PBDB Taxon

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Myrmicinae (ant)

Insecta - Hymenoptera - Formicidae

Taxonomy

Myrmicinae was named by Lepeletier de Saint-Fargeau (1835). It is extant.

It was reranked as the subfamily Myrmicidae by Scudder (1878).

It was assigned to Hymenoptera by Scudder (1878); and to Formicidae by Hong (1984), Hong and Wang (1985), Bolton (2003), Wappler (2003), Dlussky et al. (2004), Archibald et al. (2006), Bolton (2012), Dlussky and Perfilieva (2014), Dlussky et al. (2015), Ward et al. (2015), Zharkov and Dubovikoff (2022).

Synonyms

Aphaenogaster longaeva was named by Scudder (1877). Its type specimen is GSC 33, an exoskeleton, and it is a compression fossil. Its type locality is Quesnel, which is in a Priabonian lacustrine shale in the Australian Creek Formation of Canada.

It was considered a nomen dubium by Carpenter (1930); it was considered a nomen dubium by Radchenko and Perkovsky (2016).
Sinomyrmex was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Sinomyrmex brunnea was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002); it was corrected as Sinomyrmex brunneus by Bolton (2012).
Sinomyrmex furvis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (late 1970s collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Fushunomyrmex was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Fushunomyrmex liaoningensis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Orbigastrula was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Orbigastrula guchengziensis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Quineangulicapito was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Quineangulicapito fushunensis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Sphaerogasterites was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Sphaerogasterites longipetiolatus was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (late 1970s collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Brachytarsites was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Brachytarsites dongzhouheensis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Quadrulicapito was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Quadrulicapito longa was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).
Clavipetiola was named by Hong (2002).

It was considered a nomen nudum by Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Clavipetiola asiatica was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (late 1970s collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002).
Wumyrmex was named by Hong (2002).

It was considered a nomen nudum by Hong (2002), Hong (2002).

It was assigned to Myrmicinae by Bolton (2012), Ward et al. (2015).
Wumyrmex furvis was named by Hong (2002). It is an inclusion in amber. Its type locality is Xilutian coal mine, Fushun amber (post-1980 collection), which is in a Ypresian terrestrial amber in China.

It was considered a nomen nudum by Hong (2002), Hong (2002).

Synonymy list

Year	Name and author
1835	Myrmicinae Lepeletier de Saint-Fargeau p. 169
1877	Aphaenogaster longaeva Scudder p. 267
1878	Myrmicidae Scudder p. 748
1890	Aphaenogaster longaeva Scudder p. 615 figs. Pl 3 fig 28
1984	Myrmicinae Hong p. 8
1985	Myrmicinae Hong and Wang p. 20
2002	Sinomyrmex Hong p. 546
2002	Sinomyrmex brunnea Hong p. 547 figs. Pl 41, fig 2; text-fig 2.8.350-357
2002	Sinomyrmex furvis Hong p. 548 figs. Pl 41, fig 4; text-fig 2.8.358-364
2002	Fushunomyrmex Hong p. 550
2002	Fushunomyrmex liaoningensis Hong p. 550 figs. 2.8.365-374
2002	Orbigastrula Hong p. 552
2002	Orbigastrula guchengziensis Hong p. 553 figs. Pl 41, fig 3; text-fig 2.8.375-381
2002	Quineangulicapito Hong p. 554
2002	Quineangulicapito fushunensis Hong p. 555 figs. Pl 43, fig 1; text-fig 2.8.382-391
2002	Sphaerogasterites Hong p. 557
2002	Sphaerogasterites longipetiolatus Hong p. 557 figs. Pl 44, fig 1; text-fig 2.8.392-398
2002	Brachytarsites Hong p. 559
2002	Brachytarsites dongzhouheensis Hong p. 559 figs. 2.8.399-404
2002	Quadrulicapito Hong p. 561
2002	Quadrulicapito longa Hong p. 562 figs. Pl 43, fig 2; text-fig 2.8.405-412
2002	Clavipetiola Hong p. 564
2002	Clavipetiola asiatica Hong p. 564 figs. 2.8.413-422
2002	Wumyrmex Hong p. 566
2002	Wumyrmex furvis Hong p. 566 figs. Pl 44, fig 2; text-fig 2.8.423-434
2003	Myrmicinae Bolton p. 52
2003	Myrmicinae Wappler p. 132
2004	Myrmicinae Dlussky et al. p. 9
2006	Myrmicinae Archibald et al. p. 492
2012	Aphaenogaster longaeva Bolton
2012	Brachytarsites Bolton
2012	Brachytarsites dongzhouheensis Bolton
2012	Clavipetiola Bolton
2012	Clavipetiola asiatica Bolton
2012	Fushunomyrmex Bolton
2012	Fushunomyrmex liaoningensis Bolton
2012	Myrmicinae Bolton
2012	Orbigastrula Bolton
2012	Orbigastrula guchengziensis Bolton
2012	Quadrulicapito Bolton
2012	Quadrulicapito longa Bolton
2012	Quineangulicapito Bolton
2012	Quineangulicapito fushunensis Bolton
2012	Sinomyrmex Bolton
2012	Sinomyrmex brunneus Bolton
2012	Sinomyrmex furvis Bolton
2012	Sphaerogasterites Bolton
2012	Sphaerogasterites longipetiolatus Bolton
2012	Wumyrmex Bolton
2012	Wumyrmex furvis Bolton
2014	Myrmicinae Dlussky and Perfilieva p. 432
2015	Myrmicinae Dlussky et al. p. 143
2015	Myrmicinae Ward et al. p. 76
2015	Brachytarsites Ward et al. p. 77
2015	Clavipetiola Ward et al. p. 77
2015	Fushunomyrmex Ward et al. p. 77
2015	Orbigastrula Ward et al. p. 77
2015	Quadrulicapito Ward et al. p. 77
2015	Quineangulicapito Ward et al. p. 77
2015	Sinomyrmex Ward et al. p. 77
2015	Sphaerogasterites Ward et al. p. 77
2015	Wumyrmex Ward et al. p. 77
2022	Myrmicinae Zharkov and Dubovikoff

