], Plecia vulpina Statz 1943 [synonym], Protomyia lapidaria von Heyden and von Heyden 1865 [synonym]
†Plecia pisaurensis Gentilini 1991
†Plecia platoptera Zhang 1993
†Plecia platyptera Handlirsch 1910
†Plecia prisca Meunier 1899
†Plecia pristina Hardy 1971
†Plecia proserpina von Heyden and von Heyden 1865
†Plecia pulchra Handlirsch 1910
†Plecia quaesita Novák 1878
†Plecia rectivenia Zhang 1989
†Plecia redempta Cockerell 1925
†Plecia reducta Handlirsch 1910
†Plecia refracta Cockerell 1925
†Plecia rhenana von Heyden and von Heyden 1865
†Plecia rhodopterina Cockerell 1925
†Plecia similkameena Scudder 1879
†Plecia sluiteri Meunier 1917
†Plecia solaris Zhang 1989
†Plecia spinula Zhang 1989
†Plecia sturmi Statz 1943
†Plecia stygia von Heyden and von Heyden 1865
+
Invalid names: Protomyia veterana von Heyden and von Heyden 1865 [synonym]
†Plecia tessella Melander 1949
†Plecia theobaldi Skartveit and Nel 2017
+
Invalid names: Bibio nigripennis Theobald 1937 [replaced]
†Plecia transitoria Handlirsch 1910
+
Invalid names: Penthetria falcatula Handlirsch 1910 [synonym], Penthetria fragmentum Handlirsch 1910 [synonym]
†Plecia tulameenensis Rice 1959
†Plecia undans Zeuner 1941
†Plecia vergnei Piton 1935
†Plecia villosa Zhang 1989
†Plecia volgeri Von Heyden 1859
†Plecia vulcania Zhang 1989
†Plecia winchesteri Cockerell 1917
†Plecia winnertzi von Heyden and von Heyden 1865
†Plecia woodruffi Cockerell 1916
Invalid names: Bibio cylindratus Oustalet 1870 [nomen dubium], Clothonopsis Hong and Wang 1987 [nomen dubium], Clothonopsis miocenica Hong and Wang 1987 [nomen dubium], Epiplecia Giard 1879 [synonym], Plecia joannis Oustalet 1870 [nomen dubium], Plecia larteti Oustalet 1870 [nomen dubium], Plecia longipennis Oustalet 1870 [nomen dubium], Plecia painvini Meunier 1915 [nomen dubium], Plecia pallida Oustalet 1870 [nomen dubium], Plecia retracta Theobald 1937 [nomen dubium], Plecia rubescens Oustalet 1870 [nomen dubium], Plecia sauvagei Oustalet 1870 [nomen dubium]
Invalid names: Cascopleciinae Poinar 2010 [synonym], Fushunoplecia Hong 2002 [nomen nudum]
Invalid names: Bibio elongatus Heer 1849 [nomen dubium], Bibio incrassatus Heer 1849 [nomen dubium], Bibio linearis Heer 1849 [nomen dubium], Bibio robustus Oustalet 1870 [nomen dubium], Protomyia adusta Oustalet 1870 [nomen dubium], Protomyia formicoides Oustalet 1870 [nomen dubium], Protomyia fusca Oustalet 1870 [nomen dubium]
G. †Prohousea Jarzembowski 1996
hide
†Prohousea pachygaster Handlirsch 1939
Invalid names: Pleciomyia Handlirsch 1939 [replaced]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
|
|
||||
|
|||||
|
|
||||
| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Berriasian to the top of the Late/Upper Pleistocene or 139.10000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 137.05 Ma
Minimum age of oldest fossil (stem group age): 137.05 Ma
Collections (198 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Berriasian | United Kingdom (England) | Pleciomyia pachygaster (153649) | |
| Early/Lower Barremian | Spain | Cretobibio montsecensis (118913) | |
| Late/Upper Barremian | China (Hebei) | Lichnoplecia kovalevi (107001) | |
| Aptian | China (Shandong) | Bibionoidea indet. (154892) | |