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata

Rank	Name	Author
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Hymenopterida
order	Hymenoptera
—	Unicalcarida
suborder	Apocrita	Gerstaecker 1867
—	Aculeata	()
superfamily	Formicoidea
family	Formicidae	()
subfamily	Myrmicinae
subfamily	Myrmicinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Myrmicinae Lepeletier de Saint-Fargeau 1835 [ant]

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G. †Afromyrma Dlussky et al. 2004

†Afromyrma petrosa Dlussky et al. 2004

G. †Agastomyrma Dlussky et al. 2015

†Agastomyrma laticeps Dlussky et al. 2015

Tr. Attini Smith 1858

G. Acanthognathus Mayr 1887

†Acanthognathus poinari Baroni Urbani 1994

Subtr. Attina Smith 1858

G. Acromyrmex Mayr 1865

G. Apterostigma Mayr 1865

†Apterostigma electropilosum Schultz 2007

†Apterostigma eowilsoni Schultz 2007

G. Atta Fabricius 1804

G. Cyphomyrmex Mayr 1862

†Cyphomyrmex maya de Andrade 2003

†Cyphomyrmex taino de Andrade 2003

G. Mycetomoellerius Solomon et al. 2019

†Mycetomoellerius primaevus Baroni Urbani 1980

G. Trachymyrmex Forel 1893

G. Basiceros Schulz 1906

†Basiceros enana Fiorentino et al. 2025

G. Cephalotes Latreille 1802

†Cephalotes alveolatus Vierbergen and Scheven 1995

†Cephalotes bloosi Baroni Urbani and de Andrade 1999

†Cephalotes brevispineus de Andrade 1999

†Cephalotes caribicus de Andrade 1999

†Cephalotes dieteri Baroni Urbani 1999

†Cephalotes hispaniolicus de Andrade 1999

†Cephalotes integerrimus Vierbergen and Scheven 1995

†Cephalotes jansei Vierbergen and Scheven 1995

†Cephalotes maya de Andrade 1999

†Cephalotes obscurus Vierbergen and Scheven 1995

†Cephalotes olmecus de Andrade 1999

†Cephalotes poinari Baroni Urbani 1999

†Cephalotes resinae de Andrade 1999

†Cephalotes serratus Vierbergen and Scheven 1995

Invalid names: Exocryptocerus elevatus Vierbergen and Scheven 1995 [synonym], Exocryptocerus truncatus Vierbergen and Scheven 1995 [synonym]

†Cephalotes squamosus Vierbergen and Scheven 1995

Invalid names: Zacryptocerus pseudoaztecus Vierbergen and Scheven 1995 [synonym]

†Cephalotes sucinus de Andrade 1999

†Cephalotes taino de Andrade 1999

†Cephalotes ventriosus de Andrade 1999

Invalid names: Exocryptocerus Vierbergen and Scheven 1995 [synonym], Zacryptocerus Ashmead 1905 [synonym]

G. Pheidole Westwood 1840

†Pheidole anticua Casadei-Ferreira et al. 2019

†Pheidole chaan Varela-Hernández and Flores-Zapoteco 2024

†Pheidole cordata Holl 1829

Pheidole pallidula Nylander 1849

†Pheidole pauchil Varela-Hernández and Riquelme 2021

Pheidole plagiaria Smith 1860

†Pheidole primigenia Baroni Urbani 1995

†Pheidole rasnitsyni Dubovikoff 2011

†Pheidole tertiaria Carpenter 1930

†Pheidole tethepa Wilson 1985

G. Strumigenys Smith 1860

†Strumigenys electrina de Andrade 1994

†Strumigenys pilosula de Andrade 2007

†Strumigenys poinari Baroni Urbani and de Andrade 2007

†Strumigenys schleeorum Baroni Urbani 1994

Invalid names: Smithistruma Brown 1948 [synonym]

Invalid names: Cephalotini Smith 1949 [synonym], Dacetini Forel 1892 [synonym], Pheidolini Emery 1877 [synonym]

G. †Biamomyrma Dlussky et al. 2015

†Biamomyrma lata Dlussky et al. 2015

†Biamomyrma rugosa Dlussky et al. 2015

†Biamomyrma zherikhini Dlussky et al. 2015

G. †Bilobomyrma Radchenko and Dlussky 2013

†Bilobomyrma baltica Radchenko and Dlussky 2013

†Bilobomyrma ukrainica Radchenko and Dlussky 2013

G. †Boltonidris Radchenko and Dlussky 2012

†Boltonidris mirabilis Radchenko and Dlussky 2012

G. †Cephalomyrmex Carpenter 1930

†Cephalomyrmex rotundatus Carpenter 1930

G. †Cephites Heer 1849

†Cephites fragilis Heer 1849

†Cephites oeningensis Heer 1849

Tr. Crematogastrini Forel 1893

G. Cardiocondyla Emery 1869

G. Carebara Westwood 1840

†Carebara antiqua Mayr 1868

†Carebara bohemica Novák 1878

†Carebara groehni Radchenko and Dlussky 2019

†Carebara kutscheri Radchenko et al. 2019

†Carebara nitida Dlussky 2002

†Carebara rugiceps Heer 1849

†Carebara schossnicensis Assmann 1870

†Carebara sophiae Emery 1891

†Carebara thorali Théobald 1937

†Carebara ucrainica Dlussky 2002

Invalid names: Aeromyrma Forel 1891 [synonym], Erebomyrma Wheeler 1903 [synonym], Oligomyrmex Mayr 1867 [synonym], Pheidologeton Mayr 1862 [synonym]

G. Cataulacus Smith 1853

†Cataulacus planiceps Emery 1891

†Cataulacus plebeius de Andrade and Baroni Urbani 2004

†Cataulacus resinosus Viehmeyer 1913

†Cataulacus silvestrii Emery 1891

Cataulacus taprobanae Smith 1853

G. Crematogaster Lund 1831

G. †Eocenomyrma Dlussky and Radchenko 2006

†Eocenomyrma breviscapa Radchenko and Dlussky 2016

†Eocenomyrma electrina Dlussky and Radchenko 2006

†Eocenomyrma elegantula Dlussky and Radchenko 2006

†Eocenomyrma groehni Radchenko 2020

†Eocenomyrma orthospina Dlussky and Radchenko 2006

†Eocenomyrma rugosostriata Mayr 1868

†Eocenomyrma ukrainica Radchenko and Dlussky 2016

G. Gauromyrmex Menozzi 1933

G. †Hypopomyrmex Emery 1891

†Hypopomyrmex bombiccii Emery 1891

G. Leptothorax Mayr 1855

Subg. Leptothorax (Macromischa) Roger 1863

Leptothorax muscorum Nylander 1846

G. †Lonchomyrmex Mayr 1867

†Lonchomyrmex freyeri Heer 1867

†Lonchomyrmex nigritus Assmann 1870

G. Lophomyrmex Emery 1892

G. Lordomyrma Emery 1897

G. Meranoplus Smith 1853

G. Myrmecina Curtis 1829

G. Nesomyrmex Wheeler 1910

†Nesomyrmex caritatis de Andrade 1999

†Nesomyrmex dominicanus de Andrade 1999

G. †Oxyidris Wilson 1985

†Oxyidris antillana Wilson 1985

G. †Parameranoplus Wheeler 1915

†Parameranoplus primaevus Wheeler 1915

G. Podomyrma Smith 1859

†Podomyrma mayri Emery 1891

Invalid names: Acrostigma Emery 1891 [synonym]

G. Pristomyrmex Mayr 1866

†Pristomyrmex archaios Radchenko and Dlussky 2018

†Pristomyrmex elmesi Radchenko and Dlussky 2018

†Pristomyrmex rasnitsyni Dlussky and Radchenko 2011

Pristomyrmex umbripennis Smith 1863

Invalid names: Pristomyrmex parumpunctatus Emery 1887 [synonym]

G. Proatta Forel 1912

G. †Proleptothorax Radchenko et al. 2018

†Proleptothorax primitivus Radchenko et al. 2018

G. †Stigmomyrmex Mayr 1868

†Stigmomyrmex rugulosus Radchenko and Dlussky 2017

†Stigmomyrmex venustus Mayr 1868

G. †Stiphromyrmex Wheeler 1915

†Stiphromyrmex robustus Mayr 1868

G. Temnothorax Mayr 1861

†Temnothorax praecreolus de Andrade 1992

G. Tetramorium Mayr 1855

†Tetramorium kulickae Radchenko and Dlussky 2015

†Tetramorium paraarmatum Radchenko and Dlussky 2015

G. †Thanacomyrmex Chény et al. 2019

†Thanacomyrmex hoffeinsorum Chény et al. 2019

G. Vollenhovia Mayr 1865

†Vollenhovia beyrichi Mayr 1868

Invalid names: Propodomyrma samlandica Wheeler 1910 [synonym]

†Vollenhovia prisca André 1895

Invalid names: Propodomyrma Wheeler 1910 [synonym]

Invalid names: Cataulacini Emery 1895 [synonym], Formicoxenini Forel 1893 [synonym], Meranoplini Emery 1914 [synonym], Myrmecinini Ashmead 1905 [synonym], Tetramoriini Emery 1895 [synonym]