| Late/Upper Aptian | Brazil (Ceará) | Bibionidae indet. (122352) Cretobibio rasnitsyni (217013) | |
| Albian | Jordan | Bibionidae indet. (124847) | |
| Early/Lower Cenomanian | Myanmar (Kachin) | Cascoplecia insolitis (104053) | |
| Early/Lower Cenomanian | Myanmar | Burmahesperinus sp., Burmahesperinus antennatus, Burmahesperinus conicus, Burmahesperinus pedicellatus (187929) Cretobibio burmiticus, Protodilophus semispinosus (133102) Protopenthetria skartveiti, Cretpenthetria burmensis (161726) | |
| Turonian | Botswana | Bibionidae indet., Bibiodes sp. (123430) | |
| Late/Upper Campanian | Canada (Manitoba) | Plecia myersi (122354) | |
| Selandian | France (Auvergne) | Bibio sp. (113685 123463) | |
| Thanetian | United Kingdom (Scotland) | Plecia undans (113632) | |
| Wasatchian | USA (Wyoming) | Plecia pealei (122174 122192 122193 187759) | |
| Ypresian | China (Liaoning) | Longicornia tenuis, Fushunoplecia eocenica (124820) | |
| Ypresian | USA (Colorado) | Plecia winchesteri (105004) | |
| Ypresian | France (Oise) | Plecia parisiensis (123850) Plecia sp. (68174) | |
| Ypresian | Italy | Bibio sereri (152649) | |
| Ypresian | Canada (British Columbia) | Penthetria angustipennis, Penthetria reducta, Penthetria fragmentum, Penthetria curtula, Penthetria dilatata, Penthetria platyptera, Plecia transitoria (119857) Penthetria fryi, Plecia intermedia, Plecia curtula, Plecia reducta, Plecia canadensis, Plecia avus (120981) Penthetria fryi, Plecia kelownaensis, Plecia avus, Plecia transitoria, Plecia pictipennis (120934) Penthetria fryi, Plecia minutula, Plecia curtula, Plecia tulameenensis, Plecia reducta, Plecia similkameena, Plecia canadensis, Plecia transitoria, Plecia pictipennis (120943) Penthetria fryi, Plecia tulameenensis, Plecia reducta, Plecia avus, Plecia pictipennis (120940) Penthetria pictipennis (119896) Penthetria pictipennis, Penthetria pulchra, Plecia curtula (119863) Penthetria pulla, Penthetria brevipes, Penthetria pictipennis, Penthetria elatior, Penthetria falcatula, Penthetria nana, Penthetria separanda, Penthetria avunculus, Penthetria avus, Penthetria lambei, Penthetria ovalis, Penthetria transitoria, Penthetria canadensis, Plecia reducta (119864) Penthetria similkameena, Plecia canadensis, Plecia transitoria (112781) Penthetria whipsawensis (120624) Plecia angustipennis, Plecia canadensis (120988) Plecia avus (121037) Plecia cairnesi (120992) Plecia cairnesi, Plecia reducta, Plecia avus, Plecia pictipennis (120991) Plecia curtula (120989 120990) Plecia dilatata (121041) Plecia kelownaensis (119464) Plecia pulchra (121046) Plecia reducta, Plecia pictipennis (121009) Plecia transitoria (121040) Plecia tulameenensis, Plecia similkameena, Plecia canadensis, Plecia avus, Plecia transitoria (119895) Pleciinae indet. (31677) | |
| Waipawan - Bortonian | New Zealand | Dilophus campbelli (119708) | |
| Bridgerian | USA (Wyoming) | Plecia dejecta (122090) Plecia sp. (21851) | |
| Bridgerian | USA (Colorado) | Plecia rhodopterina (122746) Plecia winchesteri (124625) | |
| Bridgerian | USA (Utah) | Plecia woodruffi (124691) | |
| Lutetian | USA (Montana) | Bibiodes kishenehnensis (117614) Dilophus idanos (135666) | |
| Uintan | USA (Colorado) | Plecia akerionana (138790) | |