G. †Electromyrmex Wheeler 1910

†Electromyrmex klebsi Wheeler 1910

†Electromyrmex wheeleri Radchenko and Dlussky 2018

G. †Enneamerus Mayr 1868

†Enneamerus costatus Radchenko and Dlussky 2017

†Enneamerus reticulatus Mayr 1868

G. †Eocenidris Wilson 1985

†Eocenidris crassa Wilson 1985

G. †Eomyrmex Hong 1974

†Eomyrmex guchengziensis Hong 1974

G. †Eotemnothorax Radchenko 2024

†Eotemnothorax armatum Radchenko 2024

†Eotemnothorax balticus Radchenko 2024

†Eotemnothorax foveocephalus Radchenko 2024

†Eotemnothorax glaesarius Wheeler 1915

†Eotemnothorax gracilis Mayr 1868

†Eotemnothorax groehni Radchenko 2024

†Eotemnothorax hystriculus Wheeler 1915

†Eotemnothorax khomychi Radchenko 2024

†Eotemnothorax longaevus Wheeler 1915

†Eotemnothorax odontos Radchenko 2024

†Eotemnothorax pedunculatus Radchenko 2024

†Eotemnothorax perkovskii Radchenko 2024

†Eotemnothorax petiolatus Mayr 1868

†Eotemnothorax placivus Wheeler 1915

†Eotemnothorax punctatus Radchenko 2024

†Eotemnothorax rav Radchenko 2024

†Eotemnothorax rectispinus Radchenko 2024

†Eotemnothorax rhytidus Radchenko 2024

G. †Fallomyrma Dlussky and Radchenko 2006

†Fallomyrma anodonta Radchenko and Dlussky 2018

†Fallomyrma marginata Radchenko and Dlussky 2018

†Fallomyrma robusta Radchenko and Dlussky 2018

†Fallomyrma transversa Dlussky and Radchenko 2006

G. †Ilemomyrmex Wilson 1985

†Ilemomyrmex caecus Wilson 1985

G. †Incertogaster Boudinot 2024

†Incertogaster aurora LaPolla and Greenwalt 2015

†Incertogaster praecursor Emery 1891

†Incertogaster primitiva Radchenko and Dlussky 2019

G. †Lelejus Radchenko and Proshchalykin 2021

†Lelejus venustus Radchenko and Proshchalykin 2021

G. Melissotarsus Emery 1877

†Melissotarsus ethiopiensis Coty et al. 2016

G. †Miosolenopsis Zhang 1989

†Miosolenopsis fossilis Zhang 1989

G. †Myrmecites Dlussky and Rasnitsyn 2002

†Myrmecites dubius Dlussky et al. 2015

†Myrmecites latus Dlussky and Putyatina 2014

†Myrmecites major Dlussky et al. 2015

†Myrmecites pusillus Heer 1849

†Myrmecites rotundiceps Dlussky and Rasnitsyn 2002

†Myrmecites sibiricus Dlussky et al. 2015

Tr. Myrmicini Lepeletier de Saint-Fargeau 1835

G. Manica Jurine 1807

†Manica andrannae Zharkov and Dubovikoff 2022

G. Myrmica Latreille 1804

†Myrmica damzeni Radchenko 2023

†Myrmica dictyosa Radchenko 2023

†Myrmica electrina Radchenko 2023

†Myrmica eocenica Radchenko et al. 2007

Myrmica incompleta Provancher 1881

†Myrmica intermedia Wheeler 1915

Myrmica lobifrons Pergande 1900

†Myrmica longispinosa Mayr 1868

†Myrmica nungesseri Jessen 2020

†Myrmica paradoxa Radchenko et al. 2007

†Myrmica rudis Mayr 1868

†Myrmica saxonica Radchenko 2023

†Myrmica schaefersi Jessen 2020

Invalid names: Myrmica aemula Heer 1849 [nomen dubium], Myrmica angusticollis Heer 1849 [nomen dubium], Myrmica macrocephala Heer 1849 [nomen dubium], Myrmica molassica Heer 1849 [nomen dubium], Myrmica nebulosa Novák 1878 [nomen dubium], Myrmica obsoleta Heer 1849 [nomen dubium], Nothomyrmica Wheeler 1915 [synonym]

G. †Plesiomyrmex Dlussky and Radchenko 2009

†Plesiomyrmex tubulatus Dlussky and Radchenko 2009

G. †Protomyrmica Dlussky and Radchenko 2009

†Protomyrmica atavia Dlussky and Radchenko 2009

Tr. Pogonomyrmecini Ward et al. 2014

G. Pogonomyrmex Mayr 1868

†Pogonomyrmex fossilis Carpenter 1930

Tr. Solenopsidini Forel 1893

G. Monomorium Mayr 1855

†Monomorium kugleri Radchenko and Perkovsky 2009

†Monomorium mayrianum Wheeler 1915

Invalid names: Lampromyrmex gracillimus Mayr 1868 [replaced]

†Monomorium pilipes Mayr 1868

Invalid names: Lampromyrmex Mayr 1868 [synonym]

G. Myrmicaria Saunders 1842

G. Rogeria Emery 1894

G. Solenopsis Westwood 1840

†Solenopsis alena Özdikmen 2010

Invalid names: Solenopsis major Theobald 1937 [replaced]

†Solenopsis atavinus Perfilieva et al. 2017

†Solenopsis blanda Förster 1891

Invalid names: Attopsis acuta Förster 1891 [synonym]

†Solenopsis foersteri Theobald 1937

†Solenopsis maxima Förster 1891

†Solenopsis moesta Förster 1891

†Solenopsis privata Förster 1891

†Solenopsis superba Förster 1891

Invalid names: Attopsis extensa Förster 1891 [synonym]

†Solenopsis valida Förster 1891

Invalid names: Eurytarsites Hong 2002 [nomen nudum], Monomoriini Wheeler 1915 [synonym]