| Late/Upper Eocene | France (Gard) | Plecia longiventris, Plecia angustiventris, Plecia foersteri, Plecia splendida, Plecia dumasi, Bibio conformans (122406) | |
| Late/Upper Eocene | France | Bibio celasensis (184102) | |
| Priabonian | Russian Federation (Kaliningrad) | Bibio sp. (124201) Dilophus sp., Penthetria montanaregis, Plecia clavifemur, Plecia hoffeinsorum, Dilophus crassicornis, Bibio succineus (217198) Penthetria montanaregis (123420) Plecia borussica, Plecia prisca (123927) Plecia sp. 1, Plecia sp. 2, Plecia sp. 3, Dilophus sp., Penthetria montanaregis, Plecia brunniptera, Plecia clavifemur, Plecia prisca, Bibiodes nanus, Dilophus crassicornis, Dilophus succineus (124517) | |
| Priabonian | Ukraine | Plecia hoffeinsorum (217199) | |
| Priabonian | Russian Federation | Penthetria sp., Hesperinus electrus, Hesperinus hyalopterus, Hesperinus macroculatus, Penthetria integroneura, Plecia brunniptera, Plecia clavifemur, Plecia hoffeinsorum, Plecia prisca, Plecia tenuicornis, Bibiodes balticus, Dilophus crassicornis, Dilophus palaeofebrilis (123419) | |
| Priabonian | United Kingdom (England) | Bibio gurnetensis, Bibio oligocenus, Bibiodites confluens, Protoberis obliteratus, Bibio oblitus, Bibio extremus, Dilophus andrewrossi (123959) Penthetria sp., Bibio oligocenus, Bibiodes confluens, Dilophus andrewrossi (123960) Plecia acourti (21976) | |
| Priabonian | France (Gard) | Bibio celasensis, Bibio obtusa (122405) Dilophus sp., Bibio celasensis, Penthetria longiventris, Penthetria graciliventris (144529) | |
| Priabonian | France | Bibionidae indet., Plecia longiventris, Plecia superba, Plecia angustiventris, Plecia foersteri, Plecia longa, Plecia splendida, Plecia graciliventris, Bibio celasensis, Bibio tenuiapacalis, Bibio elongatipennis, Bibio macerata, Plecia varia (113709) | |
| Priabonian | Poland | Dilophus priscus (109625) | |
| Chadronian | USA (Colorado) | Bibio cockerelli, Bibio jamesi, Bibio podager (124681) Bibio cockerelli, Bibiodes intermedia (106382) Bibio wickhami (124680) Bibio wickhami, Bibio cockerelli, Bibio explanatus, Bibio vulcanius (122975) Bibio wickhami, Bibio cockerelli, Bibio vetus (106381) Bibio wickhami, Bibiodes intermedia, Plecia gradata (106379) Bibiodes intermedia (106380) Mycetophaetus intermedius (106384) Plecia axeliana, Bibio wickhami (113749) Plecia decapitata (122977) Plecia explanata, Bibio wickhami, Bibio cockerelli, Bibio vetus, Hesperinus axelianus, Hesperinus immutabilis, Hesperinus melanderi, Penthetria longifurca, Plecia orycta, Plecia tessella, Bibio capnodes, Bibio excurvatus, Bibio jamesi, Bibio vulcanius (124624) Plecia melanderi (144782) | |
| Early/Lower Oligocene | France (Auvergne Region) | Plecia splendida, Dichaneurum infossum, Dichaneurum primaevum (31767) | |
| Early/Lower Oligocene | France | Plecia bucklandi, Plecia grossa, Plecia exposititia (121474) Plecia bucklandi, Plecia rhenana, Plecia luctuosa (122408) Plecia rhenana, Plecia grossa, Plecia lapidaria, Plecia gracillima, Plecia lygaeoides (121476) Plecia rubescens (121475) Plecia sp. 1, Plecia sp. 2, Plecia rhenana, Plecia pallida, Plecia stygia, Plecia grossa (121473) | |