G. †Solenopsites Dlussky and Rasnitsyn 2002

†Solenopsites abdita LaPolla and Greenwalt 2015

†Solenopsites minutus Dlussky and Rasnitsyn 2002

†Solenopsites rossi Dlussky and Perfilieva 2014

Tr. Stenammini Ashmead 1905

G. Aphaenogaster Mayr 1853

†Aphaenogaster amphioceanica de Andrade 1995

†Aphaenogaster antiqua Dlussky 2002

†Aphaenogaster archaica Meunier 1915

†Aphaenogaster dlusskyana Radchenko and Perkovsky 2016

†Aphaenogaster donisthorpei Carpenter 1930

†Aphaenogaster enspelensis Jessen 2020

†Aphaenogaster mayri Carpenter 1930

†Aphaenogaster mersa Wheeler 1915

†Aphaenogaster oligocenica Wheeler 1915

†Aphaenogaster praerelicta de Andrade 1995

†Aphaenogaster sommerfeldti Mayr 1868

Invalid names: Deromyrma Forel 1913 [synonym], Rectorhagas Lin 1982 [synonym], Sinaphaenogaster Zhang 1989 [synonym]

G. Goniomma Emery 1895

†Goniomma poschmanni Jessen 2020

G. Messor Forel 1890

†Messor sculpturatus Carpenter 1930

G. †Paraphaenogaster Dlussky 1981

†Paraphaenogaster avita Fujiyama 1970

†Paraphaenogaster bizeri Jessen 2020

†Paraphaenogaster dumetorum Lin 1982

†Paraphaenogaster freihauti Jessen 2020

†Paraphaenogaster hooleyana Dlussky and Perfilieva 2014

†Paraphaenogaster jurinei Heer 1849

Invalid names: Myrmica concinna Heer 1867 [synonym]

†Paraphaenogaster lapidescens Zhang 1989

†Paraphaenogaster loosi Jessen 2020

†Paraphaenogaster maculata Theobald 1937

†Paraphaenogaster maculipes Theobald 1937

†Paraphaenogaster microphthalmus Dlussky 1981

†Paraphaenogaster paludosa Zhang 1989

†Paraphaenogaster pannonica Bachmayer 1960

†Paraphaenogaster schindleri Jessen 2020

†Paraphaenogaster shanwangensis Hong 1984

†Paraphaenogaster tertiaria Heer 1849

Invalid names: Myrmica bicolor Heer 1867 [synonym]

†Paraphaenogaster ussuriensis Perfilieva 2022

†Paraphaenogaster wettlauferi Jessen 2020

†Paraphaenogaster wuttkei Jessen 2020

G. Stenamma Westwood 1840

†Stenamma berendti Mayr 1868

G. Veromessor Forel 1917

Veromessor andrei Mayr 1886

G. †Zhangidris Bolton 2003

†Zhangidris atopogaster Zhang 1989

Invalid names: Heteromyrmex Zhang 1989 [replaced]

Invalid names: Aphaenogaster longaeva Scudder 1877 [nomen dubium], Brachytarsites Hong 2002 [nomen nudum], Clavipetiola Hong 2002 [nomen nudum], Fushunomyrmex Hong 2002 [nomen nudum], Orbigastrula Hong 2002 [nomen nudum], Pseudomyrmini Emery 1914 [empty], Quadrulicapito Hong 2002 [nomen nudum], Quineangulicapito Hong 2002 [nomen nudum], Sinomyrmex Hong 2002 [nomen nudum], Solenopsidiini Forel 1893 [empty], Sphaerogasterites Hong 2002 [nomen nudum], Wumyrmex Hong 2002 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Holocene or 93.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma

Collections (149 total)