| MP 23 | France | Bibionidae indet., Bibio sp., Plecia sp., Plecia bucklandi, Plecia splendida, Bibio painvini, Plecia livida, Plecia acutiventris, Bibio inflexus, Plecia flichei, Plecia fusciventris, Plecia guinieri, Plecia goreti, Plecia brevithoracis, Plecia cerestensis, Plecia maculitarsis, Plecia maculiventris, Bibio detractus (122535) Bibionidae indet., Plecia sp., Plecia lugens, Plecia nervisinuata, Plecia constricta, Plecia goreti, Plecia maculiventris (122536) Plecia livida, Penthetria nervisinuata, Bibio major, Dilophus cerestensis (157327) | |
| Rupelian | France (Alpes-de-Haute) | Dilophus sp., Hesperinus sp., Plecia livida, Plecia maimensis, Plecia assonensis, Plecia larguensis (164936) Plecia conica, Plecia maimensis, Plecia assonensis, Plecia larguensis (184087) | |
| Rupelian | France (Auvergne) | Bibio gigas, Protomyia adusta, Protomyia globularis (152488) Bibionidae indet., Bibio edwardsii, Plecia major, Bibio gigas, Protomyia joannis (152487) Bibionidae indet., Plecia major, Bibio gigas, Bibio robustus, Bibio lartetii, Protomyia incerta (152491) Bibionidae indet., Plecia pallida, Protomyia longa, Protomyia lugens, Protomyia inflata, Bibio edwardsii, Plecia major, Bibio marginatus, Bibio macer, Bibio alacris, Bibio robustus, Bibio cylindratus, Bibio minor, Bibio lartetii, Protomyia longipennis, Protomyia joannis, Protomyia fusca, Protomyia adusta, Protomyia sauvagei, Protomyia blanchardi, Protomyia rubescens, Protomyia formicoides, Protomyia incerta, Bibio obsoletus (127630) | |
| Rupelian | USA (Montana) | Plecia sp., Plecia inflata (132747) | |
| Rupelian | France | Dilophus sp., Protomyia oustaleti (139949) Plecia conica (184088) Plecia livida (184089) Plecia lygaeoides, Plecia livida (184090) Plecia maimensis, Plecia assonensis, Plecia larguensis (156997) | |
| Rupelian | Germany | Bibionidae indet., Plecia sp., Bibiodes sp., Dilophus sp., Plecia lapidaria, Plecia foersteri, Plecia splendida, Bibio baumbergeri, Bibio parenti, Bibio dislocata, Bibio consolida, Plecia gigantea, Plecia crassiventris, Plecia lugens, Plecia inflata, Plecia claripennis, Plecia macroventris, Plecia oviformis, Plecia conica, Plecia miegi, Plecia luteipennis, Plecia subterranea, Plecia variegata, Plecia varia, Plecia difficilis, Plecia graciosa, Plecia rubescens (122407) | |
| Oligocene | France | Plecia haennii (157308) | |
| Whitneyan | USA (Montana) | Bibionidae indet., Bibio sp., Plecia sp., Hesperinus sp., Penthetria sp. (120623) Penthetria rubiensis, Penthetria abacula, Penthetria alderensis (120622) | |
| Chattian | Germany (North Rhine-Westphalia) | Dilophus sp. 1, Penthetria sp., Penthetria sp 2, Plecia grossa, Plecia hypogaea, Plecia grandaeva, Plecia philippi, Plecia satyrus, Plecia imperialis, Plecia pennata, Plecia sturmi, Plecia exigua, Plecia vulpina, Penthetria nigra, Penthetria tropica, Penthetria aestimata, Penthetria rottensis, Penthetria scita, Bibio roehli, Bibio comosella, Bibio germari, Bibio rottensis, Bibio vulpina, Bibio aerosa, Bibio spadicea, Bibio auripes, Bibio compta, Plecia sp. 4, Bibio sp 1, Plecia sp. 7, Plecia sp. 12, Plecia sp. 1, Plecia sp. 2, Plecia rhenana, Plecia sp. 10, Plecia sp. 11, Plecia sp. 3, Plecia sp. 5, Plecia sp. 6, Plecia sp. 13, Plecia sp. 14, Penthetria sp 1, Bibio sp 4, Bibio sp 2, Bibio sp 6, Bibio sp 3, Bibio sp 5, Plecia sp. 8, Plecia sp. 15, Plecia lapidaria (122010) | |