Time interval	Ma	Country or state	Original ID and collection number
Turonian	93.9 - 89.8	Botswana	Afromyrma petrosa (123430)
Ypresian	56.0 - 48.07	China (Liaoning)	Eomyrmex guchengziensis (125469) Sinomyrmex brunnea, Fushunomyrmex liaoningensis, Orbigastrula guchengziensis, Quineangulicapito fushunensis, Brachytarsites dongzhouheensis, Quadrulicapito longa, Wumyrmex furvis (124820) Sinomyrmex furvis, Sphaerogasterites longipetiolatus, Clavipetiola asiatica, Eurytarsites fushunensis (124530)
Ypresian	56.0 - 48.07	Canada (British Columbia)	Leptothorax sp. (123795)
Bridgerian	50.5 - 46.2	USA (Colorado)	Myrmecites rotundiceps, Solenopsites minutus (138664)
Bridgerian	50.5 - 46.2	USA (Wyoming)	Myrmica sp. (121306)
Lutetian	48.07 - 41.03	Russian Federation	Aphaenogaster dlusskyana (107340)
Lutetian	48.07 - 41.03	USA (Montana)	Crematogaster aurora, Solenopsites abdita (117614)
MP 11	48.07 - 41.03	Germany (Hessen)	Myrmicinae indet. (120798)
Bartonian	41.03 - 37.71	USA (Arkansas)	Eocenidris crassa (122391)
Late/Upper Eocene	37.71 - 33.9	USA (Colorado)	Aphaenogaster mayri, Messor sculpturatus (122093)
Priabonian	37.71 - 33.9	Germany (Halle)	Eocenomyrma rugosostriata, Fallomyrma transversa, Stigmomyrmex rugulosus, Electromyrmex wheeleri, Myrmica saxonica, Eotemnothorax gracilis (129227) Eotemnothorax odontos (149487) Monomorium mayrianum, Stigmomyrmex venustus, Fallomyrma transversa, Plesiomyrmex tubulatus, Pristomyrmex archaios, Carebara kutscheri, Eotemnothorax gracilis, Eotemnothorax longaevus, Eotemnothorax rectispinus, Eotemnothorax pedunculatus (129228) Myrmica paradoxa, Eocenomyrma rugosostriata (123910)
Priabonian	37.71 - 33.9	Poland	Aphaenogaster sommerfeldti, Enneamerus reticulatus, Aphaenogaster berendti, Macromischa petiolata, Lampromyrmex gracillimus, Stigmomyrmex robustus (109625) Monomorium pilipes, Stigmomyrmex venustus, Enneamerus reticulatus, Pheidologeton antiquus, Lampromyrmex gracillimus (120811) Monomorium sp., Myrmica rudis, Stiphromyrmex robustus, Enneamerus reticulatus, Eocenomyrma rugosostriata, Eocenomyrma orthospina, Tetramorium paraarmatum, Tetramorium kulickae (120813) Stigmomyrmex rugulosus, Myrmica dictyosa (110737)
Priabonian	37.71 - 33.9	Russian Federation	Nothomyrmica rudis (123930) Thanacomyrmex hoffeinsorum (123419)
Priabonian	37.71 - 33.9	United Kingdom (England)	Paraphaenogaster hooleyana (123960) Solenopsites rossi (123959)
Priabonian	37.71 - 33.9	Russian Federation (Primorskiy)	Biamomyrma zherikhini (123794) Myrmecites sibiricus, Myrmecites dubius, Myrmecites major (172678) Myrmicinae indet., Agastomyrma laticeps, Biamomyrma lata, Biamomyrma rugosa (172679)
Priabonian	37.71 - 33.9	Denmark	Myrmica eocenica, Eocenomyrma electrina, Fallomyrma transversa, Pristomyrmex rasnitsyni, Eotemnothorax armatum (123762)
Priabonian	37.71 - 33.9	Canada (British Columbia)	Aphaenogaster longaeva (121480)
Priabonian	37.71 - 33.9	Ukraine	Eocenomyrma rugosostriata, Eotemnothorax balticus (196704) Eotemnothorax gracilis, Eotemnothorax glaesarius, Eotemnothorax hystriculus, Eotemnothorax armatum, Eotemnothorax rhytidus, Eotemnothorax punctatus, Eotemnothorax khomychi (238338) Eotemnothorax rectispinus (154710 185682) Monomorium kugleri, Enneamerus reticulatus, Eocenomyrma orthospina, Fallomyrma transversa, Bilobomyrma ukrainica, Boltonidris mirabilis, Eocenomyrma ukrainica, Eocenomyrma breviscapa, Fallomyrma anodonta, Fallomyrma marginata, Fallomyrma robusta, Pristomyrmex elmesi, Proleptothorax primitivus, Crematogaster primitiva, Myrmica electrina, Eotemnothorax gracilis, Eotemnothorax hystriculus, Eotemnothorax placivus, Eotemnothorax armatum, Eotemnothorax punctatus, Eotemnothorax pedunculatus, Eotemnothorax perkovskii (122997) Myrmica rudis, Fallomyrma transversa (81780) Myrmicinae indet., Aphaenogaster antiqua, Oligomyrmex nitidus, Oligomyrmex ucrainicus, Fallomyrma transversa, Lelejus venustus, Eotemnothorax rav (113621)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	Aphaenogaster sommerfeldti (138654) Eotemnothorax balticus (123420 124003) Eotemnothorax petiolatus (234648) Eotemnothorax rectispinus (229453) Eotemnothorax rectispinus, Eotemnothorax foveocephalus, Eotemnothorax punctatus (147612) Macromischa beyrichi (123757) Macromischa prisca (138940) Macromischa rudis, Aphaenogaster sommerfeldti (138663) Macromischa rugosostriata (123962) Manica andrannae (192258) Monomorium mayrianum, Erebomyrma antiqua, Aphaenogaster