| Chattian | France (Bouches-Du-Rhone) | Bibio sp., Dilophus sp., Plecia bucklandi, Plecia splendida, Plecia lugens, Bibio spectabilis, Bibio romani, Bibio disjectus, Bibio serresi, Plecia dilatativentris, Plecia livida, Plecia gracilis, Plecia elegans, Bibio curtisi, Plecia guinieri, Plecia brevipennis, Bibio curtisii, Penthetria nervisinuata, Bibio aquaesextiae, Bibiodes provincialis (122501) Bibio sp., Plecia bucklandi, Plecia splendida, Bibio conformans, Plecia lugens, Bibio morio, Bibio painvini, Bibio spectabilis, Bibio romani, Bibio disjectus, Bibio fusiformis, Bibio detractus, Bibio serresi, Plecia dilatativentris, Plecia nervisinuata, Plecia livida, Plecia gracilis, Plecia elegans, Plecia constricta, Bibio curtisi, Plecia guinieri, Plecia brevipennis, Plecia lygaeoides (138838) Bibionidae indet. (153412) Plecia bucklandi, Plecia splendida, Bibio morio, Bibio detractus, Plecia livida, Plecia gracilis, Plecia elegans, Plecia retracta, Plecia constricta, Bibio curtisi, Plecia brevipennis (122509) Plecia livida, Plecia elegans (122511) Plecia rhenana, Plecia lapidaria, Bibio baumbergeri, Bibio painvini, Bibio spectabilis, Penthetria subterranea (113712) Protomyia bucklandi (122878) Protomyia gracilis, Protomyia elegans, Bibio curtisii, Plecia macilenta, Penthetria nervisinuata (122876) Protomyia lygaeoides, Protomyia bucklandi, Bibio morio, Protomyia gracilis, Protomyia brevipennis, Bibio fusiformis, Empis sp., Penthetria sp., Diptera indet., Bibio moestus (122877) | |
| Chattian | France | Plecia livida, Penthetria luberonica (183981) | |
| Chattian | France (Provence-Alpes-Côte d'Azur) | Bibionidae indet., Plecia splendida, Bibio conformans, Plecia lugens, Bibio painvini, Bibio spectabilis, Bibio luteipennis, Bibio romani, Bibio disjectus, Bibio detractus, Bibio serresi, Plecia dilatativentris, Plecia nervisinuata, Plecia livida, Plecia gracilis, Plecia elegans, Plecia retracta, Plecia acutiventris, Bibio curtisi, Plecia brevipennis, Bibio fusiformis, Plecia lygaeoides (122488) Plecia splendida, Bibio conformans, Bibio serresi, Plecia livida, Plecia gracilis, Plecia elegans, Plecia constricta, Bibio curtisi (122487) | |
| Late/Upper Oligocene | USA (Colorado) | Plecia creedensis (117882 121977) | |
| Late/Upper Oligocene | France (Bouches-du-Rhone) | Bibio celasensis (113989) Bibio nigripennis (122464) | |
| Late/Upper Oligocene | Germany | Bibio tertiarius (152951) | |
| MP 28 | Germany (Rheinland-Pfalz) | Bibio xylophilus, Penthetria nigra, Plecia dubia, Bibio castaneipennis (71942) | |
| MP 30 | Germany | Helophilus primarius, Bibio lignarius, Bibio xylophilus, Phthiria dubia (115802) Plecia rhenana, Protomyia luctuosa, Protomyia lapidaria, Bibio deletus, Protomyia hypogaea, Protomyia antenata, Bibio pannosus, Protomyia abava, Protomyia collossea, Protomyia winnertzi, Protomyia grossa, Protomyia proserpina, Protomyia macrocephala, Protomyia exposititia, Protomyia stygia, Protomyia pinguis, Protomyia veterana, Protomyia grandaeva, Protomyia luteola, Protomyia schineri, Protomyia elongata, Protomyia gracilenta, Protomyia heeri, Plecia heroica, Dilophus krantzii, Bibio mimas (123945) Protomyia luctuosa, Protomyia lapidaria, Bibio janus, Protomyia winnertzi, Protomyia macrocephala, Protomyia exposititia, Protomyia stygia, Protomyia grandaeva, Dilophus krantzii (154946) | |
| MP 30 | Germany (Nordrhein-Westfalen) | Bibiopsis volgeri, Bibio lignarius (127628) Plecia superba, Bibio infumatus, Bibio brachypteroides, Bibio janus, Plecia pulchella, Protomyia hypogaea, Protomyia antenata, Protomyia sluiteri, Protomyia veterana, Protomyia elongata, Bibio germari, Bibio heydeni, Protomyia expositia, Protomyia elegantula, Plecia carbonum (138696) | |
| Aquitanian | Türkiye | Bibio anatolicus, Bibio nigricosta, Penthetria beskonakensis (123307) | |
| Aquitanian | Germany | Penthetria sp. (190126) Plecia sp., Bibio sticheli (123872) | |
| Early/Lower Miocene | Spain (Castellon) | Bibio sp. (225781) | |
| Early/Lower Miocene | Germany | Bibio antiquus (152937) | |
| Early/Lower Miocene | Japan | Bibio sadoensis (114003) | |
| Early/Lower Miocene | Mexico (Chiapas) | Plecia pristina (140210) | |
| Early/Lower Miocene | Slovenia | Bibio sticheli (152681) | |
| Burdigalian | Russian Federation | Plecia amagua, Plecia kudiella, Plecia refracta, Plecia redempta, Plecia kuznetzovi, Plecia obsitula (113705) | |
| Burdigalian | Austria | Bibionidae indet. (139943) | |
| Burdigalian | China (Shandong) | Bibio shanwangensis, Bibio flavissimus (130412) Bibio shanwangensis, Bibio miocenica, Plecia ludongensis, Plecia solaris, Bibio ventricosus, Bibio expansus, Bibio deformatus, Fungivora bullata, Bibio brunneus, Bibio cratopodus, Bibio ruptineurus, Clothonopsis miocenica (128550) Bibio shanwangensis, Plecia diatoma, Plecia vulcania, Plecia villosa, Plecia rectivenia, Bibio brunneifemus, Bibio peraffinis, Bibio brachypodus, Bibio tranopterus, Bibio cratopodus, Bibio varineurus, Bibio longineurus, Bibio eximius (128551) Plecia capitata (138944) Plecia fumosa, Plecia bivalvula, Plecia gracilenta (138946) Plecia platoptera, Bibio penisularis, Bibio udus (138945) Plecia spinula, Plecia aculeolata, Plecia villosa, Bibio ventricosus, Bibio expansus, Bibio colossoceps, Bibio abdominalis, Bibio tranopterus, Bibio varineurus (128552) | |
| Burdigalian | Greece | Penthetria fuchsi (144526) Penthetria marinosi (130761) | |
| Burdigalian - Langhian | Dominican Republic | Dilophus matilei (156229) Plecia pristina (122474) | |
| MN 4 | Czechia | Bibiopsis egerana, Bibiopsis imperialis, Bibio formosus, Bibio elegantulus, Protomyia bohemica, Plecia quaesita (117612) | |
| Langhian | USA (Idaho) | Bibio latahensis, Bibio testeus (97616) | |
| Langhian | Japan | Bibio sp., Plecia kanetakii (103101) | |
| Langhian | USA (Washington) | Bibio sp. (97619) | |
| MN 5 - MN 6 | Bosnia and Herzegovina | Bibio sp., Plecia sp. (211309) | |
| Middle Miocene | Bulgaria | Plecia sp. (181730) | |
| Middle Miocene | Germany | Bibiopsis carbonum (152936) | |
| Middle Miocene - Late/Upper Miocene | Thailand | Plecia sp. (140983) | |
| Badenian | Austria (Steiermark) | Lithobibio styriacus (139945) | |