sommerfeldti, Electromyrmex klebsi, Nothomyrmica rudis, Leptothorax gracilis, Parameranoplus primaevus, Enneamerus reticulatus, Propodomyrma samlandica (124073) Myrmica eocenica, Bilobomyrma baltica, Eotemnothorax hystriculus (122591) Myrmica intermedia, Stigmomyrmex venustus, Carebara antiqua, Eocenomyrma rugosostriata, Eocenomyrma elegantula, Enneamerus costatus, Electromyrmex wheeleri, Carebara groehni, Eotemnothorax gracilis, Eotemnothorax hystriculus, Eotemnothorax groehni, Eotemnothorax balticus, Eotemnothorax rhytidus, Eocenomyrma orthospina (110735) Myrmica longispinosa, Macromischa rudis, Nothomyrmica intermedia, Monomorium pilipes, Stenamma berendti, Aphaenogaster sommerfeldti, Aphaenogaster oligocenica, Aphaenogaster mersa, Leptothorax gracilis, Leptothorax glaesarius, Leptothorax longaevus, Leptothorax hystriculus, Leptothorax placivus, Stigmomyrmex venustus, Enneamerus reticulatus, Macromischa beyrichi, Macromischa rugosostriata, Macromischa petiolata, Pheidologeton antiquus, Lampromyrmex gracillimus, Stigmomyrmex robustus (123927) Myrmica longispinosa, Myrmica eocenica, Myrmica electrina, Eotemnothorax gracilis, Eotemnothorax rectispinus (123764) Myrmica longispinosa, Myrmica rudis, Myrmica intermedia, Myrmica damzeni (123964) Protomyrmica atavia (123766)
Chadronian	37.0 - 33.9	USA (Colorado)	Aphaenogaster mayri (113749 123119 124680) Aphaenogaster mayri, Aphaenogaster donisthorpei, Messor sculpturatus, Pheidole tertiaria, Pogonomyrmex fossilis, Cephalomyrmex rotundatus (106384) Aphaenogaster mayri, Messor sculpturatus (124002 124624 132362) Aphaenogaster mayri, Messor sculpturatus, Pogonomyrmex fossilis (127617)
Early/Lower Oligocene	33.9 - 27.3	France	Attopsis extensa, Attopsis moesta, Attopsis maxima, Attopsis valida, Attopsis longipennis, Attopsis longipes (121476) Attopsis privata, Attopsis blanda, Attopsis acuta, Attopsis longipennis (121473) Attopsis superba, Attopsis valida (121474) Solenopsis superba, Solenopsis foersteri, Solenopsis blanda, Aphaenogaster maculipes (122408)
Rupelian	33.9 - 27.3	Germany	Myrmicidae indet., Solenopsis maxima, Solenopsis valida, Solenopsis major, Solenopsis superba, Solenopsis foersteri, Solenopsis blanda, Aphaenogaster maculipes (122407)
Chattian	27.3 - 23.04	Russian Federation	Paraphaenogaster ussuriensis (125971)
Chattian	27.3 - 23.04	France (Bouches-Du-Rhone)	Aphaenogaster maculata (122501) Erebomyrma thorali (138838)
Chattian	27.3 - 23.04	France (Provence-Alpes-Côte d'Azur)	Aphaenogaster maculata (122487)
MP 28	27.3 - 23.04	Germany (Rheinland-Pfalz)	Myrmicinae indet., Paraphaenogaster sp., Paraphaenogaster loosi, Paraphaenogaster schindleri, Paraphaenogaster wettlauferi, Paraphaenogaster wuttkei, Goniomma poschmanni, Myrmica schaefersi, Myrmica nungesseri (160097) Paraphaenogaster schindleri (123354) Paraphaenogaster sp., Paraphaenogaster loosi, Paraphaenogaster schindleri, Paraphaenogaster bizeri, Paraphaenogaster freihauti, Aphaenogaster enspelensis, Goniomma poschmanni (211302) Paraphaenogaster sp., Stenammini indet., Paraphaenogaster loosi (211301) Stenammini indet. (215791)
MP 30	27.3 - 23.04	Germany (Nordrhein-Westfalen)	Myrmica sp., Myrmica archaica (138696)
Aquitanian	23.04 - 20.45	Germany	Myrmicinae indet., Formicinae indet., Aphaenogaster sp. (123872)
Early/Lower Miocene	23.04 - 15.98	Ethiopia	Melissotarsus ethiopiensis (183971)
Early/Lower Miocene	23.04 - 15.98	Mexico (Chiapas)	Aphaenogaster praerelicta, Cyphomyrmex maya, Cephalotes poinari, Cephalotes maya, Cephalotes olmecus (125757) Pheidole chaan (217216) Pheidole pauchil (198592)
Early/Lower Miocene	23.04 - 15.98	Mexico	Pheidole sp. 1, Pheidole sp. 2, Solenopsis sp., Crematogaster sp., Cephalotes sp. 1, Cephalotes sp. 2, Cephalotes ventriosus (21977)
Burdigalian	20.45 - 15.98	Austria	Myrmicinae indet. (139942)
Burdigalian	20.45 - 15.98	China (Shandong)	Heteromyrmex atopogaster, Paraphaenogaster shanwangensis (128550) Miosolenopsis fossilis, Aphaenogaster lapidescens, Aphaenogaster paludosa (128551) Rectorhagas dumetorum (130412)
Burdigalian	20.45 - 15.98	Japan (Gifu)	Solenopsis sp. (181074)