| Sarmatian | Croatia | Bibio gracilis, Protomyia latipennis (156758) Bibio lividus, Protomyia anthracina (113693) Bibio moestus, Bibio maculatus, Mesomyia brevis (156759) Bibio sp., Penthetria longa, Penthetria obsoleta, Bibio partschii, Bibio maculatus, Bibiopsis murchisonii, Penthetria anthracina (117247) Plecia lygaeoides, Plecia lugubris, Bibio giganteus, Bibio enterodelus, Penthetria anthracina, Bibio crassinervis, Penthetria cimicoides, Dilophus pinguis (224110) Protomyia bucklandi, Protomyia longa, Bibio morio, Plecia lugubris, Bibio linearis, Bibio gracilis, Bibio incrassatus, Bibio lividus, Bibiopsis brevicollis, Protomyia anthracina, Bibio pinguis (117761) Protomyia lygaeoides, Protomyia longa, Bibio giganteus, Bibio gracilis, Bibio maculatus, Bibio enterodelus, Bibio firmus, Bibiopsis cimicoides, Protomyia latipennis, Bibio murchisonis, Bibio pinguis (117246) | |
| Sarmatian | Germany (Baden-Württemberg) | Bibio fusiformis, Bibio brevis, Bibio moestus, Bibio pulchellus, Protomyia jucunda, Protomyia affinis (113689) Bibio fusiformis, Bibio brevis, Bibio moestus, Plecia hilaris, Bibio elongatus, Bibio angustatus, Bibio pulchellus, Protomyia jucunda, Protomyia speciosa, Bibio crassinervis, Dilophus pumilio, Penthetria brevicollis, Plecia macilenta (117252) Bibio fusiformis, Bibio brevis, Bibio moestus, Plecia hilaris, Bibio oblongus (115304) Bibio fusiformis, Bibio brevis, Bibio obsoletus, Bibio moestus, Bibio elongatus, Bibio angustatus, Bibio oblongus, Protomyia jucunda, Protomyia amoena (115305) Bibio moestus (117338) Bibio pinguis (117260) | |
| Tortonian | France | Bibio joursacencis (138766) | |
| Tortonian | Iceland | Bibio sp., Dilophus sp., Penthetria sp., Bibio edda (154940) | |
| Late/Upper Miocene | Romania | Bibio kochii (155724) | |
| Turolian | France (Ardèche) | Bibio boulayi (156164) Bibio joursacensis, Plecia vergnei (172761) | |
| Messinian | Italy | Bibio sp., Plecia spp., Penthetria sp., Plecia castellaroi, Plecia pisaurensis, Plecia baglii (157306) | |
| Messinian | Italy (Tuscany) | Bibio sp., Penthetria sp. (113751) | |
| Messinian | USA (Alaska) | Bibio sp. (103102) | |
| Piacenzian | Germany (Niedersachsen) | Bibio contererus (240307) Bibio contererus, Bibio acaptus (155691) Bibio contererus, Bibio acaptus, Bibio nebulosus, Dilophus flavipes, Dilophus plionanus (167949) Dilophus magnus, Bibio contererus, Bibio nebulosus, Dilophus flavipes (155690) Dilophus magnus, Bibio enormus, Bibio latiantennatus, Bibio medioalbus, Bibio contererus, Bibio mancus, Bibio dormitus, Bibio soldatus, Bibio tripus, Bibio acaptus, Bibio carnificus, Bibio constringtus, Bibio praecidus, Bibio deflectus, Bibio similis, Bibio discerptus, Bibio nebulosus, Bibio vixdus, Bibio cruciformis, Bibio albangulus, Bibio anasiformis, Bibio tenuis, Dilophus flavipes, Plecia sp., Bibio sp. (109196) | |
| Piacenzian | Japan | Bibionidae indet., Bibio sp., Penthetria togoensis, Plecia intima (160140) Bibionidae indet., Plecia sp., Penthetria velutina, Bibio tenebrosus (188976) | |
| Gelasian | France | Bibionidae indet., Plecia sp., Plecia brunneipennis (113996) Plecia lugubris, Plecia vergnei (113994) Plecia sp. (138769) | |
| Late/Upper Pleistocene | United Kingdom (Worcestershire) | Dilophus sp. (163070 163075) | |
| Late/Upper Pleistocene | USA (Washington) | Bibionidae indet. (167662) |