Burdigalian - Langhian	20.45 - 13.82	Dominican Republic	Apterostigma electropilosum, Apterostigma eowilsoni (122982) Basiceros enana (152045) Cephalotes integerrimus (124282 138708) Cephalotes serratus, Cephalotes jansei, Cephalotes squamosus, Acanthognathus poinari, Cephalotes brevispineus, Cephalotes bloosi, Cephalotes caribicus, Cephalotes sucinus, Leptothorax dominicanus, Strumigenys pilosula, Strumigenys poinari (122472) Cephalotes serratus, Cephalotes obscurus, Cephalotes squamosus, Leptothorax praecreolus (124281) Exocryptocerus jansei, Zacryptocerus squamosus, Zacryptocerus pseudoaztecus, Zacryptocerus obscurus, Cephalotes integerrimus, Cyphomyrmex taino, Ilemomyrmex caecus (21978) Exocryptocerus serratus, Cephalotes squamosus, Pheidole tethepa, Leptothorax caritatis (138700) Exocryptocerus serratus, Exocryptocerus elevatus, Exocryptocerus truncatus, Exocryptocerus jansei, Zacryptocerus integerrimus, Zacryptocerus squamosus, Zacryptocerus pseudoaztecus, Zacryptocerus alveolatus (124846) Leptothorax caritatis (138937) Nesomyrmex sp. (222615) Oxyidris antillana (138698) Pheidole anticua (203120) Pheidole sp., Cephalotes serratus, Cephalotes alveolatus, Cephalotes obscurus, Cephalotes integerrimus, Cephalotes squamosus, Strumigenys schleeorum, Strumigenys electrina, Aphaenogaster amphioceanica, Cephalotes hispaniolicus, Cephalotes caribicus, Cephalotes dieteri, Pheidole primigenia, Trachymyrmex primaevus (122474)
MN 4	16.9 - 15.98	Czechia	Pheidologeton bohemicus, Myrmica nebulosa (117612)
Langhian	15.98 - 13.82	Japan	Aphaenogaster avita (103101)
Langhian	15.98 - 13.82	USA (Idaho)	Messor sp. (97616)
Langhian	15.98 - 13.82	China (Fujian)	Aphaenogaster sp., Monomorium sp., Pheidole sp., Vollenhovia sp., Cataulacus sp., Crematogaster sp., Tetramorium sp., Carebara sp., Strumigenys sp., Cardiocondyla sp., Gauromyrmex sp., Lophomyrmex sp., Lordomyrma sp., Meranoplus sp., Myrmicaria sp., Proatta sp. (220273)
Langhian	15.98 - 13.82	Malaysia (Sarawak)	Cataulacus plebeius (138840)
Middle Miocene	15.98 - 11.63	Russian Federation	Paraphaenogaster microphthalmus (138756)
Badenian	13.82 - 12.8	Austria (Steiermark)	Myrmica obsoleta, Myrmica aemula (115306)
Sarmatian	12.8 - 11.63	Ukraine	Paraphaenogaster microphthalmus, Solenopsis atavinus (152283)
Sarmatian	12.8 - 11.63	Romania	Cataulacus sp. (156760)
Sarmatian	12.8 - 11.63	Croatia	Myrmica tertiaria, Myrmica jurinei, Myrmica pusilla, Formica freyeri, Myrmica bicolor, Myrmica concinna (117246) Paraphaenogaster jurinei, Myrmecites pusillus, Myrmecites latus (117247) Paraphaenogaster tertiaria (113698)
Sarmatian	12.8 - 11.63	Germany (Baden-Württemberg)	Cephites fragilis (113689) Myrmica angusticollis (115304) Myrmica macrocephala, Cephites oeningensis (115305) Myrmica macrocephala, Myrmica rugiceps, Myrmica molassica (113695) Myrmica rugiceps (122868) Myrmica tertiaria (122866 122867)
Late/Upper Miocene	11.63 - 5.333	Italy	Hypopomyrmex bombiccii, Podomyrma mayri, Cataulacus silvestrii, Cataulacus planiceps, Aeromyrma sophiae, Crematogaster praecursor (123432)
Late/Upper Miocene	11.63 - 5.333	Poland	Lonchomyrmex nigritus, Myrmica ruficeps (138936)
Turolian	8.725 - 5.09	France (Ardèche)	Solenopsis sp., Pheidole pallidula (172761)
Messinian	7.246 - 5.333	USA (Alaska)	Leptothorax sp. (123352) Myrmicinae indet., Aphaenogaster sp., Messor sp. (103102)
Messinian	7.246 - 5.333	Austria	Aphaenogaster pannonicus (127417)
Blancan	4.7 - 1.4	USA (Texas)	Myrmicinae indet. (154909)
Piacenzian	3.6 - 2.58	Germany (Niedersachsen)	Myrmicinae indet. (109196)
Gelasian	2.58 - 1.8	Greenland	Leptothorax sp. (160153) Myrmica sp. (160109 160144)
Gelasian	2.58 - 1.8	France	Erebomyrma antiqua (138769)
Rancholabrean	0.21 - 0.014	USA (California)	Veromessor andrei (152007 161529 161535)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	Myrmica lobifrons (169336 169350 169353)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Wisconsin)	Myrmica sp. (169406)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (British Columbia)	Myrmica sp. 1, Leptothorax muscorum, Myrmica incompleta (167673)
Holocene	0.0117 - 0.0	Colombia (Boyacá)	Myrmicinae indet., Pheidole sp., Solenopsis sp., Crematogaster sp., Zacryptocerus sp., Cephalotes sp., Acromyrmex sp., Rogeria sp. (215881)
Holocene	0.0117 - 0.0	Indonesia	Pristomyrmex parumpunctatus, Cataulacus resinosus, Pheidole plagiaria (182105)
Holocene	0.0117 - 0.0	Dominican Republic	Cephalotes resinae, Cephalotes taino (138880)
Holocene	0.0117 - 0.0	Tanzania	Aphaenogaster sp. (152160) Crematogaster sp., Pheidole cordata (234540) Formica cordata (147705) Pheidole rasnitsyni (124004)
Holocene	0.0117 - 0.0	Colombia (Santander)	Pheidole sp., Solenopsis sp., Crematogaster sp., Zacryptocerus sp., Strumigenys sp., Atta sp., Myrmecina sp., Smithistruma sp. (213